2015 in archosaur paleontology
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This article records new taxa of fossil archosaurs of every kind that are scheduled described during the year 2015, as well as other significant discoveries and events related to paleontology of archosaurs that are scheduled to occur in the year 2015.
Pseudosuchians
[edit]Research
[edit]- Revision of the type material of Rauisuchus tiradentes is published by Lautenschlager and Rauhut (2015).[1]
- A study on the changes in the morphology of vertebrae caused by increased adaptation to aquatic locomotion in the evolution of crocodylomorphs is published by Molnar et al. (2015).[2]
- A study of the evolutionary history of the clade Crocodyliformes is published by Bronzati, Montefeltro and Langer (2015).[3]
- A study of impact of the climate changes on the evolution and biodiversity of pseudosuchian archosaurs is published by Mannion et al. (2015).[4]
- A study on the phylogenetic position of thalattosuchians within Crocodylomorpha is published by Wilberg (2015).[5]
- A review of the diagnostic features of the species assigned to the genus Machimosaurus is published by Martin, Vincent & Falconnet (2015).[6]
- An isolated metriorhynchid tooth is described from the Early Cretaceous (Aptian) Hybla Formation (Sicily, Italy) by Chiarenza et al. (2015), extending the known geological range of Metriorhynchidae and Thalattosuchia by approximately 7–8 million years;[7] Fischer et al. (2015) subsequently caution that it might potentially be a pliosaurid tooth instead.[8]
New taxa
[edit]| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et sp. nov |
Valid |
Wu & Brinkman |
Latest Cretaceous |
A crocodylian, probably a crocodyloid. The type species is Albertosuchus knudsenii. |
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|
Sp. nov |
Valid |
Blanco et al. |
Late Cretaceous (Maastrichtian) |
A eusuchian crocodylomorph, a species of Allodaposuchus. |
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|
Sp. nov |
Valid |
Salas-Gismondi et al. |
Late Middle Miocene |
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|
Gen. et sp. nov |
Valid |
Zanno et al. |
Late Triassic (Carnian) |
An early member of Crocodylomorpha. The type species is Carnufex carolinensis. |
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|
Gen. et sp. nov |
Valid |
Lacerda, Schultz & Bertoni-Machado |
Triassic (late Ladinian/early Carnian) |
A non-crocodylomorph loricatan. The type species is Dagasuchus santacruzensis. |
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|
Gen. et sp. nov |
Valid |
Salas-Gismondi et al. |
Late Middle Miocene |
A caiman. The type species is Gnatusuchus pebasensis. |
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|
Gen. et sp. nov |
Valid |
Heckert et al. |
Late Triassic |
An aetosaur. The type species is Gorgetosuchus pekinensis. |
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|
Gen. et sp. nov |
Valid |
Sweetman, Pedreira-Segade & Vidovic |
Early Cretaceous (Barremian) |
A crocodyliform related to Bernissartia. The type species is Koumpiodontosuchus aprosdokiti. |
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|
Gen. et sp. nov |
Valid |
Salas-Gismondi et al. |
Late Middle Miocene |
A caiman. The type species is Kuttanacaiman iquitosensis. |
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|
Gen. et 2 sp. nov |
Valid |
Narváez et al. |
Late Cretaceous (late Campanian-early Maastrichtian) |
A eusuchian crocodyliform related to Allodaposuchus. The type species is Lohuecosuchus megadontos; genus also contains Lohuecosuchus mechinorum. |
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|
Gen. et sp. nov |
Valid |
Wilberg |
Middle Jurassic (Aalenian–Bajocian) |
A thalattosuchian crocodylomorph. The type species is Zoneait nargorum. |
Non-avian dinosaurs
[edit]Research
[edit]- A study of phylogenetic relationships of sauropod dinosaurs belonging to the family Diplodocidae and the taxonomic revision of the family is published by Tschopp, Mateus and Benson (2015); the authors propose to reestablish Brontosaurus as a genus distinct from Apatosaurus and to transfer Dinheirosaurus lourinhanensis to the genus Supersaurus.[18][19]
- Additional information on the anatomy of the holotype specimen of the rebbachisaurid sauropod Tataouinea hannibalis, including description of newly discovered fossil material of this specimen, and study of its phylogenetic relationships is published by Fanti et al. (2015).[20]
- A reevaluation of the estimated body mass of Dreadnoughtus schrani is published by Bates et al. (2015).[21]
- A study of phylogenetic relationships of ornithischian dinosaurs traditionally referred to as ‘basal ornithopods’ or ‘hypsilophodontids’ is published by Boyd (2015).[22]
- Carpenter & Lamanna (2015) reinterpret the ornithopod braincase from the Morrison Formation of the Carnegie Quarry at Dinosaur National Monument (Utah), formerly assigned to Uteodon aphanoecetes, as actually belonging to Dryosaurus cf. D. altus. The authors consider genera Uteodon and Cumnoria to be junior synonyms of Camptosaurus, and transfer the species Cumnoria prestwichii and Uteodon aphanoecetes to the genus Camptosaurus.[23]
New taxa
[edit]| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images | |
|---|---|---|---|---|---|---|---|---|---|
|
Gen. et comb. nov |
Valid |
Prieto-Márquez et al. |
Late Cretaceous (late Maastrichtian) |
A hadrosaurid; a new genus for "Saurolophus" morrisi Prieto-Márquez & Wagner (2013). |
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|
Gen. et sp. nov |
Valid |
Bell & Currie |
Late Cretaceous (late Campanian) |
A dromaeosaurid theropod. The type species is Boreonykus certekorum. |
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|
Gen. et sp. nov. |
Valid |
Novas et al. |
Late Jurassic (Tithonian) |
A dinosaur of uncertain phylogenetic placement. Originally classified as a theropod dinosaur (a basal member of Tetanurae), but subsequently argued to be an ornithischian.[27] The type species is Chilesaurus diegosuarezi. |
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|
Gen. et comb. nov |
Valid[29] |
Arbour & Currie |
Late Cretaceous (Cenomanian–Turonian) |
An ankylosaurine ankylosaurid; a new genus for "Crichtonsaurus" benxiensis Lü, Ji, Gao & Li (2007). |
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|
Gen. et sp. nov. |
Valid |
DePalma et al. |
Late Cretaceous (Maastrichtian) |
A member of Dromaeosauridae. The type species is Dakotaraptor steini. |
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| Eucnemesaurus entaxonis[31] |
Sp. nov |
Valid |
McPhee et al. |
Late Triassic |
Lower Elliot Formation |
A non-sauropod sauropodomorph, a species of Eucnemesaurus. |
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|
Gen. et comb. nov |
Valid |
Tschopp, Mateus & Benson |
Late Jurassic (Kimmeridgian) |
A diplodocid sauropod; a new genus for "Diplodocus" hayi Holland (1924). |
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| Gspsaurus[32] | Gen. et comb. nov | Dubious | Malkani | Late Cretaceous (Maastrichtian) | Pab Formation |  Pakistan
|
A titanosauriform sauropod. The type species is G. Pakistani. | ||
|
Gen. et comb. nov. |
Valid |
Upchurch, Mannion & Taylor |
Early Cretaceous (late Berriasian—Valanginian) |
A macronarian sauropod of uncertain phylogenetic placement; a new genus for "Pelorosaurus" becklesii Mantell (1852). |
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|
Gen. et comb. nov |
Valid |
Blows |
Early Cretaceous |
A new genus for "Polacanthus" rudgwickensis Blows (1996). |
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|
Gen. et sp. nov. |
Valid |
Han et al. |
Late Jurassic (Oxfordian) |
A basal ceratopsian, possibly a member of Chaoyangsauridae. The type species is Hualianceratops wucaiwanensis. |
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|
Gen. et sp. nov. |
Valid |
Lü et al. |
Late Cretaceous (Campanian-Maastrichtian) |
An oviraptorid theropod. The type species is Huanansaurus ganzhouensis. |
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|
Sp. nov |
Valid |
Verdú et al. |
Early Cretaceous (early Barremian) |
An ornithopod, a species of Iguanodon. |
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|
Gen. et sp. nov. |
Valid |
He et al. |
Late Cretaceous |
A leptoceratopsid ceratopsian. The type species is Ischioceratops zhuchengensis. |
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|
Gen. et sp. nov |
Valid |
Shibata & Azuma |
Early Cretaceous |
A non-hadrosaurid hadrosauroid. The type species is Koshisaurus katsuyama. |
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|
Gen. et sp. nov. |
Valid |
Leahey et al. |
Cretaceous (late Albian–(?)early Cenomanian) |
Allaru Mudstone |
An ankylosaur. The type species is Kunbarrasaurus ieversi. |
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|
Gen. et sp. nov. |
Disputed |
Alifanov & Saveliev |
Late Jurassic (Tithonian?) |
A dinosaur of uncertain phylogenetic placement; considered to be a theropod dinosaur related to Nqwebasaurus by the authors of its description,[41] but subsequently argued to be an ornithischian, possibly synonymous with Kulindadromeus.[42][43] The type species is Lepidocheirosaurus natatilis. |
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|
Gen. et sp. nov. |
Valid |
Nesbitt & Ezcurra |
Late Triassic |
A coelophysoid theropod. The type species is Lepidus praecisio. |
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|
Gen. et sp. nov. |
Valid |
Gasulla et al. |
Early Cretaceous (probably late Barremian) |
An ornithopod dinosaur, a basal member of Styracosterna. The type species is Morelladon beltrani. |
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|
Gen. et sp. nov |
Valid |
Zheng, Jin & Xu |
Late Cretaceous (early-middle Turonian—middle Campanian) |
A basal neoceratopsian. The type species is Mosaiceratops azumai. |
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|
Gen. et sp. nov |
Valid |
Xing et al. |
Middle Jurassic (Aalenian or Bajocian) |
A eusauropod sauropod. The type species is Nebulasaurus taito. |
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|
Gen. et sp. nov |
Valid |
Carballido et al. |
Early Cretaceous (Barremian) |
A brachiosaurid sauropod. The type species is Padillasaurus leivaensis. |
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|
Gen. et sp. nov |
Valid |
Freedman Fowler & Horner |
Late Cretaceous (Campanian) |
A hadrosaurid. The type species is Probrachylophosaurus bergei. |
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|
Gen. et sp. nov |
Valid |
McPhee et al. |
Early Jurassic (probably late Hettangian to Sinemurian) |
Upper Elliot Formation |
A basal sauropod. The type species is Pulanesaura eocollum. |
| |||
|
Gen. et sp. nov |
Valid |
Xing et al. |
Late Jurassic |
A mamenchisaurid sauropod. The type species is Qijianglong guokr. |
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|
Gen. et sp. nov |
Valid |
Brown & Henderson |
Late Cretaceous (probably middle Maastrichtian) |
A chasmosaurine ceratopsid. The type species is Regaliceratops peterhewsi. |
|
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|
Sp. nov |
Valid |
Jasinski |
Late Cretaceous (late Campanian) |
A member of Dromaeosauridae, a species of Saurornitholestes. |
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|
Gen. et sp. nov |
Valid |
Otero et al. |
Late Triassic or Early Jurassic |
A non-sauropod sauropodomorph. The type species is Sefapanosaurus zastronensis. |
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|
Gen. et sp. nov |
Valid |
Shibata et al. |
Early Cretaceous (Aptian) |
A basal member of Hadrosauroidea. The type species is Sirindhorna khoratensis. |
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|
Gen. et sp. nov |
Disputed |
Mori, Druckenmiller & Erickson |
Late Cretaceous (early Maastrichtian) |
A saurolophine hadrosaurid. The type species is Ugrunaaluk kuukpikensis. The genus Ugrunaaluk was considered to be a junior synonym of the genus Edmontosaurus by Takasaki et al. (2020).[57] |
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|
Gen. et sp. nov. |
Valid |
Evans & Ryan |
Late Cretaceous (Campanian) |
A centrosaurine ceratopsid. The type species is Wendiceratops pinhornensis. |
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|
Gen. et sp. nov |
Valid |
Xu et al. |
Jurassic (Callovian or Oxfordian) |
A scansoriopterygid theropod. The type species is Yi qi. |
|
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|
Gen. et sp. nov. |
Valid |
Lü & Brusatte |
Early Cretaceous |
A dromaeosaurid theropod. The type species is Zhenyuanlong suni. |
|
Birds
[edit]Research
[edit]- Balaur bondoc, originally interpreted as a member of Dromaeosauridae, is argued to be a basal member of Avialae by Cau, Brougham and Naish (2015).[61]
- Estimates of likely mass and flight capability of the extinct Chatham duck (Anas chathamica), as indicated by measurements of major bones, are published by Williams (2015).[62]
- New cranial material of the Oligo-Miocene relative of flamingos, Harrisonavis croizeti, is described by Torres et al. (2015).[63]
- New specimens of the Eocene relative of swifts, Scaniacypselus szarskii, are described by Mayr (2015).[64]
- A specimen of a stem-coliiform Masillacolius brevidactylus with preserved skull is described by Mayr (2015).[65]
- A study of the phylogenetic relationships of Psittacopes, Pumiliornis and Morsoravis, indicating their close relationship to the passerines and extinct family Zygodactylidae, is published by Mayr (2015); the author also introduced a new name, Parapasseres, for the clade containing Passeriformes and Zygodactylidae but not Psittacopes.[66]
- A study of preserved specimens of the Puerto Rican conure (previously considered to be an extinct subspecies of the Hispaniolan parakeet) is published by Olson (2015), who raises it to the rank of a separate species Psittacara maugei.[67]
- A Dwarf Emu was found in the Miocene of Alcoota in Central Australia.[68]
- Jones et al. (2015) describe a distal end of the left tibiotarsus of a member of the genus Caracara of uncertain specific assignment from the late Pleistocene of Argentina, estimated to be the largest member of the family Falconidae described thus far.[69]
- A description of the skeletal anatomy of two well-preserved specimens of the dodo is published by Claessens, Meijer and Hume (2015).[70]
New taxa
[edit]| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Sp. nov. |
Valid |
An Old World vulture, Accipitridae, related to the cinereous vulture. |
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|
Sp. nov |
Valid |
Forshaw |
Holocene |
A new scientific name for the Norfolk ground dove. The previous name used to refer to this species, Columba norfolciensis Latham (1801), was suppressed by ICZN in 2010.[73] |
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|
Sp. nov. |
Valid |
Zelenkov & Panteleyev |
Late Miocene |
An Anatidae. |
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|
Gen. et sp. nov. |
Valid |
Wang et al. |
A Hongshanornithidae. Type species A. meemannae. |
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|
Sp. nov. |
Valid |
Early Pliocene |
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|
Sp. nov. |
Valid |
Boev |
Early Pleistocene |
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|
Gen. et sp. nov. |
Valid |
Carvalho et al. |
Aptian |
Crato Member, Santana Formation |
A member of Euenantiornithes of uncertain phylogenetic placement.[78][79] |
| ||
|
Gen. et sp. nov. |
Valid |
Wang et al. |
Early Cretaceous |
A member of Enantiornithes. The type species is D. cuii. |
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|
Gen. et sp. nov. |
Valid |
Mayr |
Eocene |
A bird of uncertain phylogenetic placement. |
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|
Gen. et sp. nov |
Valid |
O’Connor et al. |
Early Cretaceous (Aptian) |
A member of Enantiornithes. The type species is Feitianius paradisi. |
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|
Gen. et sp. nov. |
Valid |
Zhao et al. |
Early Eocene |
A bird of uncertain phylogenetic placement, possibly a relative of trogons. Type species F. songi. |
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|
Gen. et sp. nov. |
Valid |
Bell & Chiappe |
Late Cretaceous (late Coniacian—early Campanian) |
The Smoky Hill Member of the Niobrara Chalk |
A member of Hesperornithiformes. The type species is Fumicollis hoffmani. |
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|
Sp. nov. |
Valid |
Zelenkov & Panteleyev |
Middle Turolian |
A snipe, Scolopacidae. |
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|
Gen. et sp. nov. |
Valid[86] |
De Pietri et al. |
Early Miocene (19–16 Ma) |
A member of Scolopaci of uncertain phylogenetic placement. The type species is Hakawai melvillei. |
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|
Gen. et sp. nov. |
Valid[88] |
Zelenkov & Averianov |
Early Cretaceous |
A member of Enantiornithes, probably a relative of Gobipteryx. Type species H. ponomarenkoi. |
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|
Gen. et comb. nov. |
Valid[90] |
Early Cretaceous |
A member of Enantiornithes of uncertain phylogenetic placement; a new genus for "Cathayornis" caudatus Hou (1997). |
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|
Sp. nov. |
Valid[92] |
Holocene |
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|
Gen. et sp. nov. |
Valid |
Wang, Wang & Hu |
Early Cretaceous |
A basal member of Ornithuromorpha. Type species J. zhangi. |
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|
Gen. et sp. nov. |
Valid |
Stucchi et al. |
Late Miocene |
A condor. Type species K. cardenasi. |
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|
Gen. et sp. nov. |
Valid[96] |
Stidham & Hilton |
Miocene (16.1–14.6 Ma) |
A member of Anatidae; a member or a relative of members of the subfamily Oxyurinae. The type species is Lavadytis pyrenae. |
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|
Gen. et sp. nov |
Valid |
Degrange et al. |
Late Pliocene |
A mesembriornithine phorusrhacid. Type species L. scagliai. |
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|
Gen. et sp. nov. |
Valid |
Latest Eocene or early Oligocene |
A member of Procellariiformes. Type species M. mirae. |
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|
Gen. et sp. nov |
Valid |
Zvonok, Zelenkov & Danilov |
Eocene (Bartonian) |
Possibly a member of Suliformes. The type species is Mangystania humilicristata. |
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|
Sp. nov |
Valid |
Zelenkov |
Miocene |
A grebe |
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|
Gen. et comb. nov |
Valid |
Zelenkov in Zelenkov & Kurochkin |
Late Miocene |
A member of Rallidae. A new genus for "Tertiariaporphyrula" lungi Kurochkin & Ganea (1972). |
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|
Gen. et sp. nov. |
Valid |
Worthy et al. |
Holocene |
A megapode. The type species is M. walterlinii. |
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|
Gen. et sp. nov. |
Valid |
Middle Eocene |
A Psittaciformes incertae sedis. Type species N. praeruptorum. |
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|
Gen. et sp. nov. |
Valid |
Elżanowski & Zelenkov |
Middle Miocene |
An Ardeidae. Type species N. robusta. |
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|
Gen. et sp. nov. |
Valid |
Late Oligocene |
A member of Charadriiformes related to the plains-wanderer. Type species O. milleri. |
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|
Sp. nov. |
Valid |
Agnolin & Chafrat |
Early Miocene |
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|
Gen. et sp. nov. |
Valid |
Hu, O’Connor & Zhou |
Early Cretaceous |
A member of Enantiornithes, a relative of Pengornis. Type species P. eurycaudatus. |
| |||
|
Gen. et sp. nov. |
Valid |
Agnolin & Chafrat |
Early Miocene |
A member of Phorusrhacidae. Type species P. terrificus. |
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|
Nom. nov |
Valid |
Zelenkov in Zelenkov & Kurochkin |
Late Miocene |
A member of Falconidae. A replacement name for Sushkinia Tugarinov (1935) (preoccupied). |
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|
Sp. nov. |
Valid |
Boev |
Early Pleistocene |
|||||
|
Sp. nov. |
Valid |
Tennyson, Cooper & Shepherd |
Holocene |
A member of Procellariidae, a species of gadfly petrel. |
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|
Gen. et sp. nov. |
Valid [111] |
Wang, Hu & Li |
Early Cretaceous |
A member of Enantiornithes. Type species P. dapingfangensis. |
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|
Sp. nov. |
Valid |
Alcover et al. |
Holocene |
|||||
|
Sp. nov. |
Valid |
Alcover et al. |
Holocene |
|||||
|
Sp. nov. |
Valid |
Alcover et al. |
Holocene |
|
||||
|
Sp. nov. |
Junior homonym |
Alcover et al. |
Holocene |
A member of Rallidae, a species of Rallus. The specific name is a junior homonym of Rallus minutus Pallas (1776), Rallus minutus Gmelin (1789) and Rallus minutus Forster (1844). Alcover et al. (2016) coined a replacement name Rallus nanus.[113] |
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|
Sp. nov. |
Valid |
Alcover et al. |
Holocene |
|||||
|
Sp. nov. |
Valid |
Stucchi, Varas-Malca & Urbina-Schmitt |
Miocene |
A member of Sulidae, a species of Ramphastosula. |
||||
|
Sp. nov. |
Valid |
Takano & Steadman |
Late Holocene |
A woodcock. |
||||
|
Gen. nov. et Sp. nov. |
Valid |
Mourer-Chauviré, Pickford, & Senut |
Middle Eocene |
A Paraortygidae, Galliformes. Type species S. klinghardtensis. |
||||
|
Gen. et sp. nov. |
Valid |
Watanabe & Matsuoka |
Pleistocene |
A duck. The type species is Shiriyanetta hasegawai. |
||||
|
Sp. nov. |
Valid |
Benson |
Late Miocene |
| ||||
|
Gen. et sp. nov. |
Valid |
Kaiser, Watanabe & Johns |
Late Oligocene |
A member of Plotopteridae. The type species is Stemec suntokum. |
| |||
|
Sp. nov. |
Valid |
Stucchi, Varas-Malca & Urbina-Schmitt |
Miocene |
A booby. |
||||
|
Sp. nov. |
Valid |
Stucchi, Varas-Malca & Urbina-Schmitt |
Miocene |
A booby. |
||||
|
Sp. nov. |
Valid |
Quaternary (probably late Pleistocene) |
||||||
|
Sp. nov |
Valid |
Pavia, Manegold, & Haarhoff |
Early Pliocene |
|||||
|
Gen. et sp. nov. |
Valid |
Hu et al. |
Early Cretaceous |
A member of Enantiornithes. Type species Yuanjiawaornis viriosus. |
Pterosaurs
[edit]Research
[edit]- A study on the terrestrial locomotion of non-pterodactyloid pterosaurs is published by Witton (2015).[121]
- A specimen of Rhamphorhynchus muensteri with preserved soft tissues or impressions of soft tissues, stomach contents and possibly a coprolite is described by Hone et al. (2015).[122]
- Eleutério et al. (2015) study the bone microstructure characterization of two pterosaurs belonging to the group Anhangueria.[123]
- The paleoenvironments of azhdarchid pterosaurs from the Late Cretaceous of Kazakhstan are studied by Averianov et al. (2015).[124]
- Martill et al. (2015) describe a jaw bone attributed to the mid-Cretaceous pterosaur species Alanqa saharica from the Kem Kem beds of Morocco.[125]
New taxa
[edit]| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
|---|---|---|---|---|---|---|---|---|
|
Gen. et comb. nov |
Valid |
Kellner |
Late Triassic (?Norian-Rhaetian) |
A non-pterodactyloid pterosaur of uncertain phylogenetic placement; a new genus for "Eudimorphodon" cromptonellus Jenkins et al. (2001). |
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|
Gen. et sp. nov |
Valid |
Kellner |
Late Triassic (late Norian) |
A non-pterodactyloid pterosaur of uncertain phylogenetic placement. The type species is Austriadraco dallavecchiai. |
||||
|
Gen. et sp. nov |
Disputed |
Headden & Campos |
Early Cretaceous |
Romualdo Formation |
The type species is Banguela oberlii. Originally described as a dsungaripterid; Pêgas, Costa & Kellner (2018) considered the genus Banguela to be a junior synonym of the genus Thalassodromeus, and transferred the species B. oberlii to the latter genus.[128] |
|||
|
Gen. et sp. nov |
Disputed |
Kellner |
Late Triassic (Alaunian) |
Originally interpreted as a relative of Campylognathoides. The type species is Bergamodactylus wildi. Dalla Vecchia (2018) considers B. wildi to be a junior synonym of Carniadactylus rosenfeldi.[129] |
| |||
|
Sp. nov |
Valid |
Myers |
Late Cretaceous (late Cenomanian) |
A member of Pterodactyloidea, probably a pteranodontoid; a species of Cimoliopterus. |
||||
|
Gen. et sp. nov |
Valid |
Cheng et al. |
Late Jurassic |
A non-pterodactyloid pterosaur. The type species is Daohugoupterus delicatus. |
||||
|
Gen. et sp. nov |
Valid |
Rodrigues et al. |
Early Cretaceous (Aptian) |
A member of Pteranodontoidea, possibly related to anhanguerians. The type species is Linlongopterus jennyae. |
||||
|
Gen. et sp. nov |
Valid |
Lü et al. |
Late Jurassic |
A rhamphorhynchid. The type species is Orientognathus chaoyngensis. |
||||
|
Sp. nov |
Valid |
O'Sullivan & Martill |
Late Jurassic |
Kimmeridge Clay Formation |
A species of Rhamphorhynchus. |
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