Paleobiota of the Green River Formation

The Green River Formation is a geological formation located in the Intermountain West of the United States, in the states of Colorado, Utah, and Wyoming. It comprises sediments deposited during the Early Eocene in a series of large freshwater lakes: Lake Gosiute, Lake Uinta, and Fossil Lake (the last containing Fossil Butte National Monument). It preserves a high diversity of freshwater fish, birds, reptiles, and mammals, with some sections of the formation (including Fossil Lake and the Parachute Creek member of Lake Uinta) qualifying as Konservat-Lagerstätten due to their extremely well-preserved fossils.

Cartilaginous fish

Family	Genus	Species	Member	Abundance	Notes	Image
†Asterotrygonidae	†Asterotrygon	†A. maloneyi Carvalho, Maisey & Grande, 2004	Fossil Lake		An Asterotrygonidae stingray of uncertain higher affinities.
†Heliobatidae	†Heliobatis	†H. radians Marsh, 1877	Fossil Lake		A Heliobatid stingray of uncertain higher affinities. H. radians synonymy Dasyatis species Haseman, 1912 Palaeodasybatis discus Fowler, 1947 Xiphotrygon acutidens Cope, 1879 Xyphotrygus species Romer, 1971

Bony fish

Primarily based on Grande (2001), with changes where necessary:

Acipenseriformes

Family	Genus	Species	Member	Abundance	Notes	Image
Polyodontidae	†Crossopholis	†C. magnicaudatus Cope, 1883	Fossil Lake		A paddlefish.

Lepisosteiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Lepisosteidae	Atractosteus	†A. atrox (Leidy, 1873)	Fossil Lake		An alligator gar relative. A. atrox is the largest Green River gar.
		†A. simplex (Leidy 1873)	Fossil Lake Lake Gosiute?		An alligator gar relative.
	†Cuneatus	†C. cuneatus (Cope, 1884)	Fossil Lake? Lake Gosiute Lake Uinta		A Cuneatini tribe gar.
		†C. wileyi Grande, 2010	Lake Uinta		A Cuneatini tribe gar.
	Lepisosteus	†L. bemisi Grande, 2010	Fossil Lake		A Lepisosteini tribe gar.
	†Masillosteus	†M. janeae Grande, 2010	Fossil Lake		A Cuneatini tribe gar.

Amiiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Amiidae	Amia	†A. pattersoni Grande & Bemis, 1998	Fossil Lake Lake Gosiute		An amiid bowfin.
	†Cyclurus	†C. gurleyi (Romer & Fryxell, 1928)	Fossil Lake	Only 8 specimens known	An amiid related to modern bowfins.

Hiodontiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Hiodontidae	Hiodon	†H. falcatus (Grande, 1979)	Fossil Lake		A hiodontid. First described as Eohiodon falcatus (1979).

Osteoglossiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Osteoglossidae	†Phareodus	†P. encaustus (Cope, 1871)	Fossil Lake Lake Gosiute		A phareodontine bonytongue.
		†P. testis (Leidy, 1873)	Fossil Lake

Ellimmichthyiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Armigatidae	†Diplomystus	†D. dentatus Cope, 1877	Fossil Lake Lake Gosiute Lake Uinta		An armigatid ellimmichthyiform.

Clupeiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Clupeidae	†Gosiutichthys	†G. parvus Grande, 1982	Lake Gosiute		A clupeine clupeid herring relative.
	†Knightia	†K. alta (Leidy, 1873)	Fossil Lake Lake Gosiute Lake Uinta		A pellonuline clupeid herring relative.
		†K. eocaena Jordan, 1907	Fossil Lake Lake Gosiute Lake Uinta	The most common articulated vertebrate fossil worldwide.	A pellonuline clupeid herring relative.

Gonorynchiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Gonorynchidae	†Notogoneus	†N. osculus Cope, 1885	Fossil Lake		A beaked sandfish. Last gonorynchid known from North America. Bottom feeding trace fossils are attributed to N. osculus.

Cypriniformes

Family	Genus	Species	Member	Abundance	Notes	Image
Catostomidae	†Amyzon	†A. gosiutensis Grande, Eastman, & Cavender, 1982	Lake Gosiute		An ictiobine sucker.	A. aggregatum, a related species

Siluriformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Hypsidoridae	†Hypsidoris	†H. farsonensis Lundberg & Case, 1970	Lake Gosiute		A hypsidorid catfish.
Ictaluridae	†Astephus	†A. antiquus (Leidy, 1873)	Fossil Lake Lake Gosiute Lake Uinta		An ictalurid catfish.

Salmoniformes

Family	Genus	Species	Member	Abundance	Notes	Image
Esocidae	Esox	†E. kronneri Grande, 1999	Fossil Lake	1 specimen	A pike or muskellunge species. Likely native to surrounding streams or rivers and not the lakes.

Percopsiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Percopsidae	†Amphiplaga	†A. brachyptera Cope, 1877	Fossil Lake		A trout-perch relative. Includes the jr synonym Erismatopterus endlichi (1877)
	†Erismatopterus	†E. levatus (Cope, 1870)	Lake Gosiute Lake Uinta		A trout-perch relative. First described as Cyprinodon levatus (1870) includes the jr synonym Erismatopterus rickseckeri (1871)

Carangiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Latidae	†Mioplosus	†M. labracoides Cope, 1877	Fossil Lake Lake Gosiute Lake Uinta		A lates perches relative.

Acanthuriformes

Family	Genus	Species	Member	Abundance	Notes	Image
Moronidae	†Cockerellites	†C. liops (Cope, 1877)	Fossil Lake	common	A planktivorous/insectivorous temperate bass. First described as Priscacara liops (1877) First moved to Cockerellites liops in 1923.
	†Hypsiprisca	†H. hypsacantha (Cope, 1886)	Fossil Lake	rare	A temperate bass. First named Priscacara hypsacantha 1886)
		†H. sp.	Fossil Lake	rare	An undescribed species of Hypsiprisca. A slenderer species with a more rounded tail fin outline.
	†Priscacara	†P. serrata Cope, 1877	Fossil Lake Lake Uinta		A temperate bass.
		†P. sp. a	Fossil Lake	1 specimen	An undescribed Priscacara Specimens have a rounder body and weak dorsal spines.
		†P. sp. b.	Fossil Lake	1 specimen	An undescribed Priscacara Possibly a new species or a variant specimen with 4 anal spines.

Incertae sedis

Family	Genus	Species	Member	Abundance	Notes	Image
†Asineopidae	†Asineops	†A. squamifrons Cope, 1870	Fossil Lake Lake Gosiute Lake Uinta		An acanthomorph of uncertain affinities Possibly a percopsiform, polymixiiform, or perciform.
		A. sp.	Fossil Lake		An undescribed Asineops species Restricted to Fossil Lake.

Amphibians

Frogs

Family	Genus	Species	Member	Abundance	Notes	Image
Pelodytidae	†Aerugoamnis	†A. paulus Henrici et al., 2013	Fossil Lake	2 specimens	A stem-relative of parsley frogs.

Salamanders

Family	Genus	Species	Member	Abundance	Notes	Image
Amphiumidae	†Paleoamphiuma	†P. tetradactylum Rieppel & Grande, 1998	Fossil Lake		A sirenid salamander.

Reptiles

Squamates

Family	Genus	Species	Member	Abundance	Notes	Image
Boidae	†Boavus	†B. idelmani Gilmore, 1938	Fossil Lake		A boa.
Dactyloidae	†Afairiguana	†A. avius Conrad et al., 2007	Fossil Lake		A stem-anole.
Shinisauridae	†Bahndwivici	†B. ammoskius Conrad, 2006	Fossil Lake		A relative of the Chinese crocodile lizard.
Tropidophiidae?	†Dunnophis	D. sp.	Fossil Lake		An undescribed possible dwarf boa.
Varanidae	†Saniwa	†S. ensidens Leidy, 1870	Fossil Lake		A stem-monitor lizard.
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A possible gecko or iguanid Preserved with soft tissue and skin, but bones dissolved.
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A relative of Shinisaurids Preserved with soft tissue and skin, but bones dissolved.

Crocodilians

Family	Genus	Species	Member	Abundance	Notes	Image
Alligatoridae	†Tsoabichi	†T. greenriverensis	Fossil Lake		A caiman.
Incertae sedis	†Borealosuchus	†B. wilsoni	Fossil Lake		A eusuchian of uncertain affinities.
	"Crocodylus"	†"Crocodylus" acer Cope, 1882	Lake Uinta (Manti beds)	1 specimen	A Crocodyloidea of unplaced affinity. Not a Crocodylus species, but not given a new genus as of 2024.

Turtles

Family	Genus	Species	Member	Abundance	Notes	Image
Baenidae	Chisternon	C. undatum	fossil Lake		A baenid.
Dermatemydidae	Baptemys	B. wyomingensis	Fossil Lake		A dermatemydid.
Geoemydidae	Echmatemys	†E. haydeni (Cope, 1873)		1 specimen	A geoemydid tortoise.
		†E. naomi (Leidy, 1871)	Fossil Lake	1 specimen	A geoemydid tortoise.
		†E. wyomingensis	Fossil Lake		A geoemydid tortoise.
Testudinidae	Hadrianus	Cf. †H. corsoni (Leidy, 1871)	Fossil Lake		A tortoise species. A single complete hatchling
Trionychidae	Axestemys	A. sp.	Fossil Lake		A softshell turtle.
	Oliveremys	O. uintaensis	Fossil Lake		A softshell turtle.
	Plastomenus	P. thomasii	Fossil Lake		A softshell turtle.

Birds

Lithornithiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Lithornithidae	†Calciavis	†C. grandei	Fossil Lake		A lithornithid.
	†Pseudocrypturus	†P. cercanaxius	Fossil Lake		A lithornithid.

Anseriformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Presbyornithidae	†Presbyornis	†P. pervetus	Fossil Lake		A presbyornithid anseriform.
Incertae sedis	†Paakniwatavis	†P. grandei	Fossil Lake		An anseriform of uncertain affinities.

Galliformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Gallinuloididae	†Gallinuloides	†G. wyomingensis	Fossil Lake		A gallinuloid galliform.

Coliiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Sandcoleidae	†Anneavis	†A. annae	Fossil Lake		A sandcoleid mousebird.

Leptosomiformes

Family	Genus	Species	Member	Abundance	Notes	Image
incertae sedis	†Plesiocathartes	P. major	Fossil Lake		A stem-cuckoo roller.
		†P. wyomingensis	Fossil Lake		A stem-cuckoo roller

Coraciiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Primobucconidae	†Primobucco	†P. mcgrewi	Fossil Lake		A primobucconid coraciiform.
Incertae sedis	†Paracoracias	†P. occidentalis			A coraciiform.

Piciformes

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Neanis	†N. schucherti	Fossil Lake		A possible piciform of uncertain affinities.
	†Pristineanis	†P. kistneri			A possible piciform of uncertain affinities.

Strisores

Family	Genus	Species	Member	Abundance	Notes	Image
Eocypselidae	†Eocypselus	†E. rowei	Fossil Lake		A eocypselid apodiform.
Incertae sedis	†Fluvioviridavis	†F. platyrhamphus	Fossil Lake		A basal strisore, possibly a stem-frogmouth.
	†Prefica	†P. nivea	Fossil Lake		A stem-oilbird.

Musophagiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Musophagidae?	†Foro	†F. panarium	Fossil Lake		A possible stem-turaco.

Mirandornithes

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Juncitarsus	†J. merkeli	Fossil Lake		A mirandornithalean, possibly an early phoenicopteriform.

Suliformes

Family	Genus	Species	Member	Abundance	Notes	Image
Fregatidae	†Limnofregata	L. azygosternon	Fossil Lake		An early frigatebird.
		L. hasegawai	Fossil Lake		An early frigatebird

Pelecaniformes

Family	Genus	Species	Member	Abundance	Notes	Image
Threskiornithidae	†Rhynchaeites	†R. sp.		1 specimen	An undescribed ibis, known from a largely complete specimen.	R. messelensis, a related species
	†Vadaravis	†V. brownae	Fossil Lake		Potentially a stem-ibis, but may be an indeterminate aequornithine.

Charadriiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Indeterminate	Indeterminate	Indeterminate	Lake Uinta	1 trace specimen	A shorebird Potentially the earliest record of recurvirostrids Known from a 3 footprint trackway.

Gruiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Messelornithidae	†Messelornis	†M. nearctica	Fossil Lake		A messelornithid.	Specimen of a related species

Eufalconimorphae

Family	Genus	Species	Member	Abundance	Notes	Image
†Halcyornithidae	†Cyrilavis	†C. colburnorum	Fossil Lake		A halcyornithid.
		†C. olsoni	Fossil Lake
†Messelasturidae	†Tynskya	†T. eocaena	Fossil Lake		A messelasturid.
†Psittacopedidae	†Eofringillirostrum	†E. boudreauxi	Fossil Lake		A psittacopedid.
†Quercypsittidae	†Avolatavis	†A. tenens	Fossil Lake		A quercypsittid.
†Zygodactylidae	†Eozygodactylus	†E. americanus	Fossil Lake		A zygodactylid.
	†Zygodactylus	†Z. grandei	Fossil Lake		A zygodactylid.

Neoaves incertae sedis

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Nahmavis	†N. grandei	Fossil Lake	1 specimen	Either a stem-gruiform or a stem-charadriiform.

Two other genera, "Eoeurypyga" (a stem-sunbittern) and "Wyomingcypselus" (an early apodiform) are mentioned only in a 2002 dissertation, and are presently nomina nuda.

Mammals

Partially based on Grande (1984). Aside from the few well-preserved mammals found in Fossil Lake, a majority of Green River mammals are based on isolated bones and teeth:

Metatheria

Family	Genus	Species	Member	Abundance	Notes	Image
†Herpetotheriidae	†Herpetotherium	†H. innominatum	Lake Gosiute Lake Uinta		An opossum-like basal metatherian Originally described as Peratherium innominatum.	H. huntii, a related species
		†H. knighti	Lake Gosiute		An opossum-like basal metatherian.

Cimolesta

Family	Genus	Species	Member	Abundance	Notes	Image
†Pantolestidae	†Palaeosinopa	†P. didelphoides	Fossil Lake		A semi-aquatic pantolestid.

Chiroptera

Family	Genus	Species	Member	Abundance	Notes	Image
†Icaronycteridae	†Icaronycteris	†I. gunnelli	Fossil Lake	2 specimens	An icaronycterid bat.
		†I. index	Fossil Lake	2 specimens	An icaronycterid bat.
†Onychonycteridae	†Onychonycteris	†O. finneyi	Fossil Lake	2 specimens	An onychonycterid bat.

Eulipotyphla

family	Genus	Species	Member	Abundance	Notes	Image
Nyctitheriidae	Nyctitherium	N. nitidum	Lake Gosiute		A nyctitheriid. Originally described as Talpavus nitidus.
		N. sp.	Lake Uinta

Pan-Carnivora

Family	Genus	Species	Member	Abundance	Notes	Image
Miacidae	Vulpavus	V. profectus	Lake Gosiute		A miacid.
Sinopidae	Sinopa	S. minor	Lake Uinta		A sinopid hyaenodont.	S. rapax, a related species
Viverravidae	Viverravus	V. minutus	Lake Uinta		A viverravid.
Incertae sedis	"Miacis"	"M." gracilis	Lake Uinta		A carnivoraform of uncertain affinities.

Pan-Perissodactyla

family	Genus	Species	Member	Abundance	Notes	Image
†Hyopsodontidae	†Hyopsodus	†H. minusculus	Lake Gosiute Lake Uinta		A hyopsodontid.
		†H. paulus (=H. vicarius)	Lake Uinta		A hyopsodontid. includes H. vicarius specimens.
		†H. wortmani	Fossil Lake		A hyopsodontid.
†Brontotheriidae	†Lambdotherium	†L. popoagicum	Fossil Lake		A small-sized basal brontothere.
Equidae	†Protorohippus	†Aff. P. venticolus	Fossil Lake	2 specimens	An equid closely affiliated with P. venticolus.
Indeterminate	Indeterminate	Indeterminate	Fossil Lake	1 specimen	A juvenile tapiromorph, the largest mammal known from the formation. Initially described as Heptodon calciculus.

Apatotheria

Family	Genus	Species	Member	Abundance	Notes	Image
Apatemyidae	Apatemys	A. chardini Jepsen, 1930	Fossil Lake	1 specimen	An apatemyid.

Rodentia

Family	Genus	Species	Member	Abundance	Notes	Image
Ischyromyidae	Paramys	P. sp.	Lake Uinta		A ischyromyid rodent.	P. delicatus
	Thisbemys	T. sp.	Lake Gosiute		An undescribed ischyromyid rodent.	T. plicatus
Sciuravidae	Sciuravus	S. eucristadens (Burke, 1937)	Lake Uinta (Douglas Creek Member)		A sciuravid rodent. Originally described as Viverravus eucristadens (1937).
		S. sp.	Lake Uinta		An undescribed sciuravid.

Primatomorpha

Family	Genus	Species	Member	Abundance	Notes	Image
Microsyopidae	Microsyops	M. elegans	Lake Gosiute		A microsyopid stem-primate.	M. cardiorestes, a related species
	Uintasorex	U. parvulus	Lake Gosiute		A microsyopid stem-primate.
Notharctidae	Notharctus	N. sp.	Lake Gosiute		A notharctid adapiform.	N. tenebrosus
	Smilodectes	S. gracilis	Lake Uinta		A notharctid adapiform. Formerly included in Notharctus matthewi.
Omomyidae	Omomys	O. carteri	Lake Uinta		An omomyid primate. Includes O. pucillus
	Tetonius	T. sp.	Lake Uinta		An omomyid primate.	T. homunculus
	Washakius	W. insignis	Lake Gosiute		An omomyid primate.

Arthropoda

Arachnida

Acariformes

Family	Genus	Species	Member	Abundance	Notes	Image
Cheyletidae	Indeterminate	Indeterminate	Lake Unita	1 specimen	A Chelytidae mite. Found in thin sectioning of rock samples.

Araneae

Family	Genus	Species	Member	Abundance	Notes	Image
Araneidae	Araneus	A. sp		6 specimens	Possible orbweb-spiders, known from partial fossils First placed as Epeira sp.
Hersiliidae	Hersiliola (sensu lato)	H. sp.	Lake Uinta (Parachute Creek Member)	1 male	A tree trunk spider.
Selenopidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	1 specimen	A female or juvenile selenopid spider.
Thomisidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	2 males, 1 juv female	Crab spiders.
Uloboridae	Miagrammopes	M. sp.	Lake Uinta (Parachute Creek Member)	1 female, 2 males	A uloborid spider.
	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)		A uloborid spider.
Unidentified	†Araneaovoius (Scudder, 1878)	†A. columbiae		1 specimen	An orb-web spider egg sack. First described as "Aranea" columbiae (1878)

Parasitiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Ixodidae	Ixodes	†"I." tertiarius Scudder, 1885	Lake Uinta (Laney Member)	1 specimen	A punitive ixodid hard tick. Type specimen declared lost species considered nomen dubium by Dunlop (2011)

Scorpiones

Family	Genus	Species	Member	Abundance	Notes	Image
Buthidae	†Uintascorpio	†U. halandrasi Perry, 1995	Lake Uinta (Parachute Creek Member)	1 specimen	A buthid scorpion.

Crustacea

Branchiopoda

Family	Genus	Species	Member	Abundance	Notes	Image
Cyclestheriidae	†Cyclestherioides	†C. wyomingensis	Lake Gosiute (Laney)		A cyclestheriid clam shrimp.
Lynceidae	†Prolynceus	†P. laneyensis	Lake Gosiute (Laney)		A clam shrimp.

Decapods

Family	Genus	Species	Member	Abundance	Notes	Image
Palaemonidae	†Bechleja	†B. rostrata	Fossil Lake		A palaemonid shrimp.
Cambaridae	Procambarus	†P. primaevus	Fossil Lake		A cambarid crayfish.

Ostracoda

Family	Genus	Species	Member	Abundance	Notes	Image
Candonidae	Candona	C. artesensis Swain, 1964	Fossil Lake Lake Unita		A candonid ostracod.
		C. rangliensis Swain, 1964	Fossil Lake Lake Unita		A candonid ostracod.
		C. whitei Scott & Smith, 1951	Fossil Lake Lake Gosiute Lake Unita		A candonid ostracod.
	Tuberocypris	T. centronotus Swain, 1964	Lake Unita		A candonid ostracod.
†Cyprideidae	†Cypridea	†C. bisulcata Swain, 1949	Lake Unita		A cyprideid ostracod.
Cyprididae	†Advenocypris	†A. umbonata Swain, 1964	Lake unita		A cypridid ostracod.
	Hemicyprinotus	†H. watsonensis Swain, 1964	Lake unita		A cypridid ostracod. One described subspecies H. watsonensis emaciata
	Heterocypris	†H. riflensis Swain, 1964	Lake unita		A cypridid ostracod.
		†H. whiteriverensis Swain, 1964	Lake unita		A cypridid ostracod.
	Potamocypris	†P. williamsi Swain, 1964	Fossil Lake Lake Unita		A cypridid ostracod.
	†Procyprois	†P. ravenridgensis]] Swain, 1964	Lake Unita		A cypridid ostracod. Male and female specimens show shell sexual dimorphism
	Pseudoeucypris	†P. pagei (Swain) Swain, 1956	Lake Unita		A cypridid ostracod. First described as Candona pagei (1949) Moved from Cypris pagei (1956)
	Scottia	†S. subquadrata Swain, 1964	Lake Unita		A cypridid ostracod.
	Stenocypris	Cf. S. sinuata G.W. Müller, 1898	Lake Unita		A cypridid ostracod.
Limnocytheridae	Limnocythere	L. sp.	Lake Unita		Indistinct limnocytherid fossils.
	Metacypris	†M. paracordata Swain, 1964	Lake Unita		A limnocytherid ostracod.
Notodromadidae	Cyprois	†C. ephraimensis Swain, 1964	Fossil Lake Lake Gosiute Lake Unita		A notodromadid ostracod.

Myriapoda

Family	Genus	Species	Member	Abundance	Notes	Image
Parajulidae	Julus?	"J." telluster Scudder, 1878		1 specimen	A punitive parajulidae millapede Considered nomen dubium by Hoffman (1969).

Insecta

Coleoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Anthribidae	†Pseudochirotenon	†P. eocaenicus Legalov, 2018	Lake Unita	1 specimen	An Ecelonerini tribe anthribine fungus weevil.
Brentidae	Perapion	†P. rasnitsyni Legalov, 2018	Lake Uinta		An Aplemonini tribe apionine primitive weevil.
Chrysomelidae	†Palaeatalasis	†P. monrosi Legalov, 2021	Lake Unita	1 specimen	A Megamerini tribe sagrine leaf beetle.
	†Pulchritudo	†P. attenboroughi Krell & Vitali, 2021	Lake Uinta	1 specimen	A sagrine leaf beetle
Curculionidae	†Archaeoheilus	†A. deleticius (Scudder, 1893)	Lake Uinta		A Molytini tribe molytine weevil. First named as Pachylobius deleticius (1893) Moved from Hylobius deleticius (2015)
		†A. ovalis Legalov, 2018	Lake Uinta		A Molytini tribe molytine weevil.
		†A. packardii (Scudder, 1893)	Lake Gosiute		A Molytini tribe molytine weevil. First named as Hylobius packardii (1893).
		†A. provectus (Scudder, 1876)	Lake Gosiute		A Molytini tribe molytine weevil. First named as Hylobius provectus (1876).
		†A. scudderi Legalov, 2018	Lake Uinta		A Molytini tribe molytine weevil.
	Eudiagogus	†E. vossi Legalov, 2018	Lake Uinta		A Eudiagogini tribe entimine weevil.
	Extinctocentrinus	†E. brevirostris Legalov, 2018	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	Geraeus	†G. anvilis Legalov, 2018	Lake Uinta		An Apostasimerini tribe conoderine weevil.
		†G. fossilis Legalov, 2018	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	†Lithogeraeus	†L. greenriverensis Legalov, 2018	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	Nicentrus	†N. curvirostris Legalov, 2018	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	†Palaeomallerus	†P. longirostris Legalov, 2018	Lake Uinta		A Palaeomallerini tribe conoderine weevil.
	†Primocentron	†P. wickhami (Scudder, 1876)	Lake Uinta		An Eustylini tribe entimine weevil. First described as Otiorhynchus perditus (1876)
	†Pseudophaops	†P. wickhami Legalov, 2018	Lake Uinta		A Tropiphorini tribe entimine weevil.
Elateridae	Cardiophorus	†C. exhumatus Cockerell, 1925	Lake Unita	1 specimen	A cardiophorine click beetle.
Mordellidae	†Petrimordella	†P. priscula (Cockerell, 1924)		1 specimen	A tumbling flower beetle morphospecies. First named as Mordella priscula (1925).
		†P. rasnitsyni (Odnosum & Perkovsky, 2016)		1 specimen	A tumbling flower beetle morphospecies. First named as Isotrilophus rasnitsyni (2016).
Scarabaeidae	Aegialia	†A. rupta Scudder, 1890			An aegialiine scaraboid beetle.
	Phyllophaga	†P. avus (Cockerell), Wickham, 1927	Lake Unita	1 specimen	A melolonthine scaraboid beetle. First described as Melolonthites avus (1921)

Dictyoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Alienopteridae	†Chimaeroblattina	†C. brevipes Barna, 2018	Lake Unita		An alienopterid Dictyopteran.
	†Grant	†G. viridifluvius Aristov, 2018	Lake Unita		An alienopterid Dictyopteran.
Blattidae	Periplaneta	†P.? smithiae Barna, Zelagin & Šmídová, 2023	Lake Uinta (Parachute Creek)	1 specimen	A possible periplanetean blattid cockroach.
Blaberidae	Diploptera	†D. gemini Barna, 2016	Lake Unita		A Diplopterine beetle cockroache.
		†D. savba Šmídová, 2016	Lake Unita		A Diplopterine beetle cockroache.
		†D. vladimir Vršanský, 2016	Lake Unita		A Diplopterine beetle cockroache.

Diptera

Family	Genus	Species	Member	Abundance	Notes	Image
Bibionidae	Plecia	†P. akerionana Fitzgerald, 1999	Lake Unita (Parachute Creek Member)		A bibonid marchfly. The smallest species from the formation.
		†P. dejecta Scudder, 1890			A bibonid marchfly.
		†P. pealei Scudder, 1890	Fossil Lake (Fossil Butte Member)	Most common insect in Fossil Lake	A bibonid marchfly.
		†P. rhodopterina Cockerell, 1925	Lake Unita		A bibonid marchfly.
		†P. woodruffi Cockerell, 1916	Lake Unita		A bibonid marchfly.
Cylindrotomidae	†Cyttaromyia	†C. fenestrata Scudder, 1877	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
		†C. freiwaldi Cockerell, 1925	Lake Unita	2 specimens	A cylindrotomine long bodied cranefly species.
		†C. fuscula Cockerell, 1925	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
		†C. gelhausi Kania-Kłosok et al., 2021	Lake Unita	3 specimens	A cylindrotomine long bodied cranefly species.
		†C. obdurescens Cockerell, 1925	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
		†C. reclusa Cockerell, 1924	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
Hippoboscidae	†Eornithoica	†E. grimaldii Nel, Garrouste & Engel, 2023	Fossil Lake (Fossil Butte Member)	1 specimen	A Hippoboscoidea fly Pupipara.
Chironomidae	Undescribed	Undescribed	Lake Unita	Numerous	undescribed midge larvae.
Mythicomyiidae	Reissa	†R. kohlsi			A mythicomyiine mythicomyiid fly
Oestridae	Cuterebra	†C. ascarides (Scudder, 1877)	Lake Unita	Numerous	A bot fly species.
Stratiomyidae	Odontomyia	Undescribed	Lake Unita	Numerous	undescribed larvae.
Mycetophilidae	Undescribed	Undescribed	Lake Unita	Numerous	undescribed larvae.
Syrphidae	Undescribed	Undescribed	Lake Unita	Numerous	undescribed larvae.
Tabanidae	Undescribed	Undescribed	Lake Unita	Numerous	undescribed larvae.
Tipulidae	Tipula	†T. americana	Lake Unita		A tipuline crane fly.

Hemiptera

Family	Genus	Species	Member	Abundance	Notes	Image
Cicadellidae	Coelidia	†A. wyomingensis Scudder, 1890	Fossil Lake (Fossil Butte Member)	1 specimen	A coelidiine leafhopper.
	Thamnotettix	†T. packardi Cockerell, 1925			An Athysanini tribe deltocephaline planthopper.
Gerridae	†Telmatrechus	†T. parallelus Scudder, 1890	Fossil Lake (Fossil Butte Member) Lake Uinta (Laney Member)		A water strider.
Reduviidae	†Aphelicophontes	†A. danjuddi Swanson et al., 2021
Incertae sedis	†Eoliarus	†E. lutensis (Scudder) Cockerell, 1925			A fulgoroid leafhopper. Frist described as Oliarus lutensis (1890)
		†E. quadristictus Cockerell, 1925			A fulgoroid leafhopper.

Hymenoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Bethylidae	Eupsenella	E. sp.			An old world Bethylidae wasp.
Formicidae	†Archimyrmex	†A. rostratus Cockerell, 1923	Lake Unita	Numerous	A myrmeciine ant.
	†Camponotites	†C. vetus (Scudder, 1877)	Green River Formation?	1 specimen	A formicine ant of uncertain affiliation. First described as Camponotus vetus (1877) The type locality was not specified by Scudder, and may Be G.R.F. or White River Formation.
	Dolichoderus	†D. kohlsi Dlussky & Rasnitsyn, 2003	Lake Uinta	Numerous	A Dolichoderus species ant.
	†Eoformica	†E. globularis Dlussky & Rasnitsyn, 2003	Lake Unita	2 specimens	An ant of unidentified affiliation.
		†E. magna Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	An ant of unidentified affiliation.
		†E. pinguis (Scudder, 1877)	Lake Gosiute Lake Uinta	Numerous	An ant of unidentified affiliation. Moved from Liometopum pingue (1877) Includes the jr synonyms Lasius terreus (1878) and Eoformica eocenica (1921)
		Indeterminate	Lake Unita	2 specimen	Two partial specimens that may be referable to Eoformica.
	†Kohlsimyrma	†K. gracilis Dlussky & Rasnitsyn, 2003	Lake Unita	2 specimens	An ant of uncertain affiliation.
		†K. laticeps Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	An ant of uncertain affiliation.
		†K. longiceps Dlussky & Rasnitsyn, 2003	Lake Unita	3 specimens	An ant of uncertain affiliation.
	†Mianeuretus	†M. eocenicus Dlussky & Rasnitsyn, 2003	Lake Unita	2 specimens	An aneuretine ant.
		Indeterminate	Lake Unita	1 specimen	A partial specimen that may belong to Mianeuretus.
	†Myrmecites	†M. rotundiceps Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A myrmicine ant of uncertain affiliation.
	Pachycondyla	†P. labandeirai Dlussky & Rasnitsyn, 2003	Lake Gosiute	1 specimen	A Pachycondyla species ant.
	†Ponerites	†P. coloradensis Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
		†P. eocenicus Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
		†P. hypoponeroides Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
	†Proiridomyrmex	†P. vetulus Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A dolichoderine of uncertain affinity.
		Indeterminate	Lake Unita	1 specimen	A partial specimen that may belong to Proiridomyrmex.
	†Solenopsites	†S. minutus Dlussky & Rasnitsyn, 2003	Lake Unita	1 specimen	A Solenopsis like ant of uncertain affiliation.
	†Titanomyrma	†T. lubei Archibald et al., 2011	Fossil Lake (Angelo Member)	3 specimens	A titan ant species.
	Indeterminate	Indeterminate	Lake Unita	1 specimen	A partial ant specimen that may belong to Formica.
	Indeterminate	Indeterminate	Lake Unita	3 specimens	Partial ant specimens of unidentified subfamily placement.
Ichneumonidae	†Carinibus	†C. molestus Spasojevic et al., 2018	Lake Unita		A darwin wasp of uncertain subfamily placement.
	†Ichninsum	†I. appendicrassum	Lake Unita		A pimpline darwin wasp.
	Mesoclistus?	†M.? yamataroti	Lake Unita		A possible Coleocentrini tribe acaenitine darwin wasp.
	Scambus?	†S.? mandibularis	Lake Unita		A possible Ephialtini tribe pimpline darwin wasp.
		†S.? parachuti	Lake Unita		A possible Ephialtini tribe pimpline darwin wasp.
	†Trymectus	†T. amasidis (Cockerell & LeVeque, 1931)	lake Unita		A darwin wasp of uncertain subfamily placement. First described as Tryphon amasidis (1931)
Siricidae	†Eoteredon	†E. lacoi Archibald, Aase & Nel, 2021	Fossil Lake	1 specimen	A siricid Horntail wood-wasp.
Tenthredinidae	Eutomostethus	†E. karimae Nel, Niu & Wei, 2022	Lake Unita		A blennocampine common sawfly.
Xyelidae	†Proxyelia	†P. pankowskii Jouault, Aase & Nel, 2021	Fossil Lake	1 specimen	A xyelid sawfly

Megaloptera

Family	Genus	Species	Member	Abundance	Notes	Image
Sialidae	Indeterminate	Indeterminate		1 specimen	A probable alderfly larvae

Neuroptera

Family	Genus	Species	Member	Abundance	Notes	Image
Berothidae	†Xenoberotha	†X. angustialata Makarkin, 2017	Lake Unita (Parachute Creek Member)	1 specimen	A berothine beaded lacewing.
Hemerobiidae	Megalomus?	†M.? coloradensis Makarkin, 2024	Lake Unita		A possible Megalomus brown lacewing.
	†Pseudonotherobius	†P. kohlsi Makarkin, 2024	Lake Unita		A brown lacewing.
Mantispidae	†Protonolima	†P. mantispinoformis Makarkin, 2019	Lake Unita	1 specimen	A calomantispine mantidfly.
Myrmeleontidae	†Epignopholeon	†E. sophiae Makarkin, 2017	Lake Unita (Parachute Creek Member)	1 specimen	A probable Gnopholeontini tribe myrmeleontine antlion.
Osmylidae	†Osmylidia	†O. taliae Makarkin, Archibald, & Mathewes, 2021	Lake Unita	1 specimen	A protosmyline Osmylid lance lacewing

Raphidioptera

Family	Genus	Species	Member	Abundance	Notes	Image
Raphidiidae	†Archiinocellia	†A. protomaculata (Engel, 2011)	Lake Unita (Parachute Creek Member)	ca. 18 specimens	A raphidiid snakefly. Moved from Agulla protomaculata (2011)

Odonata

Primarily based on Bechly et al (2020):

Family	Genus	Species	Member	Abundance	Notes	Image
Amphipterygidae	†Protamphipteryx	P. basalis Cockerell, 1921	Fossil Lake		An amphipterygine amphipterygid damselfly.
†Carleidae	†Carlea	C. eocenica Bechly et al., 2020	Fossil Lake	1 specimen	A carleid damselfly.
Cordulibellulidae	†Cordulibellula	C. inopinata Sroka, Howells & Nel, 2023	Lake Uinta (Parachute Creek)		A cordulibellulid dragonfly the last member of an ancient lineage.
Dysagrionidae	†Dysagrion	D. fredericii Scudder, 1878	Lake Gosiute (Laney Member)	16 specimens	A dysagrionine dysagrionid damselfly.
		D. integrum Bechly et al., 2020	Fossil Lake (Fossil Butte Member)	1 specimen
		D. lakesii Scudder, 1890	Lake Gosiute (Laney Member)	1 specimen
		D. packardii Scudder, 1885	Lake Gosiute (Laney Member)	1 specimen
	†Petrolestes	†P. hendersoni Cockerell, 1927	Lake Uinta (Parachute Creek Member)	1 specimen	A petrolestine dysagrionid damselfly.
		†P. inexpectatus Nel, 2024	Lake Uinta	1 specimen	A petrolestine dysagrionid damselfly.
	†Tenebragrion	†T. tynskyi Nel, Palazzo & Aase, 2023	Fossil Lake		A dysagrionid damselfly.
		†T. shermani Bechly et al., 2020	Fossil Lake		A dysagrionid damselfly.
Calopterygidae	†Eocalopteryx	E. atavina Cockerell, 1920	Lake Gosiute	1 specimen	A calopterygid damsefly.
Euphaeidae	†Eodysphaea	E. magnifica Bechly et al., 2020	Lake Uinta (Parachute Creek)		A euphaeid damselfly.
	†Epallagites	E. avus Cockerell, 1925	Lake Unita
	†Labandeiraia	L. americaborealis Petrulevičius et al., 2007	Lake Uinta (Parachute Creek)	1 specimen
		L. riveri Bechly et al., 2020	Fossil Lake	1 specimen
	†Litheuphaea	L. coloradensis Petrulevičius et al., 2007	Lake Uinta (Parachute Creek)	1 specimen
†Eolestidae	†Eolestes	E. syntheticus Cockerell, 1940	Lake Unita		An eolestid damselfly.
Megapodagrionidae	†Viridiflumineagrion	V. aasei Nel, 2022	Lake Uinta (Parachute Creek Member)	1 specimen	A "megapodagrionid" damselfly.
Thaumatoneuridae	†Gusagrion	†G. coloratum Nel, 2024	Lake Uinta	1 specimen	An eodysagrionine thaumatoneurid damselfly.
	†Oreodysagrion	O. tenebris Bechly et al., 2020	Fossil Lake		A thaumatoneurid damselfly.
	†Tynskysagrion	T. brookeae Bechly et al., 2020	Fossil Lake		A thaumatoneurid damselfly.
†Urolibellulidae	†Kohlslibellula	K. lini Nel, 2020	Lake Uinta (Parachute Creek)		A urolibellulid dragonfly
	†Urolibellula	U. eocaenica Zeiri et al., 2015	Lake Uinta (Parachute Creek)		A urolibellulid dragonfly
Zacallitidae	†Zacallites	Z. balli Cockerell, 1928			A zacallitid damselfly.
		Z. cockerelli Bechly et al., 2020	Fossil Lake
incertae sedis	†Eopodagrion	"E." abortivum (Scudder, 1878)	Lake Gosiute (Laney Member)	1 specimen	A damselfly of uncertain affinities. Deemed nomen dubium by Nel (2022).
		E. scudderi Cockerell, 1921	Lake Gosiute (Laney Member)	3 specimens	A damselfly of uncertain affinities.

Orthoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Tettigoniidae	Arethaea	†A. solterae Heads, Thomas & Hedlund, 2023	Lake Uinta (Parachute Creek)	1 specimen	A roundhead katydid.

Trichoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Limnephilidae	Limnephilus	†L. eocenicus Cockerell, 1921	Lake Unita	1 specimen	A northern caddisfly species.
Incertae sedis	†Litholimnephilops	†L. yinani Robinson, Thomas & Heads, 2018	Lake Unita		A caddis fly of uncertain affiliations.

Mollusks

Based on Grande (1984):

Bivalvia

Family	Genus	Species	Member	Abundance	Notes	Image
Sphaeriidae	Sphaerium	S. sp.	Lake Gosiute		A fingernail clam.	S. rhomboideum, a modern species
Unionidae	Plesielliptio	P. priscus			A unionid river mussel.
		P. sp. "A"

Gastropoda

Family	Genus	Species	Member	Abundance	Notes	Image
Bulimulidae	Oreoconus	O. sp.	?		An undescribed bulimulid land snail.
Gastrocoptidae	Gastrocopta	G. sp.			An undescribed gastrocoptid land snail.
Hydrobiidae	Hydrobia	†H. aff. utahensis C. A. White, 1876	Lake Gosiute (Laney, Tipton Shale, Wilkins Peak)		An undescribed mud snail. Affiliated to Hydrobia utahensis
		H. sp.	Lake Gosiute (Tipton Shale)		An undescribed mud snail.
Lymnaeidae	Lymnaea	L. similis Meek, 1860	Lake Gosiute (Laney)		A pond snail.
		L. sp.	Lake Gosiute (Tipton Shale)		A undescribed pond snail.
	†Pleurolimnaea	P. tenuicosta (Meek & Hayden, 1856)	Lake Gosiute (Tipton Shale?)		A pond snail. First named Limnaea tenuicosta (1856)
Planorbidae	Biomphalaria	B. aequalis (C. A. White, 1880)	Lake Gosiute (Laney, Tipton Shale)		A ramshorn snail.
		B. storchi (L. S. Russell, 1931)	Lake Gosiute (Laney?, Tipton Shale)		A ramshorn snail.
	†Headonia	†H. pseudoammonius (Schlotheim, 1820)	Lake Gosiute (Laney, Tipton Shale)		A ramshorn snail First reported as Biomphalaria pseudoammonius (1820)
	?Drepanotrema	?D. sp.	Lake Gosiute (Laney)		An undescribed ramshorn snail.
	Gyraulus	G. militaris (C.A. White, 1880)	Lake Gosiute (Tipton Shale)		A ramshorn snail.
	Omalodiscus	†O. cirrus (C. A. White, 1879)	Lake Gosiute (Tipton Shale)		A ramshorn snail.
Physidae	Physa	†?P. longiuscula Meek & Hayden, 1856	Lake Gosiute (Tipton Shale)		An bladder snail.
		P. sp.	Lake Gosiute (Laney)		An undescribed bladder snail.
	Physella	†P. bridgerensis (Meek, 1873)	Lake Gosiute (Laney, Tipton Shale)		A bladder snail. First named Physa bridgerensis (1873)
		†P. pleromatis (C. A. White, 1877)	Lake Gosiute (Tipton Shale)		A bladder snail. First named Physa pleromatis (1877)
Pleuroceridae	Elimia	†E. tenera (J. Hall, 1845)	Lake Gosiute (Laney, Tipton Shale, Wilkins Peak)		A pleurocerid snail. the most common snail in the formation The source of the famous "Turritella agate".
Urocoptidae	Holospira	H. sp.			An undescribed urocoptid land snail.
Valvatidae	Valvata	†V. subumbilicata (Meek & Hayden, 1856)	Lake Gosiute (Laney, Tipton Shale)		A valve snail.
		Cf. †V. filosa Whiteaves, 1885			An undescribed valve snail. Similar to Valvata filosa
Viviparidae	Viviparus	†V. meeki Wenz, 1930	Lake Gosiute (Laney, Tipton Shale)		A river snail. First named Paludina trochiformis (1856 non-von Zieten, 1830)
		†V. paludinaeformis (J. Hall, 1845)	Lake Gosiute (Tipton Shale)		A river snail.

Fungi

Genus	Species	Member	Abundance	Notes	Image
†Caenomyces	C. eucalypti Knowlton, 1923	Lake Unita	1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a Syzygioides americana leaf. Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
	†C. planerae R.W. Brown, 1929		1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a Cedrelospermum nervosum leaf. Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
	†C. sapindicola Knowlton, 1923		1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on the Toxicodendron winchesteri holotype leaf. Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
†Eoglobella	†E. longipes Bradley, 1931	Lake Unita	1 specimen	A Staurosporaen fungal spore of uncertain affiliation.
Entophlyctis	†E. willoughbyi Bradley, 1967	Lake Gosiute (Wilkins Peak Member)		An aquatic chytridialean fungi.
Pleotrachelus	†P. askaulos Bradley, 1967	Lake Gosiute (Wilkins Peak Member)		An aquatic chytridialean fungi.
†Sphaerites	†S. myricae (Lesquereux) Meschinelli, 1892	Lake Gosiute	1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a "Rhus" nigricans leaf. First described as Sphaeria myricae (1878) Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.

Unicellular microbiota

Cyanobacteria

Family	Genus	Species	Member	Abundance	Notes	Image
Chroococcaceae	Chroococcus	?Cf. C. dispersus (Von Keissler) Lemmerman	Lake Unita		An aquatic chroococcacean cyanobacteria.
		Cf. C. westii (W. West) P. Boye, 1929	Lake Unita		An aquatic chroococcacean cyanobacteria.
Coleofasciculaceae	Stigonema	†S. anchistina Bradley, 1970	Lake Gosiute (Wilkins Peak Member)		An aquatic coleofasciculacean cyanobacteria.
	Symploca	†S. hedraia Bradley, 1970	Lake Gosiute (Wilkins Peak Member)		An aquatic coleofasciculacean cyanobacteria.
Gomontiellaceae	Crinalium	?Cf. C. endophyticum Crow	Lake Unita		An aquatic gomontiellaceous cyanobacteria.
		C. sp.	Lake Unita		An aquatic gomontiellaceous cyanobacteria. First identified as Spirulina sp. (1916)
Hapalosiphonaceae	Hapalosiphon	H. confervaceus Borzi	Lake Unita		An aquatic hapalosiphonaceous cyanobacteria.

Algae

Family	Genus	Species	Member	Abundance	Notes	Image
Chaetophoraceae	Stigeoclonium?	Cf. S.? lubricum (Dillwyn) Kützing	Lake Unita	1 specimen	A possible chaetophoraceous chlorophyte green algae.
Chlorellaceae	†Chlorellopsis	†C. coloniata Reis, 1923			A unicellular stromatolite forming green algae.
	†Confervites	†C. mantiensis Bradley, 1929	Lake Uinta (Manti beds)		A filamentous stromatolite forming green algae.
Hydrodictyaceae	Tetraedron	Cf. T. regulare var. torsum (Turner) Brunnthaler	Lake Unita	1 specimen	A hydrodictyaceous chlorophyte green algae.
Microsporaceae	Microspora	Cf. M. pachyderma (Wille) Lagerheim	Lake Unita	1 specimen	A sphaeroplealean chlorophyte green algae.
Oocystaceae	Cf. Eremosphaera	E. sp.	Lake Unita		An oocystacean chlorophyte green algae.
Scenedesmaceae	Coelastrum	Cf. C. verrucosum Retnsch	Lake Unita		A sphaeroplealean chlorophyte green algae.
Schizochlamydaceae	Schizochlamys	†S. haywellensis Bradley, 1970	Lake Gosiute (Wilkins Peak Member)		A sphaeroplealean chlorophyte green algae.

Charophytes

Family	Genus	Species	Member	Abundance	Notes	Image
Zygnemataceae	Spirogyra	†S. wyomingia Bradley, 1970	Lake Gosiute (Wilkins Peak Member)		A zygnematacean charophyte green algae.

Euglenozoans

Family	Genus	Species	Member	Abundance	Notes	Image
Phacaceae	Phacus	Cf. P. caudata Hübner, 1886		1 specimen	A Euglenozoan algal flagellate.

Amoebozoans

Family	Genus	Species	Member	Abundance	Notes	Image
Difflugiidae	Difflugia	†D. calcifera Bradley, 931	Lake Unita	1 specimen	A difflugiid lobose testate amoeba.
Quadrulidae	Quadrula	†Q. minuta Bradley, 1931	Lake Unita	1 specimen	A quadrulid lobose testate amoeba.

Cercozoans

Family	Genus	Species	Member	Abundance	Notes	Image
Euglyphidae	Euglypha	†E. robusticornis Bradley, 1931	Lake Unita		A Cercozoan eukaryote.

Plants

Liverworts

Family	Genus	Species	Member	Abundance	Notes	Image
Jungermanniaceae	†Jungermannites	†J. eophilus (Cockerell) Steere, 1947	Lake Unita	1 specimen	A leafy liverwort morphospecies. First described as Lejeunea eophila (1925).

Lycophytes

Family	Genus	Species	Member	Abundance	Notes	Image
Isoetaceae	†Isoetites	†I. horridus (Dawson) R.W. Brown, 1939	Lake Unita	1 specimen	An isoetaceous quillwort morphospecies. First described as Carpolithes horridus (1884).

Ferns

Family	Genus	Species	Member	Abundance	Notes	Image
Aspleniaceae	Asplenium	†A. delicatula (R.W. Brown) MacGinitie, 1969			A spleenwort fern leaf morphospecies. First described as Aneimia delicatula (1929) Includes Phyllites furcivena (1934) Jr homonym of Asplenium delicatulum C. Presl, 1825
		†A. eoligniticum Berry, 1916	Lake Unita	2 specimens	A spleenwort fern leaf morphospecies.
Equisetaceae	Equisetum	†E. haydenii Lesquereux, 1872	Not Green River Formation		A scouring rush morphospecies. First ascribed to the formation Excluded by Knowlton (1923) as a Washakie Formation species.
		†E. winchesteri Brown, 1929	Lake Gosiute	1 specimen	A scouring rush morphospecies.
		†E. wyomingense Lesquereux, 1873	Lake Gosiute Lake Unita		A scouring rush morphospecies.
Lygodiaceae	Lygodium	†L. dentoni Lesquereux, 1875	Rejected	1? specimen	A climbing fern species. Type specimens missing by 1923 Brown (1934) rejected per Knowlton and dubious locality.
		†L. kaulfussi Heer, 1861	Fossil Lake Lake Gosiute Lake Unita	Numerous	A climbing fern species. First described as Lygodium neuropteroides (1870)
Marattiaceae	Danaea	†D. coloradensis Knowlton, 1923	Lake Unita	1 specimen	A possible marattiaceous fern leaf morphospecies. The genus affiliation was questioned by MacGinitie (1969).
Marsileaceae	Marsilea	†M. sprungerorum Hermsen, 2019	Lake Uinta (Parachute Creek Member)	Numerous	A water clover fern.
Osmundaceae	Osmunda?	†Q.? sepulta (Newberry) Knowlton, 1923	Lake Gosiute	2 specimens	A possible osmundaceous fern leaf morphospecies. First described as Pecopteris sepulta (1883)
Salviniaceae	Azolla	†A. berryi Brown, 1934	Lake Unita	1 specimen	A mosquito fern morphospecies.
Pteridaceae	Acrostichum	†A. hesperium Newberry, 1883	Lake Gosiute	3 specimens	A parkerioid leather fern leaf morphospecies.

Cycadalean palynomorphs

Genus	Species	Member	Abundance	Notes	Image
†Cycadopites	†C. sp.	Lake Uinta (Parachute Creek Member)	Numerous	Various Cycas - Zamia Cycad palynomorphs.
†Dioonipites	†D. sp.		5 specimens	A Dioon like Cycad palynomorph.

Conifers

Family	Genus	Species	Member	Abundance	Notes	Image
Cupressaceae	Sequoia	Cf. †S. affinis Lesquereux, 1876	Lake Unita		A coast redwood relative.
Pinaceae	Picea	†P. pinifructus Brown, 1929	Lake Unita	1 specimen	First identified as a spruce seed. Deemed a jr synonym of Cercidiphyllum arcticum by Brown (1939).
	Pinus	†P. balli Brown, 1934	Lake Unita	1 specimen	A 3-needle pine morphospecies.
		†P. florissanti Lesquereux, 1885	Lake Unita	1 specimen	A pine morphospecies.
incertae sedis	†Taxites	†T. eocenica Brown, 1929	Lake Uinta	1 specimen	Isolated conifer needles of uncertain affiliation. Similar to some Cupressaceae, Pinaceae, and Taxaceae genera.

Conifer palynomorphs

Genus	Species	Member	Abundance	Notes	Image
Abies	†A. concolipites Wodehouse, 1933	Lake Uinta (Parachute Creek Member)	2 specimens	A true fir palynomorph similar to Abies concolor.
†Abietipites	†A. antiquus Wodehouse, 1933		2 specimens	A generally abietine-like palyonomorph.
†Cedripites	†C. eocenicus Wodehouse, 1933		1 specimen	A Cedrus libani cedar-like palyonomorph.
†Inaperturopollenites	†I. concedipites (Wodehouse) Krutzsch, 1971		6 specimens	A Cunninghamia-like palyonomorph. First described as Cunninghamia concedipites (1933).
	†I. vacuipites (Wodehouse) Krutzsch, 1971		1 specimen	A Chinese swamp cypress-like palyonomorph. First described as Glyptostrobus vacuipites (1933).
Picea	†P. grandivescipites Wodehouse, 1933		6 specimens	A spruce palyonomorph.
Pinus	†P. scopulipites Wodehouse, 1933		Numerous	A pine palyonomorph. Less common than P. strobipites.
	†P. strobipites Wodehouse, 1933		Numerous	A pine palyonomorph.
	†P. tuberculipites Wodehouse, 1933		6 specimens	A pine palyonomorph.
Taxodium	†T. hiatipites Wodehouse, 1933		Numerous	A bald cypress palyonomorph.
†Tsugaepollenites	†T. viridifluminipites (Wodehouse) R. Potonié, 1958		1 specimen	A hemlock-like palyonomorph. First described as Tsuga viridifluminipites (1933) Moved from Zonalapollenites viridifluminipites (1953).

Gnetalean palynomorphs

Genus	Species	Member	Abundance	Notes	Image
†Gnetaceae-pollenites	†G. eocenipites (Wodehouse) R. Potonié, 1958	Lake Uinta (Parachute Creek Member)	12 specimens	An Ephedra-like palyonomorph. First described as Ephedra eocenipites (1933).

Magnoliids

Family	Genus	Species	Member	Abundance	Notes	Image
Aristolochiaceae	Aristolochia	†A. mortua Cockerell, 1908			An pipevine leaf morphospecies.
Hernandiaceae	Illigera	†I. eocenica Manchester & O'Leary, 2010	Lake Unita		An Illigera seed morphospecies.
Lauraceae	Beilschmiedia	†B. eocenica (R.W. Brown) MacGinitie, 1969 (non-Reid & Chandler, 1933)			A Beilschmiedia leaf morphospecies. Name a jr homonym of Beilschmiedia eocenica E. Reid & M. Chandler, 1933
	Lindera	†L. allardi MacGinitie, 1969			A spicebush leaf morphospecies.
	Ocotea	†O. coloradensis (R.W. Brown) LaMotte, 1952)	Lake Unita	2 specimens	An Ocotea leaf morphospecies. First described as Oreodaphne coloradensis (1934)
		†O. lancifolia (Lesquereux) LaMotte, 1952	Lake Gosiute Lake Unita	2 specimens	An Ocotea leaf morphospecies. First described as Sapindus lancifolius (1883) Moved from Oreodaphne lancifolia (1934) Also includes Oreodaphne knowltoni (1929)
	Persea	†P. coriacea MacGinitie, 1969 (non-Engelhardt, 1895)			A Persea leaf morphospecies. Name a jr homonym of Persea coriacea Engelhardt, 1895
Magnoliaceae	Talauma	†T. multiperforata Kruse, 1954	Lake Gosiute (Eden Valley)		An magnolia wood morphospecies.

Magnoliid palynomorphs

Genus	Species	Member	Abundance	Notes	Image
Liriodendron	†L. psilopites Wodehouse, 1933	Lake Uinta (Parachute Creek Member)	1 specimen	A Liriodendron tulipifera-like tulip tree palyonomorph.

Monocots

Family	Genus	Species	Member	Abundance	Notes	Image
Arecaceae	†Palmacites	†P. florissanti (Lesquereux) Read & Hickey, 1972	Fossil Lake Lake Uinta		A palmately veined palm leaf morphospecies. Moved from Sabalites florissanti (1930) First described as Flabellaria florissanti (1885).
	†Palmoxylon	†P. colei Tidwell, 1973	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
		†P. contortum Tidwell, 1973	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
		†P. edenense Tidwell, 1973	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
		†P. macginitiei Tidwell et al., 1971	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
	†Phoenicites	†P. haydenii (Newberry) Read & Hickey, 1972		1 specimen	A pinnately veined palm leaf morphospecies. Moved from Geonomites haydenii (1923) First described as Manicaria haydenii (1883).
	†Sabalites	†S. powellii (Newberry) Berry, 1930	Fossil Lake Lake Uinta		A palmate veined palm leaf morphospecies. First described as Sabal powellii (1883).
Cypreaceae?	†Cyperacites	†C.? filiferus	Lake Unita	A number of specimens	Likely rhizomes of monocots?. Oldest specimen first described as Diatryma? filifera, a Gastornis feather. Description of the species as a bird feather was questioned by Alexander Wetmore (1930) Brown (1934) moves the species to Carpolithus filiferus A second group of fossils is described as a cactus, Eopuntia douglassi (1944) Brown redescribes both as cypreaceous rhizomes, and synonymized both species as Cyperus filiferus (1959) Herman Becker (1963) agrees with the synonymy, but moves the species to the Cyperacites morphogenus as Cyperacites? filiferus.
		†C. haydenii (Lesquereux) Knowlton, 1898	Lake Gosiute	Several specimens	A sedge-like leaf morphospecies. First described as Cyperites haydenii (1885)
	Cyperus	†C. chavannesi Heer, 1855	Rejected	1 specimen	A sedge leaf morphospecies. Rejected by Knowlton (1923) due to dubious location.
Dioscoreaceae	Dioscorea	†D. lindgrenii Herrera & Manchester	Fossil Lake (Fossil Butte Member)	2 specimens	sweet potato fruits
		†D. shermanii Herrera & Manchester		1 specimen
Juncaceae	Juncus	Indeterminate	Rejected	?	Scrappy material tentatively identified as rush leaves. Discredited by Knowlton (1923) due to missing specimens
Poaceae	Arundo	†A. reperta Lesquereux, 1876	Lake Uinta	Several specimens	A giant cane grass morphospecies. The species validity of the fossils was questioned by Knowlton (1923) The species was retained by Brown (1934).
Potamogetonaceae	Potamogeton	†P. rubus R.W. Brown, 1934	Lake Unita		A pondweed morphotype.
Typhaceae	Sparganium	†S. antiquum (Newberry) Berry, 1924	Lake Gosiute Lake Uinta		A bur-reed leaf morphospecies. Moved from Brasenia? antiqua (1883) includes Pontederites hesperia (1923)
		†S. eocenicum R.W. Brown, 1929	Lake Gosiute	1 specimen	A bur-reed leaf morphospecies.
	Typha	†T. lesquereuxi Cockerell, 1906	Lake Unita		A cattail morphospecies.
Zingiberaceae	†Musophyllum	†M. complicatum Lesquereux, 1873	Lake Gosiute	5 specimens	A zingiberaceous monocot leaf morphotaxon.

Monocot palynomorphs

Genus	Species	Member	Abundance	Notes	Image
†Arecipites	†A. punctatus Wodehouse, 1933	Lake Uinta (Parachute Creek Member)	1 specimen	A Phoenix dactylifera-like palm palyonomorph.
	†A. rugosus Wodehouse, 1933		1 specimen	A palm palyonomorph.
†Peltandripites	†P. davisii Wodehouse, 1933		1 specimen	A tuckahoe-like arrow-arum palyonomorph.
Potamogeton	†P. hollickipites Wodehouse, 1933		Numerous	A Potamogeton palyonomorph.
†Smilacipites	†S. echinatus Wodehouse, 1933		2 specimen	A Smilax-like palyonomorph.
	†S. herbaceoides Wodehouse, 1933		1 specimen	A Smilax-like palyonomorph.
	†S. molloides Wodehouse, 1933		1 specimen	A Smilax mollis-like palyonomorph.

Ceratophyllales

Family	Genus	Species	Member	Abundance	Notes	Image
Ceratophyllaceae	Ceratophyllum	†C. muricatum incertum (Brown) Herendeen, Les, & Dilcher, 1990	Fossil Lake	19 Specimens	A prickly hornwort subspecies. First described as a full species by Berry (1930). Moved to subspecies by Herendeen, Les, & Dilcher (1990)

Eudicots

Basal Eudicots

Family	Genus	Species	Member	Abundance	Notes	Image
Berberidaceae	"Mahonia"	†"M." eocenica (R.W. Brown) C.A. Arnold, 1936	Lake Uinta	1 Specimen	An oregon grape leaf morphospecies. First described as Odostemon eocenica (1934)
Menispermaceae	†Menispermites	†M. limacioides MacGinitie, 1969			A moon-seed leaf morphospecies.
	Stephania	†S. wilfii	Lake Uinta (Wilkins Peak Member)	Numerous	A moon-seed fruit.
Nelumbonaceae	†Nelumbo	Undescribed	Lake Gosiute		An undescribed Lotus leaf species.
Platanaceae	†Macginitiea	†M. wyomingensis (Knowlton & Cockerell) Manchester, 1986			A sycamore relative species. First named Aralia macrophylla Newberry, 1882 (non-Lindley, 1844) Renamed to Aralia wyomingensis (1919) Moved from Platanus wyomingensis (1969)
Proteaceae	Lomatia	†L.? microphylla Lesquereux, 1876	Rejected	specimens missing	First identified as a guitarplant leaf morphospecies. Rejected by Brown (1934) due to missing specimens & dubious locality.
Trochodendraceae?	†Nordenskioldia	†N. borealis Heer, 1870	Rejected		A basal trochodendroid Rejected by Manchester, Crane & Dilcher (1991) considered Carpites cocculoides like.

Superasterids

Family	Genus	Species	Member	Abundance	Notes	Image
Apocynaceae	†Apocynospermum	†A. coloradensis Brown, 1929	Lake Unita	1 specimen	An apocynaceous seed morphospecies.
	†Aspidospermoxylon	†A. uniseriatum Kruse, 1954	Lake Gosiute		An apocynaceous wood morphospecies.
Araliaceae	†Allopanax	†A. quinus (Macginitie) Doweld	Lake Uinta (Parachute Creek Member)		An Apialean leaf mophogenus. Moved from Araliophyllum quinus (1969).
	Oreopanax	†O. elongatum MacGinitie, 1969	Lake Unita		An Oreopanax leaf morphospecies. Includes Apocynophyllurn wilcoxense (of Berry, 1929), Ternstroemites viridifluminis (1929) and ?Celastrophyllum lesquereuxii (1929)
Asteraceae	†Achaenites	†A. cichorioides Knowlton, 1923	Lake Uinta	1 specimen	An asteraceous seed morphospecies.
Caryophyllaceae	†Alsinites	†A. revelatus Cockerell, 1925	Lake Unita	1 specimen	A caryophyllacous flower morphotaxon.
Ericaceae	Andromeda	†A. delicatula Lesquereux, 1885		2 specimens	A possible bog rosemary ericaceous morphospecies.
Oleaceae	Osmanthus	†O. praemissa (Lesquereux) Cockerell, 1906	Lake Uinta		An Osmanthus leaf morphospecies. Moved from Olea praemissa (1885)
		†O. viridiflumensis (Knowlton) MacGinitie, 1969)	Lake Uinta	1 specimen	An Osmanthus leaf morphospecies. Moved from Ocotea viridiflumensis (1952) First described as Oreodaphne viridiflumensis (1923)
Polemoniaceae	†Gilisenium	†G. hueberi Lot, Manchester, & Dilcher, 1998	Fossil Lake	1 specimen	A polemoniaceous herb species.
Solanaceae	†Lycianthoides	†L. calycina Deanna et al.	Lake Uinta (Parachute Creek Member)	3 specimens	A nightshade family fruit
Styracaceae	Styrax	†S. transversa MacGinitie, 1969	Lake Unita		A snowbell leaf morphotype.
Symplocaceae	Symplocos	†S. exilis MacGinitie, 1969	Lake Unita		A Symplocaceous leaf morphotype.

Superrosids

Family	Genus	Species	Member	Abundance	Notes	Image
Altingiaceae	Liquidambar	†L. callarche Cockerell, 1925	Lake Unita	2 specimens	A sweetgum species.
Anacardiaceae	†Barkleya	†B. schinoloxa (R. W. Brown) Manchester & Judd	Lake Gosiute (Laney Member) Lake Uinta (Parachute Creek Member)	Numerous	An anacardiaceous samara. Moved from Anacardites schinoloxus (1929). Includes Anacardites schinus R. W. Brown, 1934 Possibly the fruits of "Rhus" nigricans.
	†Edenoxylon	†E. parviareolatum Kruse, 1954	Lake Gosiute (Eden Valley)		An anacardiaceous wood morphospecies.
	Rhus	†R. longepetiolata (Lesquereux) R.W. Brown, 1934	Lake Gosiute		An anacardiaceous leaf morphotype. Moved from Ailanthus longepetiolatus (1934)
		†"R." nigricans (Lesquereux) Knowlton, 1923	Lake Gosiute (Laney Member) Lake Uinta (Parachute Creek Member)	Numerous	An anacardiaceous leaf morphotype. Moved from Myrica nigricans (1929). Possibly the leaves of Barkleya schinoloxa. "R." nigricans synonymy Comptonia insignis Lesquereux of Brown, 1934 Cupanites nigricans (Lesquereux) Berry, 1924 Koelreuteria nigricans (Lesquereux) Brown, 1946 Koelreuteria viridifluminis (Hollick) Brown, 1934 Myrica alkalina Lesquereux, 1883 Myrica ludwigii Schimper of Lesquereux, 1872 Myrica undulata? Schimper of Lesquereux,1878 (pro part) Myrica zachariensis Saporta of Lesquereux, 1883 (pro part) Planera variabilis Newberry, 1883 (pro part) Rhus myricoides Knowlton, 1923 Rhus variabilis (Newberry) Knowlton, 1923 Staphylea viridiftuminis Hollick, 1929
	Schmaltzia	†S. vexans Cockerell, 1906	Lake Gosiute		A sumac leaf morphospecies. Moved from Rhus vexans (1885) Considered nomen dubium by MacGinitie (1969)
	†Schinoxylon	†S. actinoporosum Kruse, 1954	Lake Gosiute (Eden Valley)		An anacardiaceous wood morphospecies.
	Toxicodendron	†T. winchesteri (Knowlton) MacGinitie, 1969	Lake Gosiute		A poison oak leaf morphospecies. First described as Sapindus winchesteri (1923)
Burseraceae	Bursera	†B. inaequalateralis (Lesquereux) MacGinitie, 1969			A bursera leaf morphotype. First named as Alnites unaequalateralis (1876) B. inaequalateralis synonymy Alnites inaequalateralis Lesquereux, 1878 Alnus inaequalateralis (Lesquereux) Lesquereux, 1883 Comptonia? anomala Knowlton, 1923 Planera inaequalateralis (Lesquereux) Knowlton, 1923 Planera variabilis Newberry, 1883
Cannabaceae	Celtis	†C. mccoshii Lesquereux, 1883	Lake Gosiute Lake Unita	2 specimens	A hackberry leaf morphospecies. Includes Celtis debequensis (1928)
Capparaceae	†Forchhammerioxylon	†F. scleroticum Kruse, 1954	Lake Gosiute (Eden Valley)		A capparaceous wood morphospecies.
Celastraceae	Celastrus	†C. winchesteri (Knowlton) MacGinitie, 1969	Lake Unita	1 specimen	A staff vine leaf morphospecies. Moved from Juglans winchesteri (1923) Includes Juglans alkalina (1875 [1876] )
	†Celastrophyllum	†C. emarginatum	Lake Unita		A staff vine relative leaf morphospecies. MacGinitie (1969) considered the species nomen dubium.
Crassulaceae	Sedum?	†S.? hesperium Knowlton, 1923	Lake Uinta	1 specimen	A possible stonecrop leaf morphospecies. Considered nomen dubium by MacGinitie (1969)
Euphorbiaceae	†Heveoxylon	†H. microporosum Kruse, 1954	Lake Gosiute (Eden Valley)		An euphorb wood morphospecies.
Fabaceae	Amorpha	†A. utensis Cockerell, 1925	Lake Unita	1 specimen	A legume leaf morphospecies.
	Arcoa	†A. lindgreni Herendeen & Herrera, 2019	Fossil Lake	3 specimens	A caesalpinioid legume.
	Caesalpinia	†C. flumen-viridensis Herendeen & Dilcher, 1991	Lake Gosiute	2 specimens	A Caesalpinia subg. Mezoneuron seed morphospecies.
		†"C." pecorae R.W. Brown, 1956	Lake Unita	2 specimens	First named as a Caesalpinia leaf morphospecies. Considered a Detarieae tribe species by Herendeen & Dilcher (1991).
	†Caesalpinites	†C. falcatus MacGinitie, 1969			A Caesalpinia-like leaf morphospecies.
	Cercis	†C. tenuinervis (Lesquereux) R.W. Brown, 1934	Lake Gosiute	1 specimen	A redbud leaf morphospecies. Considered nomen dubium by MacGinitie (1969).
	Erythrina	†E. roanensis MacGinitie, 1969	Lake Unita		A coral tree leaf morphospecies.
	Gymnocladus	†G. hesperia (R.W. Brown) MacGinitie, 1969	Lake Uinta		A coffeetree leaf morphospecies. Moved from Cassia hesperia (1929) Includes Chrysobalanus lacustris (1929) & Banisteria bradleyi (1929)
	†Leguminosites	†L. alternans Lesquereux, 1875 [1876]	Rejected		A legume leaf morphospecies. Rejected due to missing specimens by Knowlton (1923) and Brown (1934)
		†L. lesquereuxiana Knowlton, 1898	Lake Unita		A legume leaf morphospecies. L. lesquereuxiana synonymy Bumelia coloradensis Cockerell, 1925 Dalbergia knowltoni Cockerell, 1925 Dalbergia retusa Knowlton, 1923 [non-Hemsl. 1878] Leguminosites cassioides Lesquereux, 1878 [non-Engelhardt, 1870] Sophora coloradensis Knowlton, 1923
		†L. regularis MacGinitie, 1969	Lake Unita		A legume leaf morphospecies.
	↑Parvileguminophyllum	†P. coloradensis (Knowlton) Call & Dilcher, 1994	Lake Uinta		A legume leaf morphospecies. Moved from Mimosites coloradensis (1923). P. coloradensis synonymy Podogonium americanum Lesquereux, 1878 (of Brown 1929) Mimosites debequensis R.W. Brown, 1929 Mimosites falcatus R.W. Brown, 1929
	Swartzia	†S. wardelli MacGinitie, 1969	Lake Uinta		A swartzieous leaf morphotype.
Fagaceae	Quercus	†Q. castaneopsis Lesquereux, 1883	Lake Gosiute Lake Uinta	Several fragmentary specimens	An oak morphospecies. P. cinnamomoides synonymy Quercus castanoides Newberry, 1883 Ilex maculata Lesquereux, 1883 Ilex? affinis Lesquereux, 1872 Considered nomen dubium by MacGinitie (1969).
		†"Q. cuneatus" MacGinitie, 1969 (non-Wangenh. 1787	Lake Unita		An oak leaf morphospecies. Name a jr homonym of Quercus cuneata Wangenheim, 1787
		†Q. drymeja Unger, 1847	Rejected		An oak morphospecies. First described from Europe, Rejected from flora by MacGinitie (1969).
		†Q. petros MacGinitie, 1969	Lake Unita		An oak leaf morphospecies.
		†Q. utensis Cockerell, 1927	Lake Unita		An oak morphospecies. Considered nomen dubium by MacGinitie (1969).
Hamamelidaceae	Distylium	†D. eocenica (R.W. Brown) MacGinitie, 1969	Lake Unita	1 specimen	A winter-hazel leaf morphotype. Moved from Betula coloradea (1934) First named Betula eocenica (1928 non Ettingshausen (1858) Also includes Grewiopsis cissifolius (1929)
Juglandaceae	Juglans	†J. crossii Knowlton, 1898	Lake Gosiute	1 specimen?	A walnut leaf morphospecies.
		†J. bilinica Unger, 1850		1 specimen	A walnut leaf morphospecies. Moved from Hicoria juglandiformis (1898)
		†J. schimperi Lesquereux, 1873	Lake Unita		A walnut leaf morphospecies. Includes Juglans occidentalis (1883) & Ficus ungeri (1872)
	†Palaeocarya	Cf. †P. clarnensis Manchester, 1987	Lake Gosiute?		A Palaeocarya nut morphospecies similar to Palaeocarya clarnensis.
		†P. uintaensis (MacGinitie) Manchester, 1987	Lake Unita		A Palaeocarya nut morphospecies.
	Platycarya	†P. castaneopsis (Lesquereux) Wing & Hickey, 1984	Fossil Lake Lake Uinta		A Platycarya species. Moved from Quercus castaneopsis (1883)
	Pterocarya	†P. macginitii Manchester & Dilcher, 1982	Lake Gosiute		A wingnut seed morphospecies.
		†P. roanensis MacGinitie, 1969	Lake Unita		A wingnut leaf morphospecies.
Malvaceae	Hibiscus	†H. roanensis R.W. Brown, 1934	Lake Unita		An Hibiscus leaf morphospecies.
	Ochroma	†O. murata R.W. Brown, 1934	Lake Unita	1 specimen	A balsa tree leaf morphospecies.
	Sterculia	†S. coloradensis R.W. Brown, 1929	Lake Unita	1 specimen	A sterculioid leaf morphospecies.
	Triumfetta	T. ovata MacGinitie, 1969 (non-De Candolle, 1824)			A burbark leaf morphospecies. Jr homonym of Triumfetta ovata De Candolle, 1824
Meliaceae	Cedrela	†C. trainii C.A. Arnold, 1936	Lake Unita		A meliaceous leaf morphospecies.
Moraceae	†Welkoetoxylon	†W. multiseriatum Boonchai, Manchester & Wheeler	Lake Gosiute (Tipton Shale Member)	1 specimen	A moraceous wood morphogenus.
Myricaceae	Morella scalariformis	†M. scalariformis (Kruse), Wheeler & Manchester, 2021	Lake Gosiute (Eden Valley)		A myricaceous wood morphospecies. First named as Myrica scalariformis (1954) Moved from Myricoxylon scalariforme (1962)
	Myrica	†M. salicina Unger, 1850	Rejected	sp missing	Identified as an European Myrica species Rejected by Knowlton (1923) due to the specimens being missing.
Myrtaceae	†Syzygioides	†S. americana (Lesquereux) Manchester, Dilcher, & Wing, 1998	Lake Unita	Circa 26 specimens	A myrtaceous species. First described as Eucalyptus americana (1872) Includes Ficus omballi (1929) & Eugenia americana (1969)
Rhamnaceae	†Berchemiopsis	†B. paucidentata MacGinitie, 1969	Lake Unita		A rhamnaceous leaf morphotype.
	Ptelea	†P. cassioides (Lesquereux) MacGinitie, 1969	Rejected		Type specimen leaf from the Florissant Formation Green River Fruits rejected by Call & Dilcher as not Ptelea.
		†P. paliuruoides (R.W. Brown) Manchester & O'Leary, 2010	Lake Unita		A rhamnaceous seed morphospecies. First describe as Carpolithus paliuroides (1934)
	†Rainbowia	†R. rhamnoides Manchester & O'Leary, 2010	Lake Unita		A rhamnaceous seed morphospecies.
Rosaceae	Prunus	†P. stewarti MacGinitie, 1969	Lake Gosiute		A cherry leaf morphospecies. First named Amygdalus gracilis (1883) Moved from jr homonym Prunus gracilis (non Engelm. & Gray, 1845)
	Rosa	†R. hillae Lesquereux, 1885			A rose leaf morphospecies.
Rutaceae	†Amyridoxylon	†A. ordinatum Kruse, 1954	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
	Fagara	†F. biseriata Kruse, 1954	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
		†F. monophylloides Kruse, 1954	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
Salicaceae	Populus	†P. cinnamomoides (Lesquereux) MacGinitie, 1969	Lake Uinta	Numerous	An Aspen/cottonwood relative. First described as Ceanothus cinnamomoides (1872) P. cinnamomoides synonymy Carpites newberryana Knowlton, 1926 Cinnamomum scheuchzeri Heer of Lesquereux, 1883 Ficus wyomingiana Lesquereux, 1878 Ilex wyomingensis Lesquereux, 1878 Ilex vyomingensis Lesquereux, 1878 Luehea newberryana (Knowlton) MacGinitie, 1974 Zizyphus cinnamomoides (Lesquereux) Lesquereux, 1878 Zizyphus longifolia Newberry, 1883
		†P. wilmattae Cockerell, 1925	Lake Uinta	Numerous	An Aspen/cottonwood relative.
		†P. tidwellii Manchester, Judd, & Handley, 2006	Lake Uinta	Numerous	An Aspen/cottonwood relative.
	†Pseudosalix	†P. handleyi Boucher, Manchester, & Judd, 2003	Lake Uinta (Parachute Creek Member)		A salicaceous species. Includes Potentilla? byrami (1925), Antholithes polemonioides (1929) & Antholithes sp.6 (1934)
	Salix	†S. cockerelli R.W. Brown, 1934	Lake Unita	4 Specimens	A willow leaf morphospecies.
		†S. longiacuminata Knowlton ex R.W. Brown, 1934	Lake Gosiute	1 specimen	A willow leaf morphospecies. Includes Banksia cockerelli (1929)
Sapindaceae	Allophylus	†A. flexifolia (Lesquereux) MacGinitie, 1969	Lake Gosiute Lake Uinta	4 Specimens	An Allophylus leaf morphospecies. First named Euonymus flexifolia (1885) A. flexifolia synonymy Fraxinus flexifolia (Lesquereux) Brown, 1934 Fraxinus petiolata Brown, 1929 Odostemon reynoldsii Hollick, 1929
	Athyana	†A. balli (R.W. Brown) MacGinitie, 1969	Lake Unita		A sapindaceous leaf morphotype. Moved from Rhus balli (1934).
	"Cardiospermum"	†"C." coloradensis (Knowlton) MacGinitie, 1969	Lake Uninta	Numerous	A possible ballon vine leaf morphospecies. Moved from Lomatia coloradensis (1929) "C." coloradensis synonymy Lomatia obtusiuscula Cockerell, 1925 Melia coloradensis (Knowlton) Cockerell, 1925 Phyllites coloradensis Knowlton, 1923 Pimelea spatulata Knowlton, 1923
	Koelreuteria	†K. allenii (Lesquereux) W.N. Edwards, 1929	Fossil Lake (Fossil Butte Member) Lake Unita (Parachute Creek Member)		A golden rain tree fruit morphospecies. Includes Koelreuteria uiridifluminis seeds
	Sapindus	†S. dentoni Lesquereux, 1876	Lake Unita		A soapberry leaf morphotype. Includes Salix media (1872) & Myrica torryi (1929)
		†S. obtusifolius Lesquereux, 1874	Lake Unita		A soapberry leaf morphotype. Includes Salix media (1872) & Myrica torryi (1929)
	Thouinia	†T. eocenica R.W. Brown, 1929	Lake Unita		A soap berry leaf morphospecies.
Simaroubaceae	Ailanthus	†A. lesquereuxii Cockerell, 1927			A tree of heaven fruit morphospecies
Surianaceae	Suriana	†S. inordinata Kruse, 1954	Lake Gosiute (Eden Valley)		A surianaceous wood morphospecies.
Tetramelaceae	†Parvaspicula	†P. lepidioides (Cockerell) Deanna et al.	Lake Uinta (Parachute Creek Member)	Numerous	A tetramelaceous fruit morphotype. The type species is Clethra (?) lepidioides Cockerell (1925) Also includes Antholithes pendula (1929)
	†Punctaphyllum	†P. glandulosa (Brown) Manchester, Judd, & Kodrul, 2023	Lake Uinta (Parachute Creek Member)	Numerous	A tetramelaceous leaf morphotype. The type species is Cucurbita glandulosa (1929) Includes Aleurites glandulosa (1969) & Dendropanax latens (1974) Also includes the Ficus mississippiensis of Brown (1928)
Ulmaceae	†Cedrelospermum	†C. nervosum (Newberry) Manchester, 1989	Lake Uinta	Numerous	An ulmaceous species. First named as Planera nervosa (1883) C. nervosum synonymy Banksites lineatulus Cockerell, 1925 Dalbergia viridiflumensis Knowlton, 1923 Lomatia lineatulus (Cockerell) MacGinitie, 1969 Myrica praedrymeja Knowlton, 1923 Myrica minuta Knowlton, 1923 Rhus myricoides (Knowlton) Brown, 1929 Salix lineara Knowlton, 1923 Zelkova nervosa (Newberry) Brown, 1929
Vitaceae	Cissus	†C. parrottiaefolia Lesquereux, 1876	Rejected	3 specimens	A grape relative leaf morphospecies. Rejected from formation by Knowlton (1923) due to distinctly different matrix.

Eudicot palynomorphs

Genus	Species	Member	Abundance	Notes	Image
†Ailanthipites	†A. berryi Wodehouse, 1933	Lake Uinta (Parachute Creek Member)		An Ailanthus glandulosa-like soapberry family palynomorph.
†Alni-pollenites	†A. speciipites (Wodehouse) Raatz, 1938		Numerous	An alder-like palynomorph. first described as Alnus speciipites (1933)
Betula	†B. claripites Wodehouse, 1933		3 specimens	A Betula lenta-like birch palynomorph.
†Caprifoliipites	†C. viridi-fluminis Wodehouse, 1933		1 specimen	A caprifoliaceous-like palynomorph.
Carpinus	†C. ancipites Wodehouse, 1933			A Carpinus-like palynomorph.
Carya	†C. juxtaporipites (Wodehouse) LaMotte, 1952		1 specimen	A hickory-like palynomorph. First described as Hicoria juxtaporipites (1933)
	†C. viridi-fluminipitis (Wodehouse) Wilson & Webster, 1944		1 specimen	A Carya myristiciformis-like hickory palynomorph. First described as Hicoria viridi-fluminipites (1933)
Ericipites	†E. brevisulcatus Wodehouse, 1933		1 specimen	A Calluna vulgaris-like heath family palynomorph.
	†E. longisulcatus Wodehouse, 1933		2 specimens	A heath family-like palynomorph.
†Juglanspollenites	†J. nigripites (Wodehouse) Wingate & Nichols, 2001		1 specimen	A walnut-like palynomorph. First described as Juglans nigripites (1933)
†Momipites	†M. corylipites (Wodehouse) D.J. Nichols, 1973		2 specimens	An Engelhardia-like palynomorph. First described as Engelhardia corylipites (1933).
	†M. coryloides Wodehouse, 1933		Numerous	A Celtis iguanaea-like hackberry palynomorph.
†Myricipites	†M. dubius Wodehouse, 1933		1 specimen	A bayberry-like palynomorph.
Myriophyllum	†M. ambiguipites Wodehouse, 1933		1 specimen	A Myriophyllum spicatum-like watermillfoil family palynomorph.
†Rhoipites	†R. bradleyi Wodehouse, 1933		4 specimens	A Rhus typhina-like cashew family palynomorph.
Salix	†S. discoloripites Wodehouse, 1933		Numerous	A Salix discolor and S. fragilis-like willow palynomorph.
†Talisiipites	†T. fischeri Wodehouse, 1933		Numerous	A Talisia depressa-like soapberry family palynomorph.
Tilia	†T. crassipites Wodehouse, 1933		1 specimen	A Tilia americana-like linden palynomorph.
	†T. tetraforaminipites Wodehouse, 1933		3 specimens	A Tilia americana-like linden palynomorph.
	†T. vescipites Wodehouse, 1933		1 specimens	A Tilia americana-like linden palynomorph.
†Vitipites	†V. dubius Wodehouse, 1933		1 specimen	A Vitis vinifera-like grape family palynomorph.

Angiosperms of uncertain affiliation

Genus	Species	Member	Abundance	Notes	Image
†Achaenites	†A. cichorioides Knowlton, 1923			A achene like structure of unidentified affiliation.
†Antholithes	†A. dubia R.W. Brown, 1929	Lake Unita	1 specimen	An 8-lobed/petaled flower of unidentified affiliation.
	†A. improbus Lesquereux, 1883		2 specimens	A flowering structure of unidentified affiliation.
	†A. vitaciflora R.W. Brown, 1929	Lake Unita	1 specimen	A cluster of possible vitaceous flowers-fruits.
	†A. winchesteri (Knowlton) MacGinitie, 1969	Lake Unita	1 specimen	A flowering structure of unidentified affiliation. First described as Sambucus? winchesteri (1923)
	†A. sp 1 R.W. Brown, 1934	Fossil Lake	1 specimen	A possibly cruciferous or orrograceous flower of unidentified affiliation.
	†A. sp 2 R.W. Brown, 1934	Lake Unita	1 specimen	A flower with many stamens of unidentified affiliation.
	†A. sp 3 R.W. Brown, 1934	Lake Unita	1 specimen	A malvaceous looking flower of unidentified affiliation.
	†A. sp 4 R.W. Brown, 1934	Lake Unita	1 specimen	A flower with a superior ovary surrounded by the five-toothed calyx.
	†A. sp 5 R.W. Brown, 1934	Lake Unita	1 specimen	A flower with tubular receptacle surmounted by a four-parted calyx.
	†A. sp 7 R.W. Brown, 1934	Fossil Lake	1 specimen	A large flower or a badly distorted leaf of a Nelumbo?
†Bonanzacarpum	†B. sprungerorum Manchester & Lott, 2019	Lake Uinta (Parachute Creek Member)	Numerous	A eudicot fruit of uncertain phylogenetic placement.
†Carpites	†C. inquirenda Knowlton, 1923	Lake Unita	1 specimen	A nutlet or fruit of unidentified affiliation.
	†C. newberryanus Knowlton, 1923		3 specimens	A nutlet of unidentified affiliation. First described as Juglans occidentalis nuts (1898). First spelled Carpites newberryana
	†C. viburni Lesquereux, 1878		1 specimen	A seed or nutlet of unidentified affiliation. Includes Carpolithes capsularis (1929)
†Carpolithus	†C. alabaccus R.W. Brown, 1934	Lake Unita	1 specimen	A winged or membranous fruit of unidentified affiliation.
	†C. caryophylloides Knowlton, 1923	Lake Unita		A dry capsular fruit of unidentified affiliation.
	†C. cassioides R.W. Brown, 1929	Lake Unita	2 specimens?	A legume like pod of unidentified affiliation.
	†C. insectoides R.W. Brown, 1934	Lake Unita	1 specimen	A winged fruit of unidentified affiliation.
	†C. palrnites R.W. Brown, 1929	Lake Unita	1 specimen	A palm fruit or husk like fossil of unidentified affiliation.
	†C. prunoides R.W. Brown, 1934	Lake Unita	1 specimen	A flattened fruit of unidentified affiliation. First named Carpolithus ellipticus Brown, 1929 (non-Sternberg, 1825)
	†C. serratifolius R.W. Brown, 1929	Lake Unita	1 specimen	Branchlet with leaves and fruits of possible salicaceous affinity.
	†C. vitaceus R.W. Brown, 1934	Lake Unita	1 specimen	A possibly vitaceous seed of unidentified affiliation.
†Caulinites	†C. acanthus R.W. Brown, 1929	Lake Unita	1 specimen	A thorny stem of unidentified affiliation.
	†C. prehensus R.W. Brown, 1929	Lake Unita	1 specimen	A climbing plant tendril of unidentified affiliation.
†Chaneya	†C. tenuis (Lesquereux) Wang & Manchester, 2000	Lake Gosiute Lake Unita		A sapindalean flower of uncertain affiliations. Includes Astronium truncatum (1953).
†Dicotylophyllum	†D. lesquereuxii (Knowlton) J.A. Wolfe & Tanai, 1987		1 specimen	A leaf of uncertain affiliation. First described as Acer indivisum Lesquereux, 1885 (non-Weber, 1852) Moved from Acer lesquereuxii Knowlton, 1898
†Palibinia	†P. comptonifolia (Brown) Manchester, Judd, & Kodrul, 2023	Lake Uinta (Parachute Creek Member)		A pentapetalean eudicot of uncertain affiliation. Moved from Vauquelinia comptonifolia (1969) Originally named Banksia comptonifolia (1934)
†Phyllites	†P. ericoides R.W. Brown, 1934	Lake Unita	1 specimen	A thick leaf of unidentified affiliation.
	†P. fremonti Unger, 1850	Rejected	1 specimen	A leaf of unidentified affiliation. R.W. Brown (1934) rejected due to doubtful locality and lost specimen.
	†P. glandiferus R.W. Brown, 1934	Lake Unita	1 specimen	An odd-pinnately compound leaf of possible fabaceous affinity.
	†P. juncoides R.W. Brown, 1929	Lake Unita	1 specimen	A leaf of unidentified affiliation.
	†P. ouratea R.W. Brown, 1934	Lake Unita	1 specimen	A possibly herbaceous leaf of unidentified affinity.
	†P. winchesteri Knowlton, 1923	Lake Unita	1 specimen	A leaf of unidentified affiliation.

Plants of uncertain affiliation

Genus	Species	Member	Abundance	Notes	Image
†Rhizocaulon	†R. dichotomum R.W. Brown, 1929	Lake Unita	1 specimen	A root of unidentified affiliation. Has a distinct branched fork.
	†R. natans R.W. Brown, 1929	Lake Unita	1 specimen	A rootlet of unidentified affiliation. small with Trapa natans-like rhizoids.