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This list is a subsection of the List of members of the National Academy of Sciences , which includes approximately 2,500 living members and 500 foreign associates of the United States National Academy of Sciences , each of whom is affiliated with one of 31 disciplinary sections. Members are listed under their primary subsection and each person's name, primary institution, and election year are given.
The designation (d) after the name means the member is deceased.
Name
Institution
Year
John Abelson
California Institute of Technology
1985
Julius Adler
University of Wisconsin–Madison
1978
Paul Ahlquist
University of Wisconsin–Madison
1993
Bruce Alberts
University of California, San Francisco
1981
Jorge Allende
University of Chile
2001
Sidney Altman
Yale University
1990
Bruce Ames
University of California, Berkeley
1972
Gilbert Ashwell
National Institutes of Health
1979
Tania Baker
Massachusetts Institute of Technology
2007
Clinton Ballou (d)
University of California, Berkeley
1975
David Bartel
Massachusetts Institute of Technology
2011
Jacqueline Barton
California Institute of Technology
2002
Brenda Bass
University of Utah
2015
Philip Beachy
Johns Hopkins University
2002
Lorena Beese
Duke University
2009
Marlene Belfort
New York State Department of Health
1999
James Berger
Johns Hopkins University
2013
Klaus Biemann
Massachusetts Institute of Technology
1993
Pamela Björkman
California Institute of Technology
2001
Elizabeth Blackburn
University of California, San Francisco
1993
Elkan Blout (d)
Harvard University
1969
Michael Botchan
UC Berkeley
2008
Herbert Boyer
University of California, San Francisco
1985
Paul D. Boyer (d)
University of California, Los Angeles
1970
Roscoe Brady (d)
National Institutes of Health
1975
Ronald Breaker
Yale University
2013
Patrick O. Brown
Stanford University
2002
John Buchanan (d)
Massachusetts Institute of Technology
1962
Giulio Cantoni (d)
National Institutes of Health
1983
John Carbon
University of California, Santa Barbara
1986
H. E. Carter (d)
University of Arizona
1953
Thomas R. Cech
Howard Hughes Medical Institute
1987
Michael Chamberlin
University of California, Berkeley
1986
Britton Chance (d)
University of Pennsylvania
1954
Aaron Ciechanover
Technion-Israel Institute of Technology
2007
W. Wallace Cleland (d)
University of Wisconsin–Madison
1985
Philip Cohen
University of Dundee
2008
Seymour Cohen (d)
Marine Biological Laboratory
1967
Mildred Cohn (d)
University of Pennsylvania
1971
Joan W. Conaway
Stowers Institute for Medical Research
2020
Minor Coon (d)
University of Michigan
1983
Nicholas R. Cozzarelli (d)
University of California, Berkeley
1989
Nancy Craig
Johns Hopkins University
2010
Benjamin Cravatt III
Scripps Research Institute
2014
James Dahlberg
University of Wisconsin–Madison
1996
Seth Darst
Rockefeller University
2008
Earl Davie (d)
University of Washington
1980
Christian de Duve (d)
Christian de Duve Institute of Cellular Pathology
1975
Hector DeLuca
University of Wisconsin–Madison
1979
Peter Dervan
California Institute of Technology
1986
Raymond Deshaies
Caltech
2016
Jack Dixon
Howard Hughes Medical Institute
2000
Russell Doolittle
University of California, San Diego
1984
Jennifer Doudna
University of California, Berkeley
2002
Gideon Dreyfuss
University of Pennsylvania
2012
Gerald Edelman (d)
The Scripps Research Institute
1969
Ronald Estabrook
University of Texas Southwestern Medical Center at Dallas
1979
Gary Felsenfeld
National Institutes of Health
1976
Edmond Fischer (d)
University of Washington
1973
Perry A. Frey
University of Wisconsin–Madison
1998
Irwin Fridovich
Duke University
1981
E. Peter Geiduschek
University of California, San Diego
1974
Quentin Gibson (d)
Rice University
1982
Alexander Glazer
University of California System
2001
Rachel Green
Johns Hopkins University
2012
Carol Greider
Johns Hopkins University
2003
Marianne Grunberg-Manago
Institut de Biologie Physico-Chimique
1982
Michael Grunstein
University of California, Los Angeles
2008
Irwin Gunsalus (d)
University of Illinois at Urbana–Champaign
1965
Lowell Hager
University of Illinois at Urbana–Champaign
1995
Sen-itiroh Hakomori
Pacific Northwest Research Institute
2000
Gordon Hammes (d)
Duke University
1973
Gregory Hannon
Cold Spring Harbor Laboratory
2012
F. Ulrich Hartl
Max Planck Institute of Biochemistry
2011
Ari Helenius
ETH Zurich
2009
Richard Henderson
Medical Research Council
1998
Wayne Hendrickson
Columbia University
1993
Leon Heppel (d)
Cornell University
1970
Avram Hershko
Technion Israel Institute of Technology
2003
George Hess
Cornell University
1987
Robert L. Hill
Duke University
1975
Mahlon Hoagland (d)
Worcester Foundation for Biomedical Research
1984
B. L. Horecker (d)
Cornell University
1961
Arthur Horwich
Yale University
2003
Robert Huber
Max Planck Institute for Biochemistry
1995
Jerard Hurwitz (d)
Memorial Sloan-Kettering Cancer Center
1974
Clyde Hutchison
University of North Carolina at Chapel Hill
1995
William Jencks (d)
Brandeis University
1971
Gerald Joyce
Scripps Research Institute
2001
Howard Ronald Kaback
University of California, Los Angeles
1987
Paul Kaesberg (d)
University of Wisconsin–Madison
1991
Seymour Kaufman (d)
National Institutes of Health
1986
Yoshito Kaziro (d)
Kyoto University
2000
Eugene P. Kennedy (d)
Harvard University
1964
Peter Kim
Merck & Co., Inc.
1997
Robert Kingston
Harvard University
2016
Judith Klinman
University of California, Berkeley
1994
Jeremy Knowles (d)
Harvard University
1988
Edward Korn
National Institutes of Health
1990
Arthur Kornberg (d)
Stanford University
1957
Roger Kornberg
Stanford University
1993
Alberto Kornblihtt
University of Buenos Aires
2011
Daniel E. Koshland, Jr. (d)
University of California, Berkeley
1966
Stephen Kowalczykowski
University of California, Davis
2007
Joseph Kraut (d)
University of California, San Diego
1988
John Kuriyan
University of California, Berkeley
2001
Alan Lambowitz
University of Texas at Austin
2004
Arthur Landy
Brown University
1999
M. Daniel Lane
Johns Hopkins University
1987
Henry Lardy (d)
University of Wisconsin–Madison
1958
I. Robert Lehman
Stanford University
1977
William Lennarz
State University of New York at Stony Brook
1989
Christopher D. Lima
Memorial Sloan Kettering Cancer Center
2020
Laszlo Lorand (d)
Northwestern University
1987
George Lorimer
University of Maryland, College Park
1997
James Manley
Columbia University
2011
Lynne E. Maquat
University of Rochester
2011
Michael Marletta
The Scripps Research Institute
2006
Emanuel Margoliash (d)
Northwestern University
1975
Brian Matthews
University of Oregon
1986
Rowena Green Matthews
University of Michigan
2002
Stephen L. Mayo
California Institute of Technology
2004
Robert Bruce Merrifield (d)
Rockefeller University
1972
Stanley Miller (d)
University of California, San Diego
1973
Kiyoshi Mizuuchi
National Institutes of Health
1994
Paul Modrich
Duke University
1993
Elizabeth Neufeld
University of California, Los Angeles
1977
Marshall Warren Nirenberg (d)
National Institutes of Health
1967
Harry Noller
University of California, Santa Cruz
1992
Masayasu Nomura (d)
University of California, Irvine
1978
Peter Novick
University of California, San Diego
2013
Michael E. O'Donnell
The Rockefeller University
2006
Erin O'Shea
University of California, San Francisco
2004
Leslie Orgel (d)
Salk Institute for Biological Studies
1990
Roy Parker
University of Colorado Boulder
2012
Armando Parodi
Fundacion Instituto Leloir
2000
Dinshaw Patel
Memorial Sloan-Kettering Cancer Center
2009
Nikola Pavletich
Memorial Sloan-Kettering Cancer Center
2012
David Phillips (d)
University of Oxford
1985
Frank Putnam (d)
Indiana University
1976
Charles Radding
Yale University
1995
Venkatraman Ramakrishnan
Medical Research Council
2004
Linda Randall
University of Missouri
1997
Tom Rapoport
Harvard University
2005
Lester Reed (d)
University of Texas at Austin
1973
Douglas C. Rees
California Institute of Technology
2000
Peter Reichard
Karolinska Institute
1980
Danny Reinberg
New York University
2015
Alexander Rich (d)
Massachusetts Institute of Technology
1970
Charles C. Richardson
Harvard University
1983
Phillips Robbins
Boston University
1982
John D. Roberts (d)
California Institute of Technology
1956
Ellen Rothenberg
California Institute of Technology
2021
Saul Roseman (d)
Johns Hopkins University
1972
William J. Rutter
Synergenics, LLC
1984
Bengt I. Samuelsson
Karolinska Institute
1984
Anthony San Pietro (d)
Indiana University
1983
Aziz Sancar
University of North Carolina at Chapel Hill
2005
Frederick Sanger (d)
Medical Research Council
1967
Robert Sauer
Massachusetts Institute of Technology
1996
Howard Schachman (d)
University of California, Berkeley
1968
Gottfried Schatz
University of Basel
1989
Robert Schimke (d)
Stanford University
1976
Paul Schimmel
Scripps Research Institute
1990
Vern Schramm
Albert Einstein College of Medicine
2007
Brenda Schulman
St. Jude Children's Research Hospital
2014
Kevan Shokat
University of California, San Francisco
2009
Maxine Singer
Carnegie Institution of Washington
1979
Robert Sinsheimer
University of California, Santa Barbara
1967
Emil Smith (d)
University of California, Los Angeles
1962
Hamilton O. Smith
J. Craig Venter Institute
1980
Dieter Söll
Yale University
1997
Nahum Sonenberg
McGill University
2015
David Sprinson (d)
Columbia University
1990
E. R. Stadtman (d)
National Institutes of Health
1969
Thressa Stadtman (d)
National Institutes of Health
1981
George Stark
Cleveland Clinic Foundation
1987
Donald Steiner (d)
University of Chicago
1975
Joan A. Steitz
Yale University
1983
Thomas Steitz (d)
Yale University
1990
Audrey Stevens (d)
Oak Ridge National Laboratory
1998
Bruce Stillman
Cold Spring Harbor Laboratory
2000
Jack Strominger
Harvard University
1970
JoAnne Stubbe
Massachusetts Institute of Technology
1992
Julian M. Sturtevant (d)
Yale University
1973
Wesley Sundquist
University of Utah
2014
Jack Szostak
Harvard University
1998
Herbert Tabor (d)
National Institutes of Health
1977
Paul Talalay (d)
Johns Hopkins University
1987
Glauco Tocchini-Valentini
National Research Council of Italy
2009
Susan S. Taylor
University of California, San Diego
1996
Jeremy Thorner
UC Berkeley
2015
Robert Tjian
University of California, Berkeley
1991
Olke Uhlenbeck
Northwestern University
1993
P. Roy Vagelos
Merck & Co., Inc.
1972
Bert Vallee (d)
Harvard University
1974
Alexander Varshavsky
California Institute of Technology
1995
Sidney Velick (d)
University of Utah
1981
Peter von Hippel
University of Oregon
1978
Salih Wakil (d)
Baylor College of Medicine
1990
Christopher T. Walsh
Harvard University
1989
Peter Walter
University of California, San Francisco
2004
James C. Wang
Harvard University
1986
Herbert Weissbach
Florida Atlantic University
1982
Jonathan Weissman
University of California, San Francisco
2009
James A. Wells
University of California, San Francisco
1999
Stephen C. West
Francis Crick Institute
2016
Reed Wickner
National Institutes of Health
2000
Sue Hengren Wickner
National Institutes of Health
2004
William T. Wickner
Dartmouth College
1996
Richard Wolfenden
University of North Carolina at Chapel Hill
2002
Carl Wu
National Institutes of Health
2006
Wei Yang
National Institutes of Health
2013
Paul Zamecnik (d)
Harvard University
1968
Awards
Biology and Medicine Earth Sciences Engineering Physical Sciences and Mathematics Others
Fellows Associates