2009 in archosaur paleontology
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The year 2009 in Archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur paleontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2009 in paleontology included various significant developments regarding archosaurs.
This article records new taxa of fossil archosaurs of every kind that have been described during the year 2009, as well as other significant discoveries and events related to paleontology of archosaurs that occurred in the year 2009.
Crurotarsans
[edit]- Delfino, M.; Smith, T. (2009). "A reassessment of the morphology and taxonomic status of 'Crocodylus' depressifrons Blainville, 1855 (Crocodylia, Crocodyloidea) based on the Early Eocene remains from Belgium". Zoological Journal of the Linnean Society. 156 (1): 140–167. doi:10.1111/j.1096-3642.2008.00478.x.
Newly named crurotarsans | |||||||
---|---|---|---|---|---|---|---|
Name | Status | Authors | Age | Unit | Location | Notes | Images |
Armadillosuchus[1] |
Valid |
|
Late Cretaceous |
Notosuchian with heavy, armadillo-like body armor consisting of flexible bands and rigid shields |
|||
Barcinosuchus[2] |
Valid |
|
Aptian – Albian |
||||
Collilongus[3] |
Valid |
|
Early Olenekian |
Possible rauisuchian |
|||
Coringasuchus[4] |
Valid |
|
Early Cenomanian |
||||
Duerosuchus[5] |
Valid |
|
Middle Eocene |
||||
Hypselorhachis[6] |
Valid |
|
Middle Triassic |
Possible ctenosauriscid |
|||
Kaprosuchus[7] |
Valid |
|
Upper Cretaceous |
Unusual large mahajangasuchid with hypertrophied caniniform teeth and posteriorly projecting horns |
|||
Kemkemia[8] |
Valid |
|
Cenomanian |
Initially identified as a theropod dinosaur,[8] but subsequently discovered to be a crocodyliform.[9] |
|||
Khoratosuchus[10] |
Valid |
|
Early Cretaceous |
Youngest Mesozoic crocodyliform yet known from Thailand |
|||
Laganosuchus [7] |
Valid |
|
Cenomanian |
||||
Miadanasuchus [11] |
Valid |
|
Campanian |
A new genus for "Trematochampsa" oblita (Buffetaut & Taquet, 1979) |
|||
Morrinhosuchus[12] |
Valid |
|
Late Cretaceous |
Notosuchian from Brazil |
|||
Penghusuchus[13] |
Valid |
|
Late Miocene |
A tomistomine crocodilian. |
|||
Polonosuchus[14] |
Valid |
|
Late Carnian |
A new genus for "Teratosaurus" silesiacus (Sulej, 2005) |
|||
Yacarerani [15] |
Valid |
|
Late Cretaceous |
Cajones Formation |
Non-avian dinosaurs
[edit]Research
[edit]- A new study on theropod furculae is published.[16]
- A "detailed description of the skull and mandible of the Chinese cerapodan ornithischian dinosaur Jeholosaurus shangyuanensis" is published.[17]
- Knoll, F.; Padian, K.; de Ricqles, A. (2009). "Ontogenetic change and adult body size of the early ornithischian dinosaur Lesothosaurus diagnosticus: Implications for basal ornithischian taxonomy". Gondwana Research. 17: 171–179. doi:10.1016/j.gr.2009.03.010.
- Matthews, J. C.; Brusatte, S. L.; Williams, S. A.; Henderson, M. D. (2009). "The first Triceratops bonebed and its implications for gregarious behavior". Journal of Vertebrate Paleontology. 29 (1): 286–290. Bibcode:2009JVPal..29..286M. doi:10.1080/02724634.2009.10010382. S2CID 196608646.
- Williamson, T. E.; Carr, T. D.; Williams, S. A.; Tremaine, K. (2009). "Early ontogeny of pachycephalosaurine squamosals as revealed by juvenile specimens from the Hell Creek Formation, eastern Montana". Journal of Vertebrate Paleontology. 29 (1): 291–294. Bibcode:2009JVPal..29..291W. doi:10.1671/039.029.0111. S2CID 84516840.
- Bittencourt, J.S.; Kellner, A.W.A. (2009). "The anatomy and phylogenetic position of the Triassic dinosaur Staurikosaurus pricei Colbert, 1970". Zootaxa. 2079: 1–56. doi:10.11646/zootaxa.2079.1.1. S2CID 90574494.
- Chin, K.; Hartman, J.H.; Roth, B. (2009). "Opportunistic exploitation of dinosaur dung: fossil snails in coprolites from the Upper Cretaceous Two Medicine Formation of Montana". Lethaia. 42 (2): 185–198. doi:10.1111/j.1502-3931.2008.00131.x.
- Maidment, S.C.R.; Porro, L.B. (2009). "Homology of the palpebral and origin of supraorbital ossifications in ornithischian dinosaurs". Lethaia. 43: 95–111. doi:10.1111/j.1502-3931.2009.00172.x.
- Gates, T.A.; Farke, A.A. (2009). "Biostratigraphic and biogeographic implications of a hadrosaurid (Ornithopoda: Dinosauria) from the Upper Cretaceous Almond Formation of Wyoming, USA". Cretaceous Research. 30 (5): 1157–1163. Bibcode:2009CrRes..30.1157G. doi:10.1016/j.cretres.2009.05.001.
- Moratalla, J.J.; Hernán, J. (2008). "Los Cayos S y D: dos afloramientos con icnitas de saurópodos, terópodos y ornitópodos en el Cretácico Inferior del área de Los Cayos (Cornago, La Rioja, España)". Estudios Geológicos. 64 (2): 161–173. doi:10.3989/egeol.08642.043.
- Taylor, M.P.; Wedel, M.J.; Naish, D. (2009). "Head and neck posture in sauropod dinosaurs inferred from extant animals". Acta Palaeontologica Polonica. 54 (2): 213–220. doi:10.4202/app.2009.0007. S2CID 7582320.
Hadrosaur chewing study
[edit]A study titled "Quantitative analysis of dental microwear in hadrosaurid dinosaurs, and the implications for hypotheses of jaw mechanics and feeding" is published by British paleontologists Mark Purnell, Paul Barrett and student Vince Williams. The paper examined the chewing methods and diet of hadrosaurid ("duck billed") dinosaurs from the Late Cretaceous period. The scientists analyzed hundreds of microscopic scratches on the teeth of a fossilized Edmontosaurus jaw, and believe they determined exactly how a hadrosaur broke down and ate its food, which had previously eluded researchers.
The study found hadrosaurs had a unique way of eating unlike any creature living today. In contrast to a flexible lower jaw joint prevalent in today's mammals, hadrosaurs had a unique hinge between the upper jaws and the rest of its skull. The team found the dinosaur's upper jaws pushed outwards and sideways while chewing, as the lower jaw slid against the upper teeth.
The study also concluded that hadrosaurs likely grazed on horsetails and vegetation close to the ground, rather than browsing higher-growing leaves and twigs. However, Purnell said these conclusions were less secure than the more conclusive evidence regarding the motion of teeth while chewing. Previous studies found contradictory conclusions, and the issue remains a subject of debate.
The findings were published on June 30, 2009 in the journal, The Proceedings of the National Academy of Sciences. Purnell said no previous study had ever employed this method of analyzing microscopic teeth scratches, and that the method could be used to study other areas of scientific research.
New taxa
[edit]Data courtesy of George Olshevky's dinosaur genera list.[18] ~44 dinosaur genera were erected in 2009.
Name | Status | Authors | Discovery year | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Adeopapposaurus[19] | Valid |
|
||||||
Aerosteon[20] | Valid | |||||||
Valid |
|
|||||||
Albertonykus[22] | Valid |
|
lower Maastrichtian |
|||||
Anchiornis[23] | Valid |
|
||||||
Valid |
|
|||||||
Valid |
|
A Spanish Lambeosaurine. | ||||||
Valid |
|
An Australian Megaraptor. Specimen named "Banjo" | ||||||
Baotianmansaurus[27] |
Valid |
|
||||||
Valid |
|
|||||||
Ceratonykus[29] |
Valid |
|
||||||
Diamantinasaurus[30] |
Valid |
|
An Australian Titanosaur. | |||||
Valid |
|
|||||||
Helioceratops[32] |
Valid |
|
||||||
Hesperonychus[33] |
Valid |
|
Smallest known dinosaur from North America. |
|||||
Valid |
|
|||||||
Kinnareemimus[35] |
Valid |
|
||||||
Kol[36] |
Valid |
|
||||||
Valid |
|
|||||||
Levnesovia[38] | Valid |
|
The oldest Hadrosauroidean |
|||||
Limusaurus[39] | Valid |
|
The first Asian ceratosaur to be discovered |
|||||
Luoyanggia[40] | Valid |
|
An oviraptorosaur |
|||||
Malarguesaurus[41] | Valid |
|
||||||
Minotaurasaurus[42] | Valid |
|
||||||
Miragaia[43] | Valid |
|
Long-necked stegosaur. |
|||||
Nothronychus graffami[44] | Valid |
|
A therizinosaurid. |
|||||
Valid |
|
|||||||
Panphagia[45] | Valid |
|
One of the most basal known sauropodomorph.[45] |
|||||
Valid |
|
|||||||
Ruyangosaurus[47] |
Valid |
|
||||||
Shaochilong[48] |
Valid |
|
||||||
Shidaisaurus[49] |
Valid |
|
||||||
Valid |
|
|||||||
Skorpiovenator[51] | Valid |
|
||||||
|
||||||||
Valid |
|
|||||||
Valid |
|
|
||||||
Valid |
|
|||||||
Valid |
|
|||||||
Valid |
|
|||||||
Xianshanosaurus[40] | Valid |
|
A sauropod |
|||||
|
Name published without scientific description in Japanese guidebook "Dinosaur Expo 2009: The Miracle of Deserts" | |||||||
Valid |
|
Aves
[edit]Research
[edit]- Anfinson, O.A.; Lockley, M.G.; Kim, S.H.; Kim, K.S.; Kim, J.Y. (2009). "First report of the small bird track Koreanaornis from the Cretaceous of North America: implications for avian ichnotaxonomy and paleoecology" (PDF). Cretaceous Research. 30 (4): 885–894. Bibcode:2009CrRes..30..885A. doi:10.1016/j.cretres.2009.02.001.
- Lockley, M.; Chin, K.; Houck, K.; Matsukawa, M.; Kukihara, R. (2009). "New interpretations of Ignotornis, the first-reported Mesozoic avian footprints: implications for the paleoecology and behavior of an enigmatic Cretaceous bird" (PDF). Cretaceous Research. 30 (4): 1041–1061. Bibcode:2009CrRes..30.1041L. doi:10.1016/j.cretres.2009.04.001.
- Bell, A.; Everhart, M.J. (2009). "A new specimen of Parahesperornis (Aves: Hesperornithiformes) from the Smoky Hill Chalk (Early Campanian) of western Kansas". Transactions of the Kansas Academy of Science. 112 (1/2): 7–14. doi:10.1660/062.112.0202. S2CID 86083098.
Newly described birds
[edit]Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images | |
---|---|---|---|---|---|---|---|---|---|
Gen. nov. et Sp. nov. |
Valid |
Placed in ?Patagopterygiformes by Agnolin and Martinelli. |
|||||||
Gen. nov. et Sp. nov. |
Valid |
Late Oligocene |
An Anatidae. |
||||||
Sp. nov. |
Valid |
Early Pleistocene |
Lower Elefante, TE 13 layer |
A Phasianidae. |
|||||
Sp. nov. |
Valid |
Late Miocene |
Originally described as a member of Scolopacidae belonging to the genus Calidris;[61] Zelenkov, Volkova and Gorobets (2016) reinterpreted it as a member of Turnicidae and transferred it to the genus Ortyxelos.[62] |
||||||
Sp. nov. |
Valid |
Middle Miocene |
A Procellariidae, the smallest of the genus. |
||||||
Sp. nov. |
Disputed |
Early-Middle Miocene |
Estancia La Costa Member, |
Federico L. Agnolin, 2009, made it the type species of the separate genus Noriegavis Agnolin, 2009.[65] Originally interpreted as a seriema; Noriega & Mayr (2017) reinterpreted it as a member of the falconid genus Thegornis of uncertain specific assignment, on the basis of a reexamination of the holotype specimen.[66] |
|||||
Sp. nov. |
Valid |
Late Miocene |
A Scolopacidae. |
||||||
Sp. nov. |
Valid |
Middle Miocene |
MN 6-8 |
An Anatidae. |
|||||
Sp. nov. |
Disputed |
A member of the family Confuciusornithidae. Considered to be a junior synonym of Confuciusornis sanctus by Wang, O'Connor & Zhou (2018).[69] |
|||||||
Sp. nov. |
Valid |
Middle Miocene |
MN 7-8 |
An Anatidae. |
|||||
Sp. nov. |
Valid |
Early Oligocene |
A Diomedeoididae Fischer, 1985, placed in Rupelornis van Beneden, 1871 by Gerald Mayr and Thierry Smith, 2012 |
||||||
Sp. nov. |
Disputed |
Late Miocene |
Originally described as a member of Ardeidae and a species of Egretta; however, Zelenkov (2017) considered this species to be a junior synonym of the barn-owl species Tyto campiterrae Jánossy (1991).[72] |
||||||
Gen. nov. et Sp. nov. |
Valid |
An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002, this is the type species of the new genus. |
|||||||
Gen. nov. et Sp. nov. |
Valid |
Riverleigh World Heritage Property |
An Anseranatidae, this is the type species of the new genus. |
||||||
Gen. nov. et Sp. nov. |
Valid |
Late Miocene |
Upper Liushu Formation |
An Accipitridae, this is the type species of the new genus. |
|||||
Sp. nov. |
Valid |
Late Pliocene |
MN 15-16 |
||||||
Sp. nov. |
Valid |
Late Miocene |
MN 11-12 |
||||||
Gen. nov. et Sp. nov. |
Valid |
A basal Ornithurae Haeckel, 1866, this is the type species of the new genus. |
|||||||
Sp. nov. |
Valid |
An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002. |
|||||||
Sp. nov. |
Valid |
An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002. |
|||||||
Sp. nov. |
Valid |
An Enantiornithes Walker, 1981, Euenantiornithes Chiappe, 2002. |
|||||||
Gen. nov. et Sp. nov. |
Valid |
Early Pliocene |
A Phasianidae, this is the type species of the new genus. |
||||||
Sp. nov. |
Valid |
Late Miocene |
A Phoenicopteriformes, Palaelodidae Stejneger, 1885. |
||||||
Sp. nov. |
Valid |
Middle Miocene |
MN 6-8 |
An Anatidae. |
|||||
Gen. nov. et Sp. nov. |
Valid |
Late Miocene |
Breda Formation Mill |
An Alcidae. |
|||||
Sp. nov. |
Valid |
Late Miocene |
Breda Formation Mill |
|
An Alcidae. |
||||
Sp. nov. |
Valid |
Late Miocene |
An Alcidae. |
||||||
Sp. nov. |
Valid |
Late Pliocene |
A Phasianidae. |
||||||
Gen. nov. et Sp. nov. |
Valid |
A stem Coraiiformes. |
|||||||
Sp. nov. |
Valid |
Late Oligocene |
An Anatidae. |
||||||
Gen. nov. et Sp. nov. |
Valid |
Late Oligocene |
An Anatidae, this is the type species of the new genus. |
||||||
Sp. nov. |
Valid |
Late Oligocene |
An Anatidae. |
||||||
Sp. nov. |
Valid |
Late Pliocene |
MN 15-16 |
||||||
Sp. nov. |
Valid |
Late Miocene |
A member of the family Rallidae. Originally described as a species Porzana; Zelenkov (2017) transferred this species to the genus Zapornia.[72] |
||||||
Sp. nov. |
Valid |
Middle Miocene |
MN 6-8 |
A Rallidae. |
|||||
Sp. nov. |
Valid |
Middle Eocene |
A Zygodactylidae Brodkorb, 1971. |
||||||
Gen. nov. et Sp. nov. |
Valid |
Late Miocene |
An Alcidae, this is the type species of the new genus. |
||||||
Gen. nov. et Sp. nov. |
Valid |
An Enantiornithes Walker, 1981, Longipterygidae Zhang, Zhou, Hou et Gu, 2000, this is the type species of the new genus. |
|||||||
Gen. nov. et Sp. nov. |
Valid |
Early Miocene, |
A Columbidae, this is the type species of the new genus. |
||||||
Sp. nov. |
Valid |
A Sapeornithidae Zhou et Zhang, 2006. |
|||||||
Sp. nov. |
Valid |
Early Oligocene, MP ?23-24 |
A Tytonidae, Selenornithinae Mourer-Chauviré, 1987. |
||||||
Gen. nov. et Sp. nov. |
Valid |
An Enantiornithes Walker, 1981, Familia Longipterygidae Zhang, Zhou, Hou et Gu, 2000, this is the type species of the new genus. |
|||||||
Gen. nov. et Sp. nov. |
Valid |
An Anatidae, Kaua'i Mole Duck, a Hawaiian duck that probably lived a kiwi-like lifestyle, this is the type species of the new genus. |
|||||||
Gen. nov. et Sp. nov. |
Valid |
Middle Miocene |
A Phasianidae, this is the type species of the new genus. |
||||||
Sp. nov. |
Valid |
Middle Miocene |
A Phasianidae. |
Pterosaurs
[edit]- Lü, J. (2009). "A new non-pterodactyloid pterosaur from Qinglong County, Hebei Province of China". Acta Geologica Sinica (English Edition). 83 (2): 189–199. doi:10.1111/j.1755-6724.2009.00062.x. S2CID 247739978.
- Vullo, R.; Neraudeau, D. (2009). "Pterosaur remains from the Cenomanian (Late Cretaceous) Paralic Deposits of Charentes, Western France". Journal of Vertebrate Paleontology. 29 (1): 277–282. Bibcode:2009JVPal..29..277V. doi:10.1080/02724634.2009.10010380. S2CID 197536660.
Newly named pterosaurs | |||||||
---|---|---|---|---|---|---|---|
Name | Status | Authors | Age | Unit | Location | Notes | Images |
Valid |
|
Late Triassic |
A campylognathoidid with a wingspan of about 70 cm. |
||||
Valid |
|
Middle Jurassic |
A primitive long-tailed pterosaur related to Dorygnathus. |
||||
Valid |
|
Early Cretaceous |
A pterodactyloid known from a juvenile specimen whose fossils preserved details of the flight membrane and fur. |
||||
Valid |
|
Late Jurassic |
A primitive long tailed pterosaur. The type specimen shows evidence of the animal having broken its shin while alive. |
See also
[edit]- 2010 in archosaur paleontology
- 2011 in archosaur paleontology
- 2012 in archosaur paleontology
- 2013 in archosaur paleontology
Notes
[edit]References
[edit]- ^ Marinho, T.S.; Carvalho, I.S. (2009). "An armadillo-like sphagesaurid crocodyliform from the Late Cretaceous of Brazil". Journal of South American Earth Sciences. 27 (1): 36–41. Bibcode:2009JSAES..27...36M. doi:10.1016/j.jsames.2008.11.005.
- ^ Juan Martín Leardi; Diego Pol (2009). "The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia". Cretaceous Research. 30 (6): 1376–1386. Bibcode:2009CrRes..30.1376L. doi:10.1016/j.cretres.2009.08.002. hdl:11336/90736.
- ^ Magdalena Borsuk−Białynicka; Andriej G. Sennikov (2009). "Archosauriform postcranial remains from the Early Triassic karst deposits of southern Poland" (PDF). Paleontologica Polonica. 65: 283–328. Retrieved 13 August 2010.
- ^ Kellner, A.W.A.; Pinheiro, A.E.P.; Azevedo, S.A.K.; Henriques, D.D.R.; de Carvalho, L.B.; Oliveira, G.R. (2009). "A new crocodyliform from the Alcântara Formation (Cenomanian), Cajual Island, Brazil". Zootaxa. 2030: 49–58. doi:10.11646/zootaxa.2030.1.4.
- ^ Santiago, L.A.; and Andrés, L.A. (2009). "Nuevo género de Crocodylia del Eoceno medio de la Península Ibérica (Zamora, España): Duerosuchus piscator nov. gen., nov. sp". Studia Geologica Salmanticensia. 45 (2): 149–173.
- ^ Butler, R.J.; Barrett, P.M.; Abel, R.L.; Gower, D.J. (2009). "A Possible Ctenosauriscid Archosaur from the Middle Triassic Manda Beds of Tanzania". Journal of Vertebrate Paleontology. 29 (4): 1022–1031. Bibcode:2009JVPal..29.1022B. doi:10.1671/039.029.0404. S2CID 86267617.
- ^ a b Sereno, P. C.; Larsson, H. C. E. (2009). "Cretaceous crocodyliforms from the Sahara". ZooKeys (28): 1–143. doi:10.3897/zookeys.28.325.
- ^ a b Cau, Andrea; Maganuco, Simone (2009). "A new theropod dinosaur, represented by a single unusual caudal vertebra from the Kem Kem Beds (Cretaceous) of Morocco". Atti Soc. It. Sci. Nat. Museo Civ. Stor. Nat. Milano. 150 (II): 239–257.
- ^ Lio, G., Agnolin, F., Cau, A. and Maganuco, S. (2012). "Crocodyliform affinities for Kemkemia auditorei Cau and Maganuco, 2009, from the Late Cretaceous of Morocco." Atti della Società Italiana di Scienze Naturali e del Museo di Storia Naturale di Milano, 153 (I), s. 119–126.
- ^ Lauprasert, K.; Cuny, G.; Thirakhupt, K. and Suteethorn, V. 2009. Khoratosuchus jintasakuli gen. et sp. nov., an advanced neosuchian crocodyliform from the Early Cretaceous (Aptian-Albian) of NE Thailand. In E. Buffetaut, G. Cuny, J. Le Loeuff, V. Suteethorn (eds.), Late Palaeozoic and Mesozoic Ecosystems in SE Asia. The Geological Society of London, Special Publication 315:175-187.
- ^ Erin L. Rasmusson Simons; Gregory A. Buckley (2009). "New Material of "Trematochampsa" Oblita (Crocodyliformes, Trematochampsidae) from the Late Cretaceous of Madagascar". Journal of Vertebrate Paleontology. 29 (2): 599–604. Bibcode:2009JVPal..29..599R. doi:10.1671/039.029.0224. S2CID 140171041.
- ^ Iori, F.V.; Carvalho, I.S. (2009). "Morrinhosuchus luziae, um novo Crocodylomorpha Notosuchia da Bacia Bauru, Brasil". Revista Brasileira de Geociências. 39 (4): 717–725. doi:10.25249/0375-7536.2009394717725. Archived from the original on 6 July 2011. Retrieved 7 August 2010.
- ^ Shan, Hsi-yin; Wu, Xiao-chun; Cheng, Yen-nien; Sato, Tamaki (2009). "A new tomistomine (Crocodylia) from the Miocene of Taiwan". Canadian Journal of Earth Sciences. 46 (7): 529–555. Bibcode:2009CaJES..46..529S. doi:10.1139/E09-036. Retrieved 15 September 2010.
- ^ Brusatte, Stephen L.; Butler, Richard J.; Sulej, Tomasz; Niedźwiedzki, Grzegorz; Brusatte, Stephen L.; Butler, Richard J.; Sulej, Tomasz; Niedźwiedzki, Grzegorz (2009). "The taxonomy and anatomy of rauisuchian archosaurs from the Late Triassic of Germany and Poland" (PDF). Acta Palaeontologica Polonica. 54 (2): 221–230. doi:10.4202/app.2008.0065. S2CID 54044937. Retrieved 13 August 2010.
- ^ Novas, F. E.; Pais, D. F.; Pol, D.; Carvalho, I. D. S.; Mones, A.; Scanferla, A.; Riglos, M. S. (2009). "Bizarre notosuchian crocodyliform with associated eggs from the Upper Cretaceous of Bolivia". Journal of Vertebrate Paleontology. 29 (4): 1316–1320. Bibcode:2009JVPal..29.1316N. doi:10.1671/039.029.0409. hdl:11336/54001. S2CID 140589478.
- ^ Nesbitt, S.J.; Turner, A.H.; Spaulding, M.; Conrad, J.L.; Norell, M.A. (2009). "The theropod furcula". Journal of Morphology. 270 (7): 856–879. doi:10.1002/jmor.10724. PMID 19206153. S2CID 11847975.
- ^ Barrett, P.M.; Han, F.-L. (2009). "Cranial anatomy of Jeholosaurus shangyuanensis (Dinosauria: Ornithischia) from the Early Cretaceous of China". Zootaxa. 2072: 31–55. doi:10.11646/zootaxa.2072.1.2.
- ^ Olshevsky, George. "Dinogeorge's Dinosaur Genera List". Archived from the original on 2011-07-15. Retrieved 2009-01-05.
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