Y-DNA haplogroups in populations of the Caucasus
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Various Y-DNA haplogroups have differing frequencies within each ethnolinguistic group in the Caucasus region.
Table
[edit]The table below lists the frequencies – identified by major studies – of various haplogroups amongst selected ethnic groups from the Caucasus. The first two columns list the ethnic and linguistic affiliations of the individuals studied, the third column gives the sample size studied, and the other columns give the percentage of the particular haplogroup.
Language family abbreviations:
IE Indo-European
NEC Northeast Caucasian
NWC Northwest Caucasian
Kartvelian South Caucasian
| Population | Language | n | E1b1b | G | I | J1 | J2 | L | R1a | R1b | R2a | T | Others | Reference |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| Abaza (Abazins/Abazinians) | NWC (Abkhaz-Ubykh) | 14 | 0 | 29 | 0 | See "Others". | 7 | See "Others". | 14 | 0 | 0 | See "Others". | K[xP]=14 (possible L or T)[1] C*=7 (1/14) F[xI,G,J2,K]=29 (possible J1) |
Nasidze 2004a[2] |
| Abaza (Abazins/Abazinians) | NWC (Abkhaz-Ubykh) | 88 | 4.5 | 40.9 | I2*=1.1 I2a=2.3 |
J1*=2.3 J1e=3.4 |
M67=1.1 other=10.2 |
2.3=L2 | 23.9 | 3.4 | 1.1 | 0.0 | 3.4=Q | Yunusbayev 2012[3] |
| Abkhaz | NWC (Abkhaz-Ubykh) | 12 | 0 | 0 | 33 | 0 | 25 | 0 | 33.0 | 8 | 0 | 0 | Nasidze 2004a[2] | |
| Abkhaz | NWC (Abkhaz-Abaza) | 58 | 1.7 | 56.9[4] | 0 | 0 | J2*=5.2 J2-M67=8.6 |
3.4=L2 | 10.3 | 12.1 | 1.7 | Balanovsky 2011[5] | ||
| Abkhaz | NWC (Abkhaz-Abaza) | 162 | 0.6 | 47.5 | 3.1 (P37.2=0.6, M223=1.9) | 3.1 (P58=1.2) | 26.5 (M67=11.1) | 2.5 | 13.0 | 7.4 | 0 | 0 | N=0.6 | Yunusbaev2012[6] |
| Adygei (Kabardin) | NWC (Adigei) | 59 | 0 | 28.8 | 10.2 | See "Others". | 11.9 | See "Others". | 1.7 | 1.7 | 0 | See "Others". | F[xG,I,J2,K]=23.7 (possible J1) K[xP]=15.3 (possible L) P[xR1,R2]=6.8 |
Nasidze 2004a[2] |
| Adygei (Kabardin) | NWC (Adygei) | 140 | 2.1 | 43.6 | I1=1.4 I2*=0.7 I2a=2.1 |
J1e=2.8 J1*=6.4 |
M67=5.7 J2a*=9.3 J2b=0.7 |
0.7=L3 | 15.0 | 3.6 | 0.0 | 0.7 | C=2.1, H=0.7 N1c=1.4, Q=0.7 |
Yunusbayev 2012[3] |
| Adygei/ "Circassians"[a] |
NWC (Adygei) | 0.0[7] | 31.3[8] | I*=1.4 I2a=2.9[9] |
various | |||||||||
| Adygei/ ("Adygea") |
NWC (Adygei) | 154 | 0.6 | 47.4 | I2*=1.3 I2a=3.2 |
J1e=1.3 J1*=1.9 |
M67=3.2 J2a*=5.8 J2b=1.9 |
L2=1.9 L1=0.6 |
14.3 | 7.1 | 0.0 | 0.0 | C=2.6 N=0.6 |
Yunusbayev 2012[3] |
| Adygei (Shapsugs) | NWC (Adygei) | 106 | 81.1 | Dibirova 2009[8] | ||||||||||
| Adygei (Shapsugs) | NWC (Adygei) | 100 | 1.0 | 87.0[10] | 0 | 0 | 6.0 | 2.0=L* | 4.0 | 0 | 0 | Balanovsky 2011[5] | ||
| Adygei ("Cherkess")[b] |
NWC (Adygei) | 142 | 1.4 | 39.8[11] | 0.7=I* | J1e=0.7 J1*=4.9 |
J2-M67=7.7 J2b=0.7 other=13.4 |
0.7=L1 | 19.7 | 4.9 | 0.7 | N1*=1.4 N1c1=2.1 Q=0.7 |
Balanovsky 2011[5] | |
| Adygei ("Cherkess") |
NWC (Adygei) | 126 | 0.8 | 45.2 | I1=0.8 I2b=0.8 |
J1e=4.0 J1*=0.8 |
M67=13.5 other=11.1 |
0.0 | 15.1 | 1.6 | 0.8 | 0.0 | C=0.8, K*=1.6 N1c1=2.4, Q=0.8 |
Yunusbayev 2012[3] |
| Andis | NEC (Avar-Andic, Andic) | 49 | 2.0 | 6.1 | I1=2.0 I2a=24.5 |
J1*=36.7 | M67=14.3 other=4.1 |
0.0 | 2.0 | 6.1 | 0.0 | 2.0 | Yunusbayev 2012[3] | |
| Armenians | IE (Armenian) | 89 | 3.4 | 29.2 | 5.6 | 24.7 | 3.4 | Rosser 2000[12] | ||||||
| Armenians | IE (Armenian) | 47 | 4.3 | F* | 4.3 | F* | 21.3 | 4.3 | 8.5 | 36.2 | 0 | 6.4 | F[xI,J2,K]=12.8 N=2 |
Wells 2001[13] |
| Armenians | IE (Armenian) | 100 | 6.0 | 11.0 | 5.0 | See "Others". | 24.0 | See "Others". | 6.0 | 19.0 | 2.0 | 4.0[1] | F[xI,G,J2,K]=18.0 (possible J1) K[xT,P]=3.0 (possible L)[1] P[xR1a,R1b,R2]=2.0 |
Nasidze 2004a[2] |
| Armenians | IE (Armenian) | 734 | 5.4 | 5.3 | 32.4 | 1.6 | Weale 2001[14] | |||||||
| Armenians (TOTAL) |
IE (Armenian) | 413 | 5.1 | P16= 0.5 G2a*= 8.2 G1=0.7 |
I2=3.6 | J1e= 4.4 J*=6.3 |
M67= 10.7 J2a* = 13.3 J2b= 1.5 |
1.9 | 1.7 | 29.1 | 4.6 | 8.5 | N=0.2 Q=0.2 |
Herrara2012 |
| Armenians (Ararat Valley) |
IE (Armenian) | 110 | 5.5 | P16=0 G2a*=9.1 G1=1.8 |
I2=2.7 | J1e=6.4 J1*=8.2 |
M67=12.7 J2a*=10.0 J2b=0.9 |
0.9 | 0.9 | 37.3 | 0.0 | 3.6 | 0.0 | Herrara2012 |
| Armenians (Sason) | IE (Armenian) | 104 | 2.9 | P16= 1.9 G2a*=10.6 |
0.0 | J1e= 2.9 J1* = 6.7 |
M67= 7.7 J2a* = 9.6 J2b = 0 |
3.8 | 1.0 | 15.4 | 17.3 | 20.2 | Herrara2012 | |
| Avars | NEC (Avar-Andic, Avar) | 42 | 7.1 | 0.0 | 0.0 | 66.7 | 4.8 | 9.5 | 2.4 | 2.4 | 2.4 | 4.8 | Yunusbayev 2012[3] | |
| Avars | NEC (Avar-Andic, Avar) | 115 | 0.0 | P18=0.9 P303=9.6 |
I*=0.9 I2a=0.9 |
J1e=0.9 J1*=58.3 |
M67=0.9 other=5.2 |
L2=2.6 | 1.7 | 14.8 | 0.9 | 0.0 | N=1.7 | Balanovsky 2011[5] |
| Avars (West)[c] |
NEC (Avar-Andic) | 20 | 0 | G[xG1,G2a, G2b]=5 |
0 | J1*=60 J1e=20 |
10 | 0 | 0 | 5 | Caciagli 2009[15] | |||
| Azeris (Azerbaijan) |
Turkic | 72 | 5.6 | 18.1 | 2.8 | F | 30.6 | 6.9[1] | 6.9 | 11.1 | 2.8 | 4.2[1] | F[xG,I,J2,K]=11 | Nasidze 2004a[2] |
| Azerbaijanis | Turkic | v | 4.1[7] | 15.2[16] | 23.9[16] | various | ||||||||
| Bagvalins | NEC (Avar-Andic, Andic) | 28 | 0.0 | 0.0 | I2b1=7.1 | 21.4 | 0.0 | 0.0 | 3.6 | 67.9 | 0.0 | 0.0 | Yunusbayev 2012[3] | |
| Balkars | Turkic | 38 | 2.6 | 28.9 | I2*=2.6 | 0 | M67=5.3 J2b=2.6 other=15.8 |
L2=5.3 | 13.2 | 13.2 | 7.9 | 0.0 | H=2.6 | Battaglia 2009[17] |
| Balkars | Turkic | 135 | 0 | 32.6 | 3.0 (all P37.2) | 3.7 (P58=0.0) | 15.6 (M67=9.6) | 0 | 28.1 | 13.3 | 0 | 0 | Q=3.7%, no others. | Yunusbaev 2012[6] |
| Chamalins | NEC (Avar-Andic, Andic) | 27 | 0.0 | 18.5 | 0.0 | 66.7 | 3.7 | 3.7 | 7.4 | 0.0 | 0.0 | 0.0 | Yunusbayev 2012[3] | |
| Chechens (East Chechnya) |
NEC (Nakh) | 19 | 0 | 5 | 0 | 26 | 5 | 5 | 0 | 16 | 5[1] | F[xG,I,J2,K]=32 | Nasidze 2004a[2] | |
| Chechens (Total)[d] | NEC (Nakh) | 330 | 0.0 | 5.4[18] | I2=0.3 | J1*=20.9 | M67=55.2 J2b=0.3 other=1.2 |
L3=7.0 | 3.9 | 1.8 | 3.3 | 0.0 | Balanovsky 2011[5] | |
| Chechens (Achkhoy-Martan, Chechnya) | NEC (Nakh) | 118 | 0 | 0.8 (P303) | I2=0.8 | J1*=24.6 | M67=56.8 other=0.8 |
L3=6.8 | 2.5 | 3.4 | 3.4 | 0 | Balanovsky 2011[5] | |
| Chechens (Malgobek, Ingushetia) |
NEC (Nakh) | 112 | 0 | 9.0[19] | 0 | 21.4 | M67=50.9 J2b=0.9 other=1.8 |
L3=0.9 | 8.0 | 0.9 | 6.3 | 0 | Balanovsky 2011[5] | |
| Chechens (mainly Akkis, in Dagestan) |
NEC (Nakh) | 100 | 0 | 7.0[20] | 0 | 16.0 | M67=58.0 other=1.0 |
L3=14.0 | 1.0 | 1.0 | 0 | 0 | N=2.0 | Balanovsky 2011[5] |
| Chechens | NEC (Nakh) | 165 | 1.2 | 1.8 | 0.0 | 24.2 P58=0.0 |
48.5 M67=46.7 |
17.6 | 0.6 | 0.6 | 0.0 | 0.0 | N=0.6 Q=4.8 |
Yunusbaev 2012[6] |
| Dargins | NEC (Dargin) | 68 | 0 | 2.9 | 0 | 91.2 | 2.9 | 0.0 | 0.0 | 2.9 | 0.0 | 0.0 | Yunusbayev 2012[3] | |
| Dargins | NEC (Dargin) | 26 | 4 | 4 | 58 | See "Others". | 4 | 0 | 0 | 4 | 0 | 0 | F[xG,I,J2,K]=27 | Nasidze 2004a[2] |
| Dargins (Dargwa) | NEC (Dargin) | 101 | 0 | P303=1.0 other=1 |
0 | J1*=69.3 J1e=1 |
M67=1.0 | 0 | 21.8 | 2.0 | 0 | 0 | O3=3 | Balanovsky 2011[5] |
| Dargins (Kaitaks) | NEC (Dargin) | 33 | 0 | 0 | 0 | J1*=84.8 | 3.3 | 0 | 3.3 | 6.7 | 3.3 | 0 | Balanovsky 2011[5] | |
| Dargins (Kubachis) | NEC (Dargin) | 65 | 0 | 0 | I2a=1.5 | J1*=98.5 | 0 | 0 | 0 | 0 | 0 | 0 | Balanovsky 2011[5] | |
| Georgians | Kartvelian (Karto-Zan) | 63 | 2.0 | 30.1 | 0.0 | J=36.5 | 1.6 | 7.9 | 14.3 | 6.3 | 1.6 | Semino 2000[21] | ||
| Georgians | Kartvelian (Karto-Zan) | 66 | 3.0 | 31.8 | I1*=1.5 | 4.5 | M67=18.2 other=13.6 |
L3=1.6 | 10.6 | 9.1 | 4.5 | 1.6 | Battaglia 2009[17] | |
| Georgians | Kartvelian (Karto-Zan) | 77 | 2.6 | 31.2 | 3.9 | F | 20.8 | 2.6[1] | 10.4 | 10.4 | 1.3 | 2.6[1] | K[xP]=2.6 P[xR1,R2]=2.6 F[xG,I,J2,K]=14.3 |
Nasidze 2004a[2] |
| Svans | Kartvelian (Svan) | 25 | 0 | F* | 0 | F* | 0 | 0 | 8 | 0 | 0 | 0 | F[xG,I,J2,K]= 92% | Wells 2001[13] |
| Georgians (Kazbegis) | Kartvelian (Karto-Zan)[e] | 25 | 0 | F | 4 | F | 72 | 0 | 4 | 8 | 0 | 0 | F[xG,I,J2,K]=12 | Wells 2001[13] |
| Ingush | NEC (Nakh) | 143 | 0 | P18=1.4 | 0.7 | 2.8 | M67=87.4 other=1.4 |
L3=2.8 | 3.5 | 0 | 0 | Balanovsky 2011[5] | ||
| Ingush | NEC (Nakh) | 22 | 0 | 5.0 | 5 | 0 | 32 | 5 | 0 | 0 | 0 | 0 | F[xG,I,J2,K]=27 P[xR1,R2]=4.5(1/22) |
Nasidze 2004a[2] |
| Ingush | NEC (Nakh) | 105 | 0.0 | 4.8 | 0.0 | 1.9 | 81.9 (all M67) | 8.6 | 2.9 | 0 | 0 | 0 | Yunusbaev 2012 [6] | |
| Juhurim | AA (Semitic) | 10 | 0 | 0 | 0 | 30 (all P58) | 50 (M67=10%) | 20 | 0 | 0 | 0 | 0 | Yunusbaev 2012[6] | |
| Karachays | Turkic | 69 | 0 | 31.9 | 8.7 (P37.2=7.2, M223=1.5) |
7.2 (P58=2.9) |
11.6 (M67=5.8) |
0 | 27.5 | 10.1 | 0 | 2.9 | 0.0 | Yunusbaev 2012[6] |
| Kumyks | Turkic | 76 | 2.6 | 11.8 | 0.0 | 21.1 | 25.0 | 0.0 | 13.2 | 19.7 | 3.9 | 1.3 | J*=1.3 O=1.3 |
Yunusbaev 2012[6] |
| Laks | NEC (Lak) | 21 | 9.5 | 4.8 | 14.3 | 42.9 | 14.3 | 0.0 | 9.5 | 4.8 | 0.0 | 0.0 | Caciagli 2009[15] | |
| Lezgins (Azerbaijan) | NEC (Lezgic) | 12 | 17 | 0 | See "Others". | 0 | 8 | 17 | 0 | F[xI,J2,K]=58 | Wells 2001[13] | |||
| Lezgins (Dagestan) | NEC (Lezgic) | 25 | 0 | 36 | 0 | See "Others". | 0 | 0 | 4 | 0 | See "Others". | F[xI,G,J2,K]=32 (possible J1) K[xR]=28 |
Nasidze 2003[1] | |
| Lezgins | NEC (Lezgic) | 31 | 6.5 | 9.7 | 9.7 | 58.1 | 0.0 | 0.0 | 0.0 | 16.1 | 0.0 | 0.0 | Yunusbaev 2012[6] | |
| Lezgins (Akhtynskiy District) | NEC (Lezgic) | 81 | 1.2 | 13.5 (P18=1.2%) |
0 | 44.4 J1* | 2.5 | 1.2 L2 | 3.7 | 29.6 | 0 | 2.5 | 1.2 N1c1 | Balanovsky 2011[5] |
| Kara Nogai | Turkic | 76 | 0 | 1.3 | 13.2 I-P37.2 | 2.6 J* | 10.5 (M67 = 1.3%) |
0 | 17.1 | 18.4 | 0 | 0 | C=10.5 D=5.3 N=14.5 O=5.3 Q=1.3 |
Yunusbayev 2012[3] |
| Kuban Nogai | Turkic | 87} | 0 | 13.8 | 0 | 21.8 | 16.1 M67=3.4% |
0 | 12.6 | 17.2 | 0 | 1.1 | C=8.0% D=1.1% N=4.6% O=3.4% Q=0.0% |
Yunusbayev 2012[3] |
| Ossetes | IE (Iranian, NE) | 47 | 6.4 | J=34.0 | 2.1 | 42.6 | Rosser 2000[12] | |||||||
| North Ossetians | IE (Iranian, NE) | 129 | 0 | 57.4 | 10.1 | See "Others". | 14.0 | 0.8 | 0.8 | 1.6 | See "Others". | F[xI,G,J2,K]=3.9 (possible J1) K[xP]=9.3 (possible L) P[xR1,R2]=2.3 |
Nasidze 2004b[22] | |
| North Ossetians (Iron) | IE (Iranian, NE) | 230 | 0.4 | 74.3[23] | 0 | 1.3 | M67=11.3 other=7.0 |
L2=0.9 | 0.4 | 3.0 | 0 | N=0.4 Q=0.9 |
Balanovsky 2011[5] | |
| North Ossetians (Digor) | IE (Iranian, NE) | 127 | 0.8 | 60.6[24] | 0 | 3.9 | M67=5.5 other=6.3 |
L1=0.8 | 0.8 | 16.5 | 0.8 | Q=3.9 | Balanovsky 2011[5] | |
| North Ossetians | IE (Iranian, NE) | 132 | 1.5 | 69.7 | 0.0 | 3.8 | 18.2 (M67=9.8) | 0.8 | 0.8 | 4.5 | 0 | 0 | Q = 0.8 | Yunusbaev 2012[6] |
| Russians (Adygea) | IE (Slavic, East) | 78 | 24.4 | Rootsi 2004[9] | ||||||||||
| Rutuls | NEC (Lezgic) | 24 | 0.0 | 37.5 | 0.0 | F | 4.2 | 0.0 | 0.0 | 0.0 | 0.0 | 0.0 | F[xG,I,J2,K]=58 | Nasidze 2004a[2] |
| Tabassarans | NEC (Lezgic) | 43 | 0.0 | 0.0 | 0.0 | 48.8 | 2.3 | 0.0 | 2.3 | 39.5 | 0.0 | 0.0 | C=7.0 | Yunusbaev 2012[6] |
See also
[edit]- Caucasus
- Y-DNA haplogroups by groups
- Y-DNA haplogroups in populations of Europe
- Y-DNA haplogroups in populations of the Near East
- Y-DNA haplogroups in populations of North Africa
- Y-DNA haplogroups in populations of Central and North Asia
- Y-DNA haplogroups in populations of South Asia
- Y-DNA haplogroups in populations of Sub-Saharan Africa
- Y-DNA haplogroups in populations of East and Southeast Asia
- Y-DNA haplogroups in populations of Oceania
- Y-DNA haplogroups in indigenous peoples of the Americas
Notes
[edit]- ^ despite the fact that Shapsugs were also technically "Circassian", Dibirova labels this population "Circassians". In Rootsi, they are called "Adygei", or some variation of the word.
- ^ These are labeled "Circassians" (at least in the English version), but the sample was taken from the Karacheyo-Cherkess Republic, so they are labeled "Cherkess" here.
- ^ This is significant, as in Western Avaria, where the population was sampled, there are large number of non-Avars- Andic and Dido peoples- who are in fact ethnically misclassified as Avars by the Dagestani census.
- ^ Made by combining the Malgobek, Achkhoy-Martan and Khasavyurt Chechen populations
- ^ Although the Kazbegi Georgians speak Georgian, and have most likely done so for centuries, it is theorized by some that their (possible) ancestors, the Tzanars, who inhabited the region in Medieval and Ancient times, were a Nakh-speaking people, like Chechens and Ingush, possibly accounting for their large frequencies of J2
References
[edit]- ^ a b c d e f g h i Nasidze 2003
- ^ a b c d e f g h i j Nasidze 2004a
- ^ a b c d e f g h i j k Yunusbayev 2012
- ^ P18=12.1
P303=20.7
other=24.1 - ^ a b c d e f g h i j k l m n o Balanovsky 2011
- ^ a b c d e f g h i j CITEREFYunusbayev2012
- ^ a b Cruciani 2004
- ^ a b Dibirova 2009
- ^ a b Rootsi 2004
- ^ P303=86.0; other G =1.0
- ^ P16=9.2
P303=29.6; other=1.4. - ^ a b Rosser 2000
- ^ a b c d Wells 2001
- ^ Weale 2001
- ^ a b Caciagli 2009
- ^ a b Di Giacomo 2004
- ^ a b Battaglia 2009
- ^ P18=3.0
P303=2.4 - ^ P18=3.6
P303=5.4 - ^ P18=6.0
P303=1.0 - ^ Semino 2000
- ^ Nasidze 2004b
- ^ P16=73.0
P303=1.3 - ^ P16=55.9
P303=4.7
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