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Ancient North Eurasian
Mal'ta–Buret' culture ivory figurines (c. 24,000 BP-c. 15,000 BP). Some of the figurines wear hooded overalls with decorative stripes.[1] [2]
Approximate location of the Ancient North Eurasians c. 24,000~16,000 BP.[3][4][5]

In archaeogenetics, the term Ancient North Eurasian (ANE) is the name given to an ancestral component that represents the lineage of the people of the Mal'ta–Buret' culture (c. 24,000 BP) and populations closely related to them, such as the Upper Paleolithic individuals from Afontova Gora in Siberia.[6][7] Genetic studies also revealed that the ANE are closely related to the remains of the preceding Yana culture (c. 32,000 BP), which were named Ancient North Siberians (ANS). Ancient North Eurasians are predominantly of West Eurasian ancestry (related to European Cro-Magnons and ancient and modern peoples in West Asia) who arrived in Siberia via the "northern route", but also derive a significant amount of their ancestry (c. 1/3) from an East Eurasian source, having arrived to Siberia via the "southern route".[8][9][10][11][12][13][14][15][16][17][a][18]

Around 20,000 to 25,000 years ago, a branch of Ancient North Eurasian people mixed with Ancient East Asians, which led to the emergence of Ancestral Native American, Ancient Beringian and Ancient Paleo-Siberian populations. It is unknown exactly where this population admixture took place, and two opposing theories have put forth different migratory scenarios that united the Ancient North Eurasians with ancient East Asian populations.[19]

Later, ANE populations migrated westward into Europe and admixed with European Western hunter-gatherer (WHG)-related groups to form the Eastern Hunter-Gatherer (EHG) group, which later admixed with Caucasus hunter-gatherers to form the Western Steppe Herder group, which became widely dispersed across Eurasia during the Bronze Age.[20]

ANE ancestry has spread throughout Eurasia and the Americas in various migrations since the Upper Paleolithic, and more than half of the world's population today derives between 5 and 42% of their genomes from the Ancient North Eurasians.[21] Significant ANE ancestry can be found in Native Americans, as well as in Europe, South Asia, Central Asia, and Siberia. It has been suggested that their mythology may have featured narratives shared by both Indo-European and some Native American cultures, such as the existence of a metaphysical world tree and a fable in which a dog guards the path to the afterlife.[22]

Genetic studies

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Definition

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The ANE lineage, also known as Paleolithic Siberians, is defined by association with the "Mal'ta boy" (MA-1), the remains of an individual who lived during the Last Glacial Maximum, 24,000 years ago in central Siberia, discovered in the 1920s. Together with the Yana Rhinoceros Horn Site samples, and Afontova Gora individuals, they are collectively referred to as 'Ancient North Siberians', although 'Ancient North Eurasian' is also used as collective name for both MA-1 and Yana remains.[23][24][17]

The Mal'ta boy (MA-1), dated 24,000 BP, with tomb artifacts, Hermitage Museum (Hall 11),[25] Saint-Petersburg.[26]

The Ancient North Eurasians represent a Paleolithic Siberian cluster, more closely related to European hunter-gatherers than to East and Southeast Asian populations.[17][27][28] It is suggested that the ANE ancestry found among modern human populations was largely contributed from a population linked to Afontova Gora (AG2/3), rather than Malta (MA1) or Yana.[29]

Ancient North Eurasian associated Y-chromosome haplogroups are P-M45, and its subclades R and Q. Haplogroup P is inferred to have originated around 44,000 years ago in Southeast Asia and is downstream to Haplogroup K2b found among the Tianyuan man in Northern China. Their maternal haplogroup belonged to subclades of haplogroup U.[30][31][32][33][34]

Formation

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Human principal component analysis (PCA) with ancient human genomes projected on top.[35]
A qpGraph model by Maier et al. 2023, showing the possible formation of Ancient North Siberians/Eurasians (ANS/ANE) and their subsequent contribution to the Ancient Paleo-Siberians and Native Americans.[36]

The formation of the Ancient North Eurasian/Siberian (ANE/ANS) gene pool likely occurred very early by the Upper Paleolithic dispersal by the admixture of an 'Ancient West Eurasian' population via the 'northern route' through Central Asia into Siberia, with an 'Ancient East Eurasian' via the 'southern route'. The West Eurasian source was distantly related to the Upper Paleolithic remains in Europe, such as the Goyet specimen, as well as the Kostenki-14 and Sungir individuals, and ultimately expanded from a population hub in the Iranian Plateau. The 'East Eurasian' source can be associated with ancestry found in the 40,000 year old Tianyuan man of Northern China.[37][28][15][38][18]

Overall, Ancient North Eurasians are best described as admixture between an Ancient West Eurasian lineage (71%), with approximately 29% geneflow from an East Eurasian source.[18] Grebenyuk et al. argues that 'Ancient North Eurasians' were "Early Upper Paleolithic tribes of hunters" and linked to similar groups associated with Southern Siberian sites. These communities of Southern Siberian and Central Asian hunters belonged to one of the earliest migration waves of anatomically modern humans into Siberia. The authors summarized that "the initial peopling of Northeastern Asia by the anatomically modern humans could have happened both from West to East and from South to North".[37]

Studies on Ancient North Eurasian remains

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The ANE/ANS-associated samples from the Yana Rhinoceros Horn Site (31,600 BP) in Northeastern Siberia and the closely related remains from the Mal'ta–Buret' culture (MA1), and the individuals associated with the Afontova Gora site (AG2/3), can be modeled to derive their ancestry from an Ancient West Eurasian lineage, with significant amounts of admixture from an East Eurasian lineage (22–50%).[b][39][40][41][42][17][43]

Lipson and Reich (2017) modeled the Mal'ta sample to be derived from a West Eurasian source (82%), with additional admixture from a lineage related to East Asians (18%), while also noting the possibility for a reversed geneflow from Mal'ta into East Asians, which however had less support with the available data.[12] Yang et al. 2020 modeled both the Yana specimens and the Mal'ta sample as a merger of a sister lineage of the 'European hunter-gatherer' Kostenki-14, contributing around 68% ancestry, and of a lineage contemporary to the 'Basal-East Asian' Tianyuan man, contributing around 32% ancestry, while finding no evidence for a reversed geneflow from ANE/ANS into Tianyuan or modern East Asians.[44][45] Mao et al. 2021 models both Yana and Afontova Gora remains with around 73% West Eurasian and 27% East Eurasian ancestry.[46] Sikora et al. 2019 analyzed the genetic remains of the Yana Rhinoceros Horn Site and found them to be closely related to the Ancient North Eurasians. The collectively named both populations as Ancient North Siberian. They modeled the ANE/ANS to derived between 71–78% West Eurasian ancestry and between 22–29% East Eurasian ancestry.[47] Sikora et al. also notes that the Ancient North Eurasians (Malta and Afontova Gora individuals) are unlikely to be direct descendants of the 'Ancient North Siberian' Yana population; rather, the study argues, both are sister lineages sharing a common ancestor. According to Sikora et al., the Malta sample may additionally also have received some 'early Caucasus hunter-gatherer' geneflow (c. 11%).[47] This scenario is questioned by Maier et al. 2023, who state that this conclusion is contradicted by other published articles, and that the direction of gene flow as well as observed affinity between ANE and CHG populations cannot be demonstrated by analysis of admixture graphs, but need further investigation.[48] By using a newly developed version of ADMIXTOOLS, they estimate around 76% West Eurasian ancestry and 24% East Eurasian ancestry for both the Yana and Mal'ta remains.[49] Zhang et al. 2023 summarized that the Ancient North Siberians (Yana remains) are best described to derive 71% ancestry from a West Eurasian lineage and 29% ancestry from an East Eurasian lineage. The Yana remains are closely related to the Mal'ta and Afontova Gora remains, but not identical with them.[18] Vallini et al. 2022/2024 described the Ancient North Eurasians as falling into "an intermediate position between the two axes, the result of a palaeolithic admixture", deriving around 50% from West Eurasian and 50% from East Eurasian sources.[28][50] Allentoft et al. 2024 modeled the Yana specimens, the Mal'ta sample as well as the Afontova Gora remains as 65% West Eurasian and 35% East Eurasian.[51]

A different but geographically close specimen, known as the Salkhit individual (c. 34,000 BP) from Northern Mongolia was found to display a complex relation to the Yana individuals. While the Yana individuals derived between 25–33% of their ancestry from a Tianyuan-like source, the Salkhit individual derived around 25% ancestry from the Yana lineage and 75% from the Tianyuan lineage, suggesting bi-directional geneflow between Ancient West and East Eurasian populations in Northeastern Siberia.[36][28][16]

Distribution

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One of the "Mal'ta figurines" with facial features, and 3D rendering.[52]

By c. 32kya, populations carrying ANE-related ancestry were probably widely distributed across northeast Eurasia. They may have expanded as far as Alaska and the Yukon, but were forced to abandon high latitude regions following the onset of harsher climatic conditions that came with the Last Glacial Maximum.[53]

Populations genetically similar to MA-1 and Afontova Gora were an important genetic contributor to Native Americans, Europeans, Ancient Central Asians, South Asians, and some East Asian groups, in order of significance.[54] Lazaridis et al. (2016:10) note "a cline of ANE ancestry across the east-west extent of Eurasia". A 2016 study found that the global maximum of ANE ancestry occurs in modern-day Kets, Mansi, Native Americans, and Selkups.[6][54]

Deer tooth pendant of an ANE woman, from Denisova Cave, dated circa 24,700 years BP.[55]

The ancient Bronze-age-steppe Yamnaya and Afanasevo cultures were found to have a significant ANE-like component at c. 25–50% via their EHG and CHG ancestry.[56][57] According to Moreno-Mayar et al. 2018 between 14% and 38% of Native American ancestry may originate from gene flow from the Mal'ta–Buret' (ANE) population. This difference is caused by the penetration of posterior "Neo-Siberian" migrations into the Americas, with the lowest percentages of ANE ancestry found in Inuit and Alaskan Natives, as these groups are the result of migrations into the Americas roughly 5,000 years ago.[58] Estimates for ANE ancestry among first wave Native Americans show higher percentages,[59] such as 41% (36-45%) for those belonging to the Andean region in South America.[59] The other gene flow in Native Americans (the remainder of their ancestry) was of an East Asian-related origin, specifically diverged from other East Asians c. 30,000 years ago.[60] Gene sequencing of another south-central Siberian people (Afontova Gora-2) dating to approximately 17,000 years ago, revealed similar autosomal genetic signatures to that of Mal'ta boy-1, suggesting that the region was continuously occupied by humans throughout the Last Glacial Maximum.[60]

Genomic studies also indicate that the ANE component was brought to Western Europe by people related to the Yamnaya culture, long after the Paleolithic.[56][6] It is reported in modern-day Europeans (10%–20%).[56][6] Earlier ANE ancestry is found in European hunter-gatherer populations through Paleolithic interactions with Eastern European Hunter-Gatherers, which resulted in populations such as Scandinavian Hunter-Gatherers. [61] Western Hunter-Gatherers of the Villabruna cluster also carried the Y-haplogroup R1b, derived from the Ancient North Eurasian haplogroup R*, indicating "an early link between Europe and the western edge of the Steppe Belt of Eurasia."[15]

A deer tooth pendant impregnated with the genetic material of an ANE woman was found in the Denisova Cave, and dated to circa 24,700 years before present. She is closely related to Mal'ta and Afontova Gora specimens, found further east.[55]

An early Neolithic Central Asian specimen (Tutkaul1) from Tajikistan was found to be primarily derived from Ancient North Eurasians with some additional Neolithic Iranian-related inputs. The sample is closely related to Afontova Gora 3 (AG3) and Mal’ta 1, as well as to the West Siberian hunter-gatherers (Tyumen and Sosnoviy). While the sample also displays affinity for Eastern hunter-gatherers (EHGs), AG3 was found to be closer to EHGs than Tutkaul1, who instead may be a good proxy for ANE-related ancestry among ancient populations from the Iran and the Turan region.[20]

The Ancient Tianyuan Man and modern East/Southeast Asian populations were found to lack Upper Paleolithic Western Eurasian or ANE-related admixture, suggesting "resistance of those groups to the incoming UP population movements", or alternatively a subsequent reexpansion from a genetically East Asian-like population reservoir.[28]

Groups partially derived from the Ancient North Eurasians

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Native American contribution

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Main article: Genetic history of the Indigenous peoples of the Americas
The "Ancient North Eurasian" (ANE) network, consisted of several Paleolithic Siberian samples and contributed ancestry towards a wide variety of populations across Eurasia.
The map shows the origin of the first major wave of Native Americans. Involved are the ANE (Ancestral Northern Eurasian, which are related to Europeans) and the NEA (Northeast Asians, which are an East Asian-related people). The admixture happened somewhere in Northeast Siberia.[62]

According to Jennifer Raff, the Ancient North Eurasian population mixed with a daughter population of ancient East Asians, who they encountered around 25,000 years ago, which led to the emergence of Native American ancestral populations. However, the exact location where the admixture took place is unknown, and the migratory movements that united the two populations are a matter of debate.[19] Vallini et al. 2024 notes that the "position of Native Americans suggests a primarily East Asian ancestry, with a smaller contribution from palaeolithic West Eurasian populations".[50]

One theory supposes that Ancient North Eurasians migrated south to East Asia, or Southern Siberia, where they would have encountered and mixed with ancient East Asians. Genetic evidence from Lake Baikal in Mongolia supports this area as the location where the admixture took place.[63]

However, a third theory, the "Beringian standstill hypothesis", suggests that East Asians instead migrated north to Northeastern Siberia, where they mixed with ANE, and later diverged in Beringia, where distinct Native American lineages formed. This theory is supported by maternal and nuclear DNA evidence.[64] According to Grebenyuk, after 20,000 BP, a branch of Ancient East Asians migrated to Northeastern Siberia, and mixed with descendants of the ANE, leading to the emergence of Ancient Paleo-Siberian and Native American populations in Extreme Northeastern Asia.[65][37] However, the Beringian standstill hypothesis is not supported by paternal DNA evidence, which may reflect different population histories for paternal and maternal lineages in Native Americans, which is not uncommon and has been observed in other populations.[66]

The descendants of admixture between ANE and ancient East Asians include Ancient Beringian/Ancestral Native American, which are specific archaeogenetic lineages, based on the genome of an infant found at the Upward Sun River site (dubbed USR1), dated to 11,500 years ago.[67] The AB and the Ancestral Native American (ANA) lineage formed about 25,000 years ago, and subsequently diverged from each other, with the AB staying in the Beringian region, while the Ancestral Native Americans populated the Americas. The ANE genetic contribution to late-Paeolithic Ancestral Native Americans (USR1 specimen, dated to 11,500 BP in Alaska, and Clovis specimen, dated to 12,600 BP in Montana) is estimated at 36.8%.[68] There are also the Ancient Paleo-Siberians, populations represented by the Late Upper Paeolithic Lake Baikal Ust'Kyakhta-3 (UKY) 14,050-13,770 BP. They carried 30% ANE ancestry and 70% East Asian ancestry.[62]

Jomon people

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Jōmon people, the pre-Neolithic population of Japan, mainly derived their ancestry from East Asian lineages, but also received geneflow from the ANE-related "Ancient North Siberians" (represented by samples from the Yana Rhinoceros Horn Site) prior to the migration from the Asian mainland to the Japanese archipelago. Jōmon ancestry is still found among the inhabitants of present-day Japan: most markedly among the Ainu people, who are considered the direct descendants of the Jōmon people, and to a small, but significant degree among the majority of the Japanese population.[69][70]

Siberian and Asian Holocene populations

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Altai hunter-gatherer is the name given to Middle Holocene Siberian hunter-gatherers within the Altai-Sayan region in Southern Siberia. They originated from the admixture of Paleo-Siberian and Ancient North Eurasian groups and show increased affinity towards Native Americans. Bronze Age groups from North and Inner Asia with significant ANE ancestry (e.g. Lake Baikal hunter-gatherers, Okunevo pastoralists) can be successfully modeled with Altai hunter-gatherers as a proximal ANE-derived ancestry source.[71][72]

West Siberian Hunter-Gatherer (WSHG) is a specific archaeogenetic lineage that was first reported by Narasimhan et al. (2019). It can be modeled as 20% EHG, 73% ANE and 6% Ancient Northeast Asian. Although only represented by three sampled hunter-gatherer individuals from Tyumen Oblast in the Russian Forest Zone east of the Urals dated ca. 5,000 BCE, high-levels of WSHG-like ancestry can be detected in various populations of Central Asia until the Bronze Age. The population of the Botai culture, while probably not directly descended from WSHG, displays a high affinity with the WSHG lineage.[73] The European-Siberian cline defined by Eastern hunter-gatherer-like ancestry stretched from Central Europe to Siberia and was already established 10,000 years ago, including the West Siberian hunter-gatherers, all deriving their ancestry primarily from Paleolithic Siberians (ANE).[74]

"Princess of Xiaohe", one of the Tarim mummies, the "best representatives" of Ancient North Eurasians.[75]

Among the Ancient Northeast Asians (ANA) of the Neolithic to Early Bronze Age period, Baikal Eneolithic (Baikal_EN) and Baikal Early Bronze Age (Baikal_EBA) derived 6.4% to 20.1% ancestry from ANE, while the rest of their ancestry was derived from ANA. Fofonovo_EN near by Lake Baikal were mixture of 12-17% ANE ancestry and 83-87% ANA ancestry.[76]

Tarim mummies

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A 2021 genetic study on the Tarim mummies found that they were primarily descended from a population represented by the Afontova Gora 3 specimen (AG3), genetically displaying "high affinity" with it.[75] The genetic profile of the Afontova Gora 3 individual represented about 72% of the ancestry of the Tarim mummies, while the remaining 28% of their ancestry was derived from a population represented by the Baikal EBA (Early Bronze Age Northeast Asian Baikal populations).[75]

The Tarim mummies are thus one of the rare Holocene populations who derive most of their ancestry from the Ancient North Eurasians (ANE, specifically the Mal'ta and Afontova Gora populations), despite their distance in time (around 14,000 years).[75] Having survived in a type of "genetic bottleneck" in the Tarim basin where they preserved and perpetuated their ANE ancestry, the Tarim mummies, more than any other ancient populations, can be considered as "the best representatives" of the Ancient North Eurasians among all sampled known Bronze Age populations.[75]

North Han Chinese populations

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Several studies reveal minor West Eurasian-derived admixture among Shaanxi Han Chinese, especially those living in Guanzhong and Shaanbei (2–5%).[77][78] Ancient North Eurasian admixture is more dominant among Shaanxi Han Chinese compared to other Han subgroups.[79]

West Asian populations

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A model has been presented by Vallini et al. 2024, suggesting that Ancient Iranians (Iranian hunter-gatherers) formed from a deep Ancient West Eurasian lineage (WEC2, around 72%), and from minor geneflow from Basal Eurasian (around 18%) and Ancient East Eurasian (around 10%) sources. The Ancient West Eurasian component associated with Iranian hunter-gatherers is inferred to have diverged from the West Eurasian Core lineage (represented by Kostenki-14; WEC), with the WEC2 component staying in the region of the Iranian Plateau, while the proper WEC component expanded into Europe.[80]

European populations

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Further information: Eastern Hunter-Gatherer, Scandinavian Hunter-Gatherer, and Western Steppe Herders

Lazaridis et al. (2014) detected ANE ancestry among modern European populations in proportions up to 20%.[81] In ancient European populations, the ANE genetic component is visible in tests of the Yamnaya people[56] but not of Western or Central Europeans predating the Corded Ware culture:[82] ANE ancestry was introduced in the European gene pool with the Eastern Hunter-Gatherer (EHG) lineage which derived significant ancestry from the ANE, c. 70%, with the remaining ancestry from a group more closely related to, but distinct from, Western Hunter-Gatherers (WHGs).[54][83][20] It is represented by multiple individuals, such as from Yuzhny Oleny in Karelia, one of Y-haplogroup R1a-M417, dated c. 8.4 kya, the other of Y-haplogroup J, dated c. 7.2 kya; and one individual from Samara, of Y-haplogroup R1b-P297, dated c. 7.6 kya, as well as individuals from Sidelkino and Popovo. After the end of the Last Glacial Maximum, the Western Hunter-Gatherers (WHG) and EHG lineages merged in Eastern Europe, accounting for early presence of ANE-derived ancestry in Mesolithic Europe. Evidence suggests that as Ancient North Eurasians migrated westward from Eastern Siberia, they absorbed Western Hunter-Gatherers and other West Eurasian populations as well.[84][20]

Villalba-Mouco et al. 2023 confirmed the strong affinity between the Eastern European Hunter-Gatherers (EHG) to the Ancient North Eurasians, and also found a low affinity to the Tianyuan man, explained by them having received significant amounts of ANE ancestry.[85]

Scandinavian Hunter-Gatherer (SHG) is represented by several individuals buried at Motala, Sweden ca. 6000 BC. They were descended from Western Hunter-Gatherers who initially settled Scandinavia from the south, and received later admixture from EHG who entered Scandinavia from the north through the coast of Norway.[86][56][87][61][88]

Western Steppe Herders (WSH) is the name given to a distinct ancestral component that represents descent closely related to the Yamnaya culture of the Pontic–Caspian steppe.[c] This ancestry is often referred to as Yamnaya ancestry or Steppe ancestry, and was formed from EHG and CHG (Caucasus hunter-gatherer) in about equal proportions.[90]

Phenotype prediction

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Genomic studies by Raghavan et al. (2014) and Fu et al. (2016) suggested that Mal'ta boy may have had brown eyes, and relatively dark hair and dark skin,[91][92] while cautioning that this analysis was based on an extremely low coverage of DNA that might not give an accurate prediction of pigmentation.[93] Mathieson, et al. (2018) could not determine if Mal'ta 1 boy had the derived allele associated with blond hair in ANE descendants, as they could obtain no coverage for this SNP.[94]

Anthropologic research

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One of the Tarim mummies, a "Beauty of Loulan" dated c. 2000 BCE

Han Kangxin (1994) Described the mummies found in the cemetery of Gumugou as possessing "clear western racial characteristics" approximating those of a so-called "proto-European type". Whereas he asserted that among the seven mummies in the suburbs of Loulan cemetery six possessed "clear European characteristics" approximating the Saka population of the southern Pamirs within the 6th century B.C.E; while one of them possessed "Mongoloid" traits.[95] Kozintsev (2020, 2022) argues that the historical Southern Siberian Okunevo population, and other Paleo-Siberians, which derive high amounts of their ancestry from Ancient North Eurasians, as possessing a distinct craniometric phenotype, which he dubbed "Americanoid", which represents the variation of the first humans in Siberia and should not be associated solely with ancient Caucasoids. The Ancient North Eurasians themselves originated among Ancient West Eurasians, and represent a "Boreal" variation of early humans. Craniometric data on ANE-rich remains (such as from Botai), show them to cluster most closely with remains from the Pit-Comb Ware culture in Eastern Europe, and to take up a more "western" position.[57][72]

Reconstruction of a female individual from Xiaohe Cemetery. Xinjiang Museum. The Tarim mummies are considered as the "best representatives" of Ancient North Eurasians.[96]

Zhang et al. (2021) proposed that the 'Western' like features of the earlier Tarim mummies could be attributed to their Ancient North Eurasian ancestry.[97] Previous craniometric analyses on the early Tarim mummies found that they formed their own cluster, and clustered with neither European-related Steppe pastoralists of the Andronovo and Afanasievo cultures, nor with inhabitants of the Western Asian BMAC culture, nor with East Asian populations further east, but displayed an affinity for two specimens from the Harappan site of the Indus Valley Civilisation.[98]

Evolution of blond hair

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Blond hair is associated with a single nucleotide polymorphism, the mutated allele rs12821256 of the KITLG gene.[99][100][101][102][103] The earliest known individual with this allele is a female south-central Siberian ANE individual from the Afontova Gora 3 site, which is dated to c. 17,000 before present (the earlier ANE Mal'ta boy lacks the sequence coverage to make this determination).[104] The allele then appears later in ANE-derived Eastern Hunter-Gatherer (EHG) populations at Samara, Motala and Ukraine, circa 10,000 BP, and then in populations with Steppe ancestry.[94] Mathieson, et al. (2018) thus argued that this allele originated in the Ancient North Eurasian population, before spreading to western Eurasia.[94]

Geneticist David Reich said that the KITLG gene for blond hair probably entered continental Europe in a population migration wave from the Eurasian steppe, by a population carrying substantial Ancient North Eurasian ancestry.[105] Hanel and Carlberg (2020) likewise report that populations derived Ancient North Eurasian ancestry, specifically the Eastern Hunter-Gatherers and the Yamnayas, were responsible for transmitting this gene to Europeans.[99] The gene was also found among the Tarim mummies.[106]

The mutation for blond hair is thought to have originated among the Afontova Gora population of the Ancient North Eurasian (ANE) cline of south-central Siberia

Comparative mythology

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Mal'ta–Buret' culture centrally perforated ivory plaques with abstract circles, and three snakes[1] According to archeologist Don Hitchcock "the snake is rare in northern hemisphere Paleolithic art, presumably because the cold conditions precluded a wide distribution of snakes. In addition, it can be seen that the snakes have very broad heads, as though they belong to the Cobra group - yet Cobras are now known only in southern asian localities." It has yet to be clarrified how the creators of these ivory plaques know snakes, or if there are other possible interpretations for these.[107]

Since the term 'Ancient North Eurasian' refers to a genetic bridge of connected mating networks, scholars of comparative mythology have argued that they probably shared myths and beliefs that could be reconstructed via the comparison of stories attested within cultures that were not in contact for millennia and stretched from the Pontic–Caspian steppe to the American continent.[22]

The mytheme of the dog guarding the Otherworld possibly stems from an older Ancient North Eurasian belief, as suggested by similar motifs found in Indo-European, Native American and Siberian mythology. In Siouan, Algonquian, Iroquoian, and in Central and South American beliefs, a fierce guard dog was located in the Milky Way, perceived as the path of souls in the afterlife, and getting past it was a test. The Siberian Chukchi and Tungus believed in a guardian-of-the-afterlife dog and a spirit dog that would absorb the dead man's soul and act as a guide in the afterlife. In Indo-European myths, the figure of the dog is embodied by Cerberus, Sarvarā, and Garmr. In Zoroastrianism, two four-eyed dogs guard the bridge to the afterlife called Chinvat Bridge. Anthony and Brown note that it might be one of the oldest mythemes recoverable through comparative mythology.[22]

A second canid-related series of beliefs, myths and rituals connected dogs with healing rather than death. For instance, Ancient Near Eastern and Turkic-Kipchaq myths are prone to associate dogs with healing and generally categorised dogs as impure. A similar myth-pattern is assumed for the Eneolithic site of Botai in Kazakhstan, dated to 3500 BC, which might represent the dog as absorber of illness and guardian of the household against disease and evil. In Mesopotamia, the goddess Nintinugga, associated with healing, was accompanied or symbolized by dogs. Similar absorbent-puppy healing and sacrifice rituals were practiced in Greece and Italy, among the Hittites, again possibly influenced by Near Eastern traditions.[22]

See also

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References

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  1. ^ a b Bednarik RG (2013). "Pleistocene Palaeoart of Asia". Arts. 2 (2): 46–76. doi:10.3390/arts2020046.
  2. ^ Lbova L (2021). "The Siberian Palaeolithic Site of Mal'ta: A Unique Source for The Study of Childhood Archaeology". Evolutionary Human Sciences. 2: e9. doi:10.1017/ehs.2021.5. PMC 10427291. PMID 37588521. S2CID 231980510.
  3. ^ "Paleolithic to Bronze Age Siberians Reveal Connections with First Americans and across Eurasia" (PDF). CELL. 2020.
  4. ^ Yang MA (6 January 2022). "A genetic history of migration, diversification, and admixture in Asia". Human Population Genetics and Genomics. 2 (1): 1–32. doi:10.47248/hpgg2202010001. ISSN 2770-5005.
  5. ^ Willerslev E, Meltzer DJ (June 2021). "Willerslev, E., Meltzer, D.J. Peopling of the Americas as inferred from ancient genomics. Nature 594, 356–364 (2021)". Nature. 594 (7863): 356–364. Bibcode:2021Natur.594..356W. doi:10.1038/s41586-021-03499-y. PMID 34135521. S2CID 235460793.
  6. ^ a b c d Flegontov et al. 2016.
  7. ^ Jeong et al. 2019
  8. ^ Grebenyuk PS, Fedorchenko AY, Dyakonov VM, Lebedintsev AI, Malyarchuk BA (2022). "Ancient Cultures and Migrations in Northeastern Siberia". Humans in the Siberian Landscapes. Springer Geography. Springer International Publishing. p. 93. doi:10.1007/978-3-030-90061-8_4. ISBN 978-3-030-90060-1.
  9. ^ Villalba-Mouco V, van de Loosdrecht MS, Rohrlach AB, Fewlass H, Talamo S, Yu H, et al. (1 March 2023). "A 23,000-year-old southern Iberian individual links human groups that lived in Western Europe before and after the Last Glacial Maximum". Nature Ecology & Evolution. 7 (4): 597–609. Bibcode:2023NatEE...7..597V. doi:10.1038/s41559-023-01987-0. ISSN 2397-334X. PMC 10089921. PMID 36859553.
  10. ^ Sikora M, Pitulko VV, Sousa VC, Allentoft ME, Vinner L, Rasmussen S, et al. (June 2019). "The population history of northeastern Siberia since the Pleistocene". Nature. 570 (7760): 182–188. Bibcode:2019Natur.570..182S. doi:10.1038/s41586-019-1279-z. hdl:1887/3198847. ISSN 1476-4687. PMID 31168093. S2CID 174809069.
  11. ^ Vallini et al. 2022, Supplementary Information, p. 17.
  12. ^ a b Lipson M, Reich D (10 January 2017). "A working model of the deep relationships of diverse modern human genetic lineages outside of Africa". Molecular Biology and Evolution. 34 (4): 889–902. doi:10.1093/molbev/msw293. ISSN 0737-4038. PMC 5400393. PMID 28074030.
  13. ^ Posth C, Nakatsuka N, Lazaridis I, Skoglund P, Mallick S, Lamnidis TC, et al. (November 2018). "Reconstructing the Deep Population History of Central and South America". Cell. 175 (5): 1185–1197.e22. doi:10.1016/j.cell.2018.10.027. ISSN 0092-8674. PMC 6327247. PMID 30415837.
  14. ^ Allentoft ME, Sikora M, Refoyo-Martínez A, Irving-Pease EK, Fischer A, Barrie W, et al. (January 2024). "Population genomics of post-glacial western Eurasia". Nature. 625 (7994): 301–311. Bibcode:2024Natur.625..301A. doi:10.1038/s41586-023-06865-0. ISSN 1476-4687. PMC 10781627. PMID 38200295.
  15. ^ a b c Gabidullina LR, Dzhaubermezov MA, Ekomasova NV, Sufyanova ZR, Khusnutdinova EK (2023). "Genetic History of Eurasia Before the Common Era". Opera Medica et Physiologica. 10 (3): 95–117. ISSN 2500-2295.
  16. ^ a b Massilani D, Skov L, Hajdinjak M, Gunchinsuren B, Tseveendorj D, Yi S, et al. (30 October 2020). "Denisovan ancestry and population history of early East Asians". Science. 370 (6516): 579–583. doi:10.1126/science.abc1166. ISSN 0036-8075. PMID 33122380. Fig. 2 Simplified demographic model including the Salkhit individual and other Eurasians older than 30,000 years: 25-33% geneflow from Salkhit to Yana, but Salkhit already had 22-26% gene flow from Ancient West Eurasians
  17. ^ a b c d Yang MA (6 January 2022). "A genetic history of migration, diversification, and admixture in Asia". Human Population Genetics and Genomics. 2 (1): 1–32. doi:10.47248/hpgg2202010001. ISSN 2770-5005. Two individuals (Yana 1 and 2) were sampled in northeastern Siberia at the Yana Rhinoceros Horn site off the Yana River dating to 31,000 years ago [69], and another three (Mal'ta 1 and Afontova Gora 2/3), dating to 24,000–17,000 years ago, were sampled in south central Siberia [51] (Figure 1B). Comparison to ancient and present-day populations in Europe and Asia using f4-statistics (non-normalized version of the D-statistic, Box 1) showed that these individuals were genetically more similar to European hunter-gatherers than to Tianyuan or present-day East and Southeast Asians [51,69], which suggests that Upper Paleolithic Siberians were split at an early point from a population that contributed to ancient European hunter-gatherers [79].
  18. ^ a b c d ZHANG M, PING W (15 June 2023). 古基因组揭示史前欧亚大陆现代人复杂遗传历史 [Ancient genomes reveal the complex genetic history of Prehistoric Eurasian modern humans]. 人类学学报 人类学学报 (in Chinese). 42 (3): 412–421. doi:10.16359/j.1000-3193/AAS.2023.0010. ISSN 1000-3193. 以发现于西伯利亚东北部约31.6 kaBP的Yana个体为代表的古西伯利亚北部人群(Ancient North Siberians, ANS)。该人群是具有约71%的欧洲祖源成分和29%的亚洲祖源成分的独立人群[2]。
  19. ^ a b Raff J (8 February 2022). Origin: A Genetic History of the Americas. Grand Central Publishing. p. 188. ISBN 978-1-5387-4970-8.
  20. ^ a b c d Posth C, Yu H, Ghalichi A, Rougier H, Crevecoeur I, Huang Y, et al. (March 2023). "Palaeogenomics of Upper Palaeolithic to Neolithic European hunter-gatherers". Nature. 615 (7950): 117–126. Bibcode:2023Natur.615..117P. doi:10.1038/s41586-023-05726-0. ISSN 1476-4687. PMC 9977688. PMID 36859578.
  21. ^ Reich 2018, p. 81
  22. ^ a b c d Anthony & Brown 2019, pp. 104–106.
  23. ^ Willerslev E, Meltzer DJ (June 2021). "Peopling of the Americas as inferred from ancient genomics". Nature. 594 (7863): 356–364. Bibcode:2021Natur.594..356W. doi:10.1038/s41586-021-03499-y. ISSN 1476-4687. PMID 34135521. S2CID 235460793.
  24. ^ Sikora M, Pitulko VV, Sousa VC, Allentoft ME, Vinner L, Rasmussen S, et al. (June 2019). "The population history of northeastern Siberia since the Pleistocene". Nature. 570 (7760): 182–188. Bibcode:2019Natur.570..182S. doi:10.1038/s41586-019-1279-z. hdl:1887/3198847. ISSN 1476-4687. PMID 31168093. S2CID 174809069.
  25. ^ "Hall 11 panorama". State Hermitage Museum.
  26. ^ Lbova L (2021). "The Siberian Paleolithic site of Mal'ta: a unique source for the study of childhood archaeology". Evolutionary Human Sciences. 3: 8, Fig. 6-1. doi:10.1017/ehs.2021.5. PMC 10427291. PMID 37588521. S2CID 231980510.
  27. ^ Kozintsev AG (31 December 2020). "The Origin of the Okunev Population, Southern Siberia: The Evidence of Physical Anthropology and Genetics". Archaeology, Ethnology & Anthropology of Eurasia. 48 (4): 135–145. doi:10.17746/1563-0110.2020.48.4.135-145. ISSN 1563-0110.
  28. ^ a b c d e Vallini et al. 2022
  29. ^ Fu Q, Posth C, Hajdinjak M, Petr M, Mallick S, Fernandes D, et al. (June 2016). "The genetic history of Ice Age Europe". Nature. 534 (7606): 200–205. Bibcode:2016Natur.534..200F. doi:10.1038/nature17993. hdl:10211.3/198594. ISSN 1476-4687. PMC 4943878. PMID 27135931.
  30. ^ Xu D, Li H (5 May 2017). Languages and Genes in Northwestern China and Adjacent Regions. Springer. ISBN 978-981-10-4169-3. In the study of Zhong et al. haplogroups O-M175, C-M130, D-M174 and N-M231 still suggests the substantial contribution of the southern route. However, the Central Asia and West Eurasia related haplogroups, such as haplogroups R-M207 and Q-M242, occur primarily in northwestern East Asia and their frequencies gradually decrease from west to east. In addition, the Y-STR diversities of haplogroups R-M207 and Q-M242 also indicate the existence of northern route migration about 18,000 years ago from Central Asia to North Asia.
  31. ^ Estes R (12 September 2020). "Y DNA Haplogroup P Gets a Brand-New Root – Plus Some Branches". DNAeXplained - Genetic Genealogy. Retrieved 27 June 2024.
  32. ^ "P YTree". www.yfull.com. Retrieved 27 July 2024. formed 44300 ybp, TMRCA 41500 ybp
  33. ^ "Allen Ancient DNA Resource (AADR): Downloadable genotypes of present-day and ancient DNA data | David Reich Lab". reich.hms.harvard.edu. Retrieved 21 January 2024.
  34. ^ Zhang F, Ning C, Scott A, Fu Q, Bjørn R, Li W, et al. (27 October 2021). "The genomic origins of the Bronze Age Tarim Basin mummies". Nature. 599 (7884): 256–261. Bibcode:2021Natur.599..256Z. doi:10.1038/s41586-021-04052-7. ISSN 1476-4687. PMC 8580821. PMID 34707286.
  35. ^ Zhang F, Ning C, Scott A, Fu Q, Bjørn R, Li W, et al. (27 October 2021). "The genomic origins of the Bronze Age Tarim Basin mummies". Nature. 599 (7884): 256–261. Bibcode:2021Natur.599..256Z. doi:10.1038/s41586-021-04052-7. ISSN 1476-4687. PMC 8580821. PMID 34707286.
  36. ^ a b Maier R, Flegontov P, Flegontova O, Işıldak U, Changmai P, Reich D (14 April 2023). Nordborg M, Przeworski M, Balding D, Wiuf C (eds.). "On the limits of fitting complex models of population history to f-statistics". eLife. 12: e85492. doi:10.7554/eLife.85492. ISSN 2050-084X. PMC 10310323. PMID 37057893.
  37. ^ a b c Grebenyuk PS, Fedorchenko AY, Dyakonov VM, Lebedintsev AI, Malyarchuk BA (2022), Bocharnikov VN, Steblyanskaya AN (eds.), "Ancient Cultures and Migrations in Northeastern Siberia", Humans in the Siberian Landscapes: Ethnocultural Dynamics and Interaction with Nature and Space, Springer Geography, Cham: Springer International Publishing, pp. 89–133, doi:10.1007/978-3-030-90061-8_4, ISBN 978-3-030-90061-8
  38. ^ Zhang X, Ji X, Li C, Yang T, Huang J, Zhao Y, et al. (July 2022). "A Late Pleistocene human genome from Southwest China". Current Biology. 32 (14): 3095–3109.e5. Bibcode:2022CBio...32E3095Z. doi:10.1016/j.cub.2022.06.016. PMID 35839766. S2CID 250502011.
  39. ^ Posth C, Nakatsuka N, Lazaridis I, Skoglund P, Mallick S, Lamnidis TC, et al. (15 November 2018). "Reconstructing the Deep Population History of Central and South America". Cell. 175 (5): 1185–1197.e22. doi:10.1016/j.cell.2018.10.027. ISSN 0092-8674. PMC 6327247. PMID 30415837.
  40. ^ Svensson E, Günther T, Hoischen A, Hervella M, Munters AR, Ioana M, et al. (26 July 2021). "Genome of Peştera Muierii skull shows high diversity and low mutational load in pre-glacial Europe". Current Biology. 31 (14): 2973–2983.e9. Bibcode:2021CBio...31E2973S. doi:10.1016/j.cub.2021.04.045. hdl:10810/52864. ISSN 0960-9822. PMID 34010592. S2CID 234793812.
  41. ^ Villalba-Mouco V, van de Loosdrecht MS, Rohrlach AB, Fewlass H, Talamo S, Yu H, et al. (April 2023). "A 23,000-year-old southern Iberian individual links human groups that lived in Western Europe before and after the Last Glacial Maximum". Nature Ecology & Evolution. 7 (4): 597–609. Bibcode:2023NatEE...7..597V. doi:10.1038/s41559-023-01987-0. hdl:10230/57143. ISSN 2397-334X. PMC 10089921. PMID 36859553.
  42. ^ Balter 2013.
  43. ^ Sikora M, Pitulko VV, Sousa VC, Allentoft ME, Vinner L, Rasmussen S, et al. (June 2019). "The population history of northeastern Siberia since the Pleistocene". Nature. 570 (7760): 182–188. Bibcode:2019Natur.570..182S. doi:10.1038/s41586-019-1279-z. hdl:1887/3198847. ISSN 1476-4687. PMID 31168093. S2CID 174809069.
  44. ^ Yang MA, Fan X, Sun B, Chen C, Lang J, Ko YC, et al. (17 July 2020). "Ancient DNA indicates human population shifts and admixture in northern and southern China". Science. 369 (6501): 282–288. Bibcode:2020Sci...369..282Y. doi:10.1126/science.aba0909. ISSN 0036-8075. PMID 32409524. S2CID 218649510.
  45. ^ Yang MA, Gao X, Theunert C, Tong H, Aximu-Petri A, Nickel B, et al. (23 October 2017). "40,000-Year-Old Individual from Asia Provides Insight into Early Population Structure in Eurasia". Current Biology. 27 (20): 3202–3208.e9. Bibcode:2017CBio...27E3202Y. doi:10.1016/j.cub.2017.09.030. ISSN 0960-9822. PMC 6592271. PMID 29033327.
  46. ^ Mao X, Zhang H, Qiao S, Liu Y, Chang F, Xie P, et al. (10 June 2021). "The deep population history of northern East Asia from the Late Pleistocene to the Holocene". Cell. 184 (12): 3256–3266.e13. doi:10.1016/j.cell.2021.04.040. ISSN 0092-8674. PMID 34048699.
  47. ^ a b Sikora M, Pitulko VV, Sousa VC, Allentoft ME, Vinner L, Rasmussen S, et al. (June 2019). "The population history of northeastern Siberia since the Pleistocene". Nature. 570 (7760): 182–188. Bibcode:2019Natur.570..182S. doi:10.1038/s41586-019-1279-z. hdl:1887/3198847. ISSN 1476-4687. PMID 31168093. S2CID 174809069.
  48. ^ Maier et al. 2023.
  49. ^ Maier R, Flegontov P, Flegontova O, Işıldak U, Changmai P, Reich D (14 April 2023). Nordborg M, Przeworski M, Balding D, Wiuf C (eds.). "On the limits of fitting complex models of population history to f-statistics". eLife. 12: e85492. doi:10.7554/eLife.85492. ISSN 2050-084X. PMC 10310323. PMID 37057893.
  50. ^ a b Vallini L, Zampieri C, Shoaee MJ, Bortolini E, Marciani G, Aneli S, et al. (25 March 2024). "The Persian plateau served as hub for Homo sapiens after the main out of Africa dispersal". Nature Communications. 15 (1): 1882. Bibcode:2024NatCo..15.1882V. doi:10.1038/s41467-024-46161-7. ISSN 2041-1723. PMC 10963722. PMID 38528002. Similarly, Mal'ta and Yana fall in an intermediate position between the two axes, the result of a palaeolithic admixture between EEC and WEC groups18.
  51. ^ Allentoft ME, Sikora M, Refoyo-Martínez A, Irving-Pease EK, Fischer A, Barrie W, et al. (10 January 2024). "Population genomics of post-glacial western Eurasia". Nature. 625 (7994): 301–311. Bibcode:2024Natur.625..301A. doi:10.1038/s41586-023-06865-0. ISSN 1476-4687. PMC 10781627. PMID 38200295. Supplementary Information: Fig. S3d.16. Admixture graph of deep Eurasian lineages
  52. ^ "Figurine of a woman with the "Spanish rose" hairdo". Art of Mal'ta. Novosibirsk State University.
  53. ^ Cassidy, Jim, Ponkratova, Irina, Fitzhugh, Ben, eds. (2022). Maritime Prehistory of Northeast Asia. The Archaeology of Asia-Pacific Navigation. Vol. 6. doi:10.1007/978-981-19-1118-7. ISBN 978-981-19-1117-0. S2CID 249355636.
  54. ^ a b c Lazaridis et al. 2016.
  55. ^ a b Essel E (3 May 2023). "Ancient human DNA recovered from a Palaeolithic pendant". Nature. 618 (7964): 328–332. Bibcode:2023Natur.618..328E. doi:10.1038/s41586-023-06035-2. PMC 10247382. PMID 37138083.
  56. ^ a b c d e Haak et al. 2015.
  57. ^ a b Kozintsev AG (31 December 2020). "The Origin of the Okunev Population, Southern Siberia: The Evidence of Physical Anthropology and Genetics". Archaeology, Ethnology & Anthropology of Eurasia. 48 (4): 135–145. doi:10.17746/1563-0110.2020.48.4.135-145. ISSN 1563-0110.
  58. ^ Moreno-Mayar et al. 2018.
  59. ^ a b Wong et al. 2017.
  60. ^ a b Raghavan et al. 2013.
  61. ^ a b Günther et al. 2018.
  62. ^ a b Yu et al. 2020.
  63. ^ Raff 2022, p. 188
  64. ^ Raff 2022, pp. 188–189
  65. ^ Grebenyuk PS, Fedorchenko AY, Dyakonov VM, Lebedintsev AI, Malyarchuk BA (2022). "Ancient Cultures and Migrations in Northeastern Siberia". Humans in the Siberian Landscapes. Springer Geography. Springer International Publishing. p. 96. doi:10.1007/978-3-030-90061-8_4. ISBN 978-3-030-90060-1.
  66. ^ Hoffecker JF (4 March 2016). "Beringia and the global dispersal of modern humans: Beringia and the Global Dispersal of Modern Humans". Evolutionary Anthropology: Issues, News, and Reviews. 25 (2): 64–78. doi:10.1002/evan.21478. PMID 27061035. S2CID 3519553.
  67. ^ Moreno-Mayar et al. 2018.
  68. ^ Gakuhari T, Nakagome S, Rasmussen S, Allentoft ME (25 August 2020). "Ancient Jomon genome sequence analysis sheds light on migration patterns of early East Asian populations". Communications Biology. 3 (1): Fig.1 A, C. doi:10.1038/s42003-020-01162-2. ISSN 2399-3642. PMC 7447786. PMID 32843717.
  69. ^ Osada & Kawai 2021.
  70. ^ Cooke NP, Mattiangeli V, Cassidy LM, Okazaki K, Stokes CA, Onbe S, et al. (2021). "Ancient genomics reveals tripartite origins of Japanese populations". Science Advances. 7 (38): eabh2419. Bibcode:2021SciA....7.2419C. doi:10.1126/sciadv.abh2419. PMC 8448447. PMID 34533991.
  71. ^ Wang K, Yu H, Radzevičiūtė R, Kiryushin YF, Tishkin AA, Frolov YV, et al. (12 January 2023). "Middle Holocene Siberian genomes reveal highly connected gene pools throughout North Asia". Current Biology. 33 (3): 423–433. Bibcode:2023CBio...33E.423W. doi:10.1016/j.cub.2022.11.062. ISSN 0960-9822. PMID 36638796. S2CID 255750546.
  72. ^ a b Kozintsev AG (4 January 2022). "Patterns in the Population History of Northern Eurasia from the Mesolithic to the Early Bronze Age, Based on Craniometry and Genetics". Archaeology, Ethnology & Anthropology of Eurasia. 49 (4): 140–151. doi:10.17746/1563-0110.2021.49.4.140-151. ISSN 1563-0110. Judging by the huge distribution area of ANE, it was a legacy of early Homo sapiens, members of the Boreal meta-population (Biasutti, 1941: 275; Kozintsev, 2013, 2014) [...] The ancestor of ANE was the ANS (Ancient North Siberian) autosomal component, represented in a male from the Upper Paleolithic Yana site, dating to 31.6 ka BP (Sikora et al., 2019). ANS is thought to have originated among West Eurasians soon after their divergence from East Eurasians about 43 ka BP. [...] The sample that includes the few Botai crania (No. 24) takes a more "western" position (Fig. 1), forming a pair with the Pit-Comb Ware group of European Russia, and this is supported by archaeological data suggesting that these cultures are related (Mosin, 2003: 97–98)
  73. ^ Narasimhan 2019.
  74. ^ Mattila TM, Svensson EM, Juras A, Günther T, Kashuba N, Ala-Hulkko T, et al. (9 August 2023). "Genetic continuity, isolation, and gene flow in Stone Age Central and Eastern Europe". Communications Biology. 6 (1): 793. doi:10.1038/s42003-023-05131-3. hdl:10852/104260. ISSN 2399-3642. PMC 10412644. PMID 37558731.
  75. ^ a b c d e Zhang F (November 2021). "The genomic origins of the Bronze Age Tarim Basin mummies". Nature. 599 (7884): 256–261. Bibcode:2021Natur.599..256Z. doi:10.1038/s41586-021-04052-7. ISSN 1476-4687. PMC 8580821. PMID 34707286.
  76. ^ Jeong et al. 2020.
  77. ^ He GL, Wang MG, Li YX, Zou X (2022). "Fine-scale north-to-south genetic admixture profile in Shaanxi Han Chinese revealed by genome-wide demographic history reconstruction". Journal of Systematics and Evolution. 60 (4): 955–972. doi:10.1111/jse.12715 – via Wiley Online Library.{{cite journal}}: CS1 maint: overridden setting (link)
  78. ^ Chiang CW, Mangul S, Robles C, Sankararaman S (2018). "A Comprehensive Map of Genetic Variation in the World's Largest Ethnic Group —Han Chinese". Molecular Biology and Evolution. 35 (11): 2736–2750. doi:10.1093/molbev/msy170. PMC 6693441. PMID 30169787.
  79. ^ He GL, Wang MG, Li YX, Zou X (2022). "Fine-scale north-to-south genetic admixture profile in Shaanxi Han Chinese revealed by genome-wide demographic history reconstruction". Journal of Systematics and Evolution. 60 (4): 955–972. doi:10.1111/jse.12715 – via Wiley Online Library.{{cite journal}}: CS1 maint: overridden setting (link)
  80. ^ Vallini L, Zampieri C, Shoaee MJ, Bortolini E, Marciani G, Aneli S, et al. (25 March 2024). "The Persian plateau served as hub for Homo sapiens after the main out of Africa dispersal". Nature Communications. 15 (1): 1882. Bibcode:2024NatCo..15.1882V. doi:10.1038/s41467-024-46161-7. ISSN 2041-1723. PMC 10963722. PMID 38528002.
  81. ^ Lazaridis I, Patterson N, Mittnik A (September 2014). "Ancient human genomes suggest three ancestral populations for present-day Europeans". Nature. 513 (7518): 409–413. arXiv:1312.6639. Bibcode:2014Natur.513..409L. doi:10.1038/nature13673. ISSN 1476-4687. PMC 4170574. PMID 25230663.
  82. ^ Lazaridis, Patterson & Mittnik 2014.
  83. ^ Wang 2019.
  84. ^ Mathieson et al. 2018.
  85. ^ Villalba-Mouco V, van de Loosdrecht MS, Rohrlach AB, Fewlass H, Talamo S, Yu H, et al. (April 2023). "A 23,000-year-old southern Iberian individual links human groups that lived in Western Europe before and after the Last Glacial Maximum". Nature Ecology & Evolution. 7 (4): 597–609. Bibcode:2023NatEE...7..597V. doi:10.1038/s41559-023-01987-0. hdl:10230/57143. ISSN 2397-334X. PMC 10089921. PMID 36859553.
  86. ^ Lazaridis et al. 2014.
  87. ^ Mathieson 2015.
  88. ^ Mittnik 2018.
  89. ^ Jeong et al. 2018.
  90. ^ Jeong et al. 2019.
  91. ^ Raghavan & Skoglund et al. 2014.
  92. ^ Fu Q, Posth C, Hajdinjak M (2 May 2016). "The genetic history of Ice Age Europe". Nature. 504 (7606): 200–5. Bibcode:2016Natur.534..200F. doi:10.1038/nature17993. hdl:10211.3/198594. PMC 4943878. PMID 27135931.
  93. ^ Raghavan M, Skoglund P, Graf KE, Metspalu M, Albrechtsen A, Moltke I, et al. (2 January 2014). "Upper Palaeolithic Siberian genome reveals dual ancestry of NativeAmericans". Nature. 505 (7481): 87–91. Bibcode:2014Natur.505...87R. doi:10.1038/nature12736. ISSN 0028-0836. PMC 4105016. PMID 24256729.
  94. ^ a b c Mathieson et al. 2018 "Supplementary Information page 52
  95. ^ Han K (November 1994). "The Study of ancient human skeletons from Xinjiang, China" (PDF). Sino-Platonic Papers. 51: 16.
  96. ^ Zhang 2021: "The Tarim mummies are among only a few known Holocene populations that derive the majority of their ancestry from Pleistocene ANE groups, who once made up the huntergatherer populations of southern Siberia, and which are represented by individual genomes from the archaeological sites of Mal'ta (MA-1)29 and Afontova Gora (AG3). (...) The Tarim mummies are currently the best representative of the pre-pastoralist ANE-related population that once inhabited Central Asia and southern Siberia (Extended Data Fig. 2A), even though Tarim_EMBA1 postdates these populations in time."
  97. ^ Zhang F, Ning C, Scott A, Fu Q, Bjørn R, Li W, et al. (November 2021). "The genomic origins of the Bronze Age Tarim Basin mummies". Nature. 599 (7884): 256–261. Bibcode:2021Natur.599..256Z. doi:10.1038/s41586-021-04052-7. ISSN 1476-4687. PMC 8580821. PMID 34707286.
  98. ^ "A Craniometric Investigation of The Bronze Age Settlement of Xinjiang - Important | PDF". Scribd. Retrieved 4 April 2023.
  99. ^ a b Carlberg & Hanel 2020.
  100. ^ Hartz P. "KIT LIGAND; KITLG". Online Mendelian Inheritance in Man.
  101. ^ Khan R. "KITLG makes you white skinned?". Discover Magazine.
  102. ^ "P21583". Uniprot.
  103. ^ Guenther et al. 2014.
  104. ^ Evans G (2019). Skin Deep: Dispelling the Science of Race (1 ed.). Simon and Schuster. p. 139. ISBN 978-1-78607-623-6.
  105. ^ Reich D (2018). Who We are and How We Got Here: Ancient DNA and the New Science of the Human Past. Oxford University Press. ISBN 978-0-19-882125-0.
  106. ^ Zhang 2021.
  107. ^ "The Mal'ta - Buret' venuses and culture in Siberia". donsmaps.com.

Notes

[edit]
  1. ^ Lipson & Reich (2017) model the ANE as 18% East Asian and the remainder West Eurasian. Posth et al. 2018 models the Mal'ta boy as 16% East Asian and remainder West Eurasian. Sikora et al. (2019) model the Yana individuals as 22–29% East Eurasian and the remainder West Eurasian ("Using admixture graphs and outgroup-based estimation of mixture proportions, we find that ANS can be modelled as early West Eurasian with an approximately 22% contribution from early East Asians"). Massilani et al. (2020): "In agreement with previous results (10), the Yana individuals are estimated to have about one-third of their ancestry from early East Eurasians and the remaining two-thirds from the early West Eurasians."Vallini et al. (2022) model Yana as 50% West Eurasian and 50% East Eurasian. Maier et al. (2023) modeled the ANE/ANS as around 25% East Eurasian and 75% West Eurasian.
  2. ^ Lipson & Reich (2017) model the ANA as 18% East Eurasian and the remainder West Eurasian. Posth et al. 2018 models the Mal'ta boy as 16% East Eurasian and remainder West Eurasian. Sikora et al. (2019) model the Yana individuals as 22% East Eurasian and the remainder West Eurasian ("Using admixture graphs and outgroup-based estimation of mixture proportions, we find that ANS can be modelled as early West Eurasian with an approximately 22% contribution from early East Asians"). Massilani et al. (2020): "In agreement with previous results (10), the Yana individuals are estimated to have about one-third of their ancestry from early East Eurasians and the remaining two-thirds from the early West Eurasians." Vallini et al. (2022) model Yana as 50% West Eurasian and 50% East Eurasian. Maier et al. (2023) modeled the ANE/ANS as around 25% East Eurasian and 75% West Eurasian.
  3. ^ "Recent paleogenomic studies have shown that migrations of Western steppe herders (WSH) beginning in the Eneolithic (ca. 3300–2700 BCE) profoundly transformed the genes and cultures of Europe and central Asia... The migration of these Western steppe herders (WSH), with the Yamnaya horizon (ca. 3300–2700 BCE) as their earliest representative, contributed not only to the European Corded Ware culture (ca. 2500–2200 BCE) but also to steppe cultures located between the Caspian Sea and the Altai-Sayan mountain region, such as the Afanasievo (ca. 3300–2500 BCE) and later Sintashta (2100–1800 BCE) and Andronovo (1800–1300 BCE) cultures."[89]

Bibliography

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