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Climax species
[edit]Climax species, also called late seral, late-successional, K-selected or equilibrium species, are plant species that can germinate and grow with limited resources, like low-sun exposure or low water availability.[1] They are the species within forest succession that are more adapted to stable and predictable environments, and will remain essentially unchanged in terms of species composition for as long as a site remains undisturbed.[2]
The seedlings of climax species can grow in the shade of the parent trees, ensuring their dominance indefinitely. The presence of climax species can also reduce the prevalence of other species within an ecosystem.[3] However, a disturbance, such as fire, may kill the climax species, allowing pioneer or earlier successional species to re-establish for a time.[4] They are the opposite of pioneer species, also known as ruderal, fugitive, opportunistic or R-selected species, in the sense that climax species are good competitors but poor colonizers, whereas pioneer species are good colonizers but poor competitors.[5]
In addition to disturbances promoting species diversity by allowing more pioneer species to develop, the age of a successional ecosystem increases species diversity.[6] As ecosystems progress toward later successional stages, the appearance of climax species allows other species to develop that cannot tolerate the high sun-exposure that pioneer species can.
Given the prevailing ecological conditions, climax species dominate the climax community. When the pace of succession slows down as a result of ecological homeostasis the maximum permitted biodiversity is reached.[7] Their reproductive strategies and other adaptive characteristics can be considered more sophisticated than those of opportunistic species.[1]
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Examples
[edit]Picea glauca (White spruce) is an example of a climax species in the Northern forests of North America due to its ability to adapt to resource scarce, stable conditions, it dominates Northern forest ecosystem in the absence of a disturbance.[8]
Other examples of climax species in old-growth forests:
- Canadian hemlock
- Pacific silver fir
- White fir
- Yellow carabeen
- Blue grama
- Douglas fir
- Coast redwood
- European beech
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Notes
[edit]- ^ a b Shimano, Koji (2000-02-01). "A power function for forest structure and regeneration pattern of pioneer and climax species in patch mosaic forests". Plant Ecology. 146 (2): 205–218. doi:10.1023/A:1009867302660. ISSN 1573-5052.
- ^ Wehenkel, Christian; Bergmann, Fritz; Gregorius, Hans-Rolf (2006-07-01). "Is there a trade-off between species diversity and genetic diversity in forest tree communities?". Plant Ecology. 185 (1): 151–161. doi:10.1007/s11258-005-9091-2. ISSN 1573-5052.
- ^ Do, Ha T. T.; Grant, John C.; Zimmer, Heidi C.; Trinh, Bon N.; Nichols, J. Doland (2020-05-29). "Site conditions for regeneration of climax species, the key for restoring moist deciduous tropical forest in Southern Vietnam". PLOS ONE. 15 (5): e0233524. doi:10.1371/journal.pone.0233524. ISSN 1932-6203. PMC 7259571. PMID 32469962.
{{cite journal}}: CS1 maint: PMC format (link) CS1 maint: unflagged free DOI (link) - ^ Wehenkel, Christian; Bergmann, Fritz; Gregorius, Hans-Rolf (2006-07-01). "Is there a trade-off between species diversity and genetic diversity in forest tree communities?". Plant Ecology. 185 (1): 151–161. doi:10.1007/s11258-005-9091-2. ISSN 1573-5052.
- ^ Brown, S.; Dockery, J.; Pernarowski, M. (2005-03-01). "Traveling wave solutions of a reaction diffusion model for competing pioneer and climax species". Mathematical Biosciences. 194 (1): 21–36. doi:10.1016/j.mbs.2004.10.001. ISSN 0025-5564.
- ^ "Succession: A Closer Look | Learn Science at Scitable". www.nature.com. Retrieved 2020-12-03.
- ^ Ernest, S. K. M. (2008-01-01), Jørgensen, Sven Erik; Fath, Brian D. (eds.), "Homeostasis", Encyclopedia of Ecology, Oxford: Academic Press, pp. 1879–1884, doi:10.1016/b978-008045405-4.00507-3, ISBN 978-0-08-045405-4, retrieved 2020-12-03
- ^ "Picea glauca". www.fs.fed.us. Retrieved 2020-12-03.