User:Kweetal nl/IOC132/Pycnonotidae

Family Pycnonotidae - Bulbuls

Sequence of Pycnonotidae is primarily based on Shakya & Sheldon (2017) with modifications from Dejtaradol et al. (2016); Fuchs et al. (2018).

Pycnonotidae   genera: 32
Pycnonotidae   species: 166
Pycnonotidae   subspecies: 264

Pycnonotidae   Breeding Range: IO, AF, PAL, OR, AU

Pycnonotidae

Common name	Species + authority	Br. range	au	yr	id
Sombre greenbul	Andropadus importunus (Vieillot, 1818)	AF	Viei	1818	1
Slender-billed greenbul	Stelgidillas gracilirostris (Strickland, 1844)	AF	Stri	1844	2
Golden greenbul	Calyptocichla serinus (Verreaux, J & Verreaux, É, 1855)	AF	Verr	1855	3
Black-collared bulbul	Neolestes torquatus Cabanis, 1875	AF	Caba	1875	4
Red-tailed bristlebill	Bleda syndactylus (Swainson, 1837)	AF	Swai	1837	5
Green-tailed bristlebill	Bleda eximius (Hartlaub, 1855)	AF	Hart	1855	6
Grey-headed bristlebill	Bleda canicapillus (Hartlaub, 1854)	AF	Hart	1854	7
Yellow-lored bristlebill	Bleda notatus (Cassin, 1856)	AF	Cass	1856	8
Yellow-eyed bristlebill	Bleda ugandae Van Someren, 1915	AF	Some	1915	9
Yellow-throated leaflove	Atimastillas flavicollis (Swainson, 1837)	AF	Swai	1837	10
Spotted greenbul	Ixonotus guttatus Verreaux, J & Verreaux, É, 1851	AF	Verr	1851	11
Swamp palm bulbul	Thescelocichla leucopleura (Cassin, 1855)	AF	Cass	1855	12
Joyful greenbul	Chlorocichla laetissima (Sharpe, 1899)	AF	Shar	1899	13
Prigogine's greenbul	Chlorocichla prigoginei De Roo, 1967	AF	Roo	1967	14
Falkenstein's greenbul	Chlorocichla falkensteini (Reichenow, 1874)	AF	Reic	1874	15
Yellow-bellied greenbul	Chlorocichla flaviventris (Smith, A, 1834)	AF	Smit	1834	16
Simple greenbul	Chlorocichla simplex (Hartlaub, 1855)	AF	Hart	1855	17
Honeyguide greenbul	Baeopogon indicator (Verreaux, J & Verreaux, É, 1855)	AF	Verr	1855	18
Sjöstedt's greenbul	Baeopogon clamans (Sjöstedt, 1893)	AF	Sjös	1893	19
Cameroon greenbul	Arizelocichla montana (Reichenow, 1892)	AF	Reic	1892	20
Western greenbul	Arizelocichla tephrolaema (Gray, GR, 1862)	AF	Gray	1862	21
Kakamega greenbul	Arizelocichla kakamegae (Sharpe, 1900)	AF	Shar	1900	22
Shelley's greenbul	Arizelocichla masukuensis (Shelley, 1897)	AF	Shel	1897	23
Uluguru greenbul	Arizelocichla neumanni Hartert, EJO, 1922	AF	Hart	1922	24
Black-browed greenbul	Arizelocichla fusciceps (Shelley, 1893)	AF	Shel	1893	25
Yellow-throated greenbul	Arizelocichla chlorigula (Reichenow, 1899)	AF	Reic	1899	26
Olive-breasted greenbul	Arizelocichla kikuyuensis (Sharpe, 1891)	AF	Shar	1891	27
Mountain greenbul	Arizelocichla nigriceps (Shelley, 1889)	AF	Shel	1889	28
Olive-headed greenbul	Arizelocichla olivaceiceps (Shelley, 1896)	AF	Shel	1896	29
Stripe-faced greenbul	Arizelocichla striifacies (Reichenow & Neumann, 1895)	AF	Reic	1895	30
Stripe-cheeked greenbul	Arizelocichla milanjensis (Shelley, 1894)	AF	Shel	1894	31
Western bearded greenbul	Criniger barbatus (Temminck, 1821)	AF	Temm	1821	32
Eastern bearded greenbul	Criniger chloronotus (Cassin, 1859)	AF	Cass	1859	33
Red-tailed greenbul	Criniger calurus (Cassin, 1856)	AF	Cass	1856	34
White-bearded greenbul	Criniger ndussumensis Reichenow, 1904	AF	Reic	1904	35
Yellow-bearded greenbul	Criniger olivaceus (Swainson, 1837)	AF	Swai	1837	36
Little greenbul	Eurillas virens (Cassin, 1857)	AF	Cass	1857	37
Yellow-whiskered greenbul	Eurillas latirostris (Strickland, 1844)	AF	Stri	1844	38
Plain greenbul	Eurillas curvirostris (Cassin, 1859)	AF	Cass	1859	39
Little grey greenbul	Eurillas gracilis (Cabanis, 1880)	AF	Caba	1880	40
Ansorge's greenbul	Eurillas ansorgei (Hartert, EJO, 1907)	AF	Hart	1907	41
Lowland tiny greenbul	Phyllastrephus debilis (Sclater, WL, 1899)	AF	Scla	1899	42
Montane tiny greenbul	Phyllastrephus albigula (Grote, 1919)	AF	Grot	1919	43
White-throated greenbul	Phyllastrephus albigularis (Sharpe, 1882)	AF	Shar	1882	44
Xavier's greenbul	Phyllastrephus xavieri (Oustalet, 1892)	AF	Oust	1892	45
Icterine greenbul	Phyllastrephus icterinus (Bonaparte, 1850)	AF	Bona	1850	46
Terrestrial brownbul	Phyllastrephus terrestris Swainson, 1837	AF	Swai	1837	47
Cameroon olive greenbul	Phyllastrephus poensis Alexander, 1903	AF	Alex	1903	48
Northern brownbul	Phyllastrephus strepitans (Reichenow, 1879)	AF	Reic	1879	49
Grey-olive greenbul	Phyllastrephus cerviniventris Shelley, 1894	AF	Shel	1894	50
Fischer's greenbul	Phyllastrephus fischeri (Reichenow, 1879)	AF	Reic	1879	51
Cabanis's greenbul	Phyllastrephus cabanisi (Sharpe, 1882)	AF	Shar	1882	52
Placid greenbul	Phyllastrephus placidus (Shelley, 1899)	AF	Shel	1899	53
Red-tailed leaflove	Phyllastrephus scandens Swainson, 1837	AF	Swai	1837	54
Sassi's olive greenbul	Phyllastrephus lorenzi Sassi, 1914	AF	Sass	1914	55
Yellow-streaked greenbul	Phyllastrephus flavostriatus (Sharpe, 1876)	AF	Shar	1876	56
Sharpe's greenbul	Phyllastrephus alfredi (Shelley, 1903)	AF	Shel	1903	57
Grey-headed greenbul	Phyllastrephus poliocephalus (Reichenow, 1892)	AF	Reic	1892	58
Toro olive greenbul	Phyllastrephus hypochloris (Jackson, FJ, 1906)	AF	Jack	1906	59
Baumann's olive greenbul	Phyllastrephus baumanni Reichenow, 1895	AF	Reic	1895	60
Pale-olive greenbul	Phyllastrephus fulviventris Cabanis, 1876	AF	Caba	1876	61
Hairy-backed bulbul	Tricholestes criniger (Blyth, 1845)	OR	Blyt	1845	62
Hook-billed bulbul	Setornis criniger Lesson, RP, 1839	OR	Less	1839	63
Yellow-bellied bulbul	Alophoixus phaeocephalus (Hartlaub, 1844)	OR	Hart	1844	64
Palawan bulbul	Alophoixus frater (Sharpe, 1877)	OR	Shar	1877	65
Grey-cheeked bulbul	Alophoixus tephrogenys (Jardine & Selby, 1833)	OR	Jard	1833	66
Penan bulbul	Alophoixus ruficrissus (Sharpe, 1879)	OR	Shar	1879	67
Brown-cheeked bulbul	Alophoixus bres (Lesson, RP, 1831)	OR	Less	1831	68
White-throated bulbul	Alophoixus flaveolus (Gould, 1836)	OR	Goul	1836	69
Ochraceous bulbul	Alophoixus ochraceus (Moore, F, 1854)	OR	Moor	1854	70
Puff-throated bulbul	Alophoixus pallidus (Swinhoe, 1870)	OR	Swin	1870	71
Striated bulbul	Alcurus striatus (Blyth, 1842)	OR	Blyt	1842	72
Finsch's bulbul	Iole finschii (Salvadori, 1871)	OR	Salv	1871	73
Sulphur-bellied bulbul	Iole palawanensis (Tweeddale, 1878)	OR	Twee	1878	74
Olive bulbul	Iole viridescens Blyth, 1867	OR	Blyt	1867	75
Buff-vented bulbul	Iole crypta Oberholser, 1918	OR	Ober	1918	76
Charlotte's bulbul	Iole charlottae (Finsch, 1867)	OR	Fins	1867	77
Cachar bulbul	Iole cacharensis (Deignan, 1948)	OR	Deig	1948	78
Grey-eyed bulbul	Iole propinqua (Oustalet, 1903)	OR	Oust	1903	79
Cream-striped bulbul	Hemixos leucogrammicus (Müller, S, 1836)	OR	Müll	1836	80
Ashy bulbul	Hemixos flavala Blyth, 1845	OR	Blyt	1845	81
Cinereous bulbul	Hemixos cinereus (Blyth, 1845)	OR	Blyt	1845	82
Chestnut bulbul	Hemixos castanonotus Swinhoe, 1870	OR	Swin	1870	83
Yellow-browed bulbul	Acritillas indica (Jerdon, 1839)	OR	Jerd	1839	84
Sumatran bulbul	Ixos sumatranus (Wardlaw-Ramsay, RG, 1882)	OR	Ward	1882	85
Javan bulbul	Ixos virescens Temminck, 1825	OR	Temm	1825	86
Streaked bulbul	Ixos malaccensis (Blyth, 1845)	OR	Blyt	1845	87
Mountain bulbul	Ixos mcclellandii (Horsfield, 1840)	OR	Hors	1840	88
Nicobar bulbul	Ixos nicobariensis (Moore, F, 1854)	OR	Moor	1854	89
Philippine bulbul	Hypsipetes philippinus (Forster, JR, 1795)	OR	Fors	1795	90
Mindoro bulbul	Hypsipetes mindorensis (Steere, 1890)	OR	Stee	1890	91
Streak-breasted bulbul	Hypsipetes siquijorensis (Steere, 1890)	OR	Stee	1890	92
Seram golden bulbul	Hypsipetes affinis (Hombron & Jacquinot, 1841)	AU	Homb	1841	93
Sangihe golden bulbul	Hypsipetes platenae (Blasius, W, 1888)	AU	Blas	1888	94
Togian golden bulbul	Hypsipetes aureus (Walden, 1872)	AU	Wald	1872	95
Banggai golden bulbul	Hypsipetes harterti (Stresemann, 1912)	AU	Stre	1912	96
Sula golden bulbul	Hypsipetes longirostris (Wallace, 1863)	AU	Wall	1863	97
Halmahera golden bulbul	Hypsipetes chloris (Finsch, 1867)	AU	Fins	1867	98
Obi golden bulbul	Hypsipetes lucasi (Hartert, EJO, 1903)	AU	Hart	1903	99
Buru golden bulbul	Hypsipetes mysticalis (Wallace, 1863)	AU	Wall	1863	100
Visayan bulbul	Hypsipetes guimarasensis (Steere, 1890)	OR	Stee	1890	101
Yellowish bulbul	Hypsipetes everetti (Tweeddale, 1877)	OR	Twee	1877	102
Camiguin bulbul	Hypsipetes catarmanensis Rand & Rabor, 1969	OR	Rand	1969	103
Zamboanga bulbul	Hypsipetes rufigularis Sharpe, 1877	OR	Shar	1877	104
Brown-eared bulbul	Hypsipetes amaurotis (Temminck, 1830)	PAL	Temm	1830	105
Reunion bulbul	Hypsipetes borbonicus (Pennant, 1781)	IO	Penn	1781	106
Malagasy bulbul	Hypsipetes madagascariensis (Müller, PLS, 1776)	AF	Müll	1776	107
Mauritius bulbul	Hypsipetes olivaceus Jardine & Selby, 1837	IO	Jard	1837	108
White-headed bulbul	Hypsipetes thompsoni (Bingham, 1900)	OR	Bing	1900	109
Black bulbul	Hypsipetes leucocephalus (Gmelin, JF, 1789)	OR	Gmel	1789	110
Square-tailed bulbul	Hypsipetes ganeesa Sykes, 1832	OR	Syke	1832	111
Grande Comore bulbul	Hypsipetes parvirostris Milne-Edwards & Oustalet, 1885	AF	Miln	1885	112
Moheli bulbul	Hypsipetes moheliensis (Benson, 1960)	AF	Bens	1960	113
Seychelles bulbul	Hypsipetes crassirostris Newton, E, 1867	IO	Newt	1867	114
Puff-backed bulbul	Euptilotus eutilotus (Jardine & Selby, 1837)	OR	Jard	1837	115
Black-and-white bulbul	Microtarsus melanoleucos Eyton, 1839	OR	Eyto	1839	116
Yellow-wattled bulbul	Poliolophus urostictus (Salvadori, 1870)	OR	Salv	1870	117
Grey-headed bulbul	Brachypodius priocephalus (Jerdon, 1839)	OR	Jerd	1839	118
Black-headed bulbul	Brachypodius melanocephalos (Gmelin, JF, 1788)	OR	Gmel	1788	119
Andaman bulbul	Brachypodius fuscoflavescens Hume, 1873	OR	Hume	1873	120
Blue-wattled bulbul	Brachypodius nieuwenhuisii (Finsch, 1901)	OR	Fins	1901	121
Spectacled bulbul	Ixodia erythropthalmos (Hume, 1878)	OR	Hume	1878	122
Grey-bellied bulbul	Ixodia cyaniventris (Blyth, 1842)	OR	Blyt	1842	123
Scaly-breasted bulbul	Ixodia squamata (Temminck, 1828)	OR	Temm	1828	124
Black-crested bulbul	Rubigula flaviventris (Tickell, 1833)	OR	Tick	1833	125
Flame-throated bulbul	Rubigula gularis (Gould, 1836)	OR	Goul	1836	126
Black-capped bulbul	Rubigula melanictera (Gmelin, JF, 1789)	OR	Gmel	1789	127
Ruby-throated bulbul	Rubigula dispar (Horsfield, 1821)	OR	Hors	1821	128
Bornean bulbul	Rubigula montis Sharpe, 1879	OR	Shar	1879	129
Bare-faced bulbul	Nok hualon (Woxvold, Duckworth & Timmins, 2009)	OR	Woxv	2009	130
Crested finchbill	Spizixos canifrons Blyth, 1845	OR	Blyt	1845	131
Collared finchbill	Spizixos semitorques Swinhoe, 1861	OR	Swin	1861	132
Cream-vented bulbul	Pycnonotus simplex Lesson, RP, 1839	OR	Less	1839	133
Olive-winged bulbul	Pycnonotus plumosus Blyth, 1845	OR	Blyt	1845	134
Asian red-eyed bulbul	Pycnonotus brunneus Blyth, 1845	OR	Blyt	1845	135
Straw-headed bulbul	Pycnonotus zeylanicus (Gmelin, JF, 1789)	OR	Gmel	1789	136
Spot-necked bulbul	Pycnonotus tympanistrigus (Müller, S, 1836)	OR	Müll	1836	137
Cream-eyed bulbul	Pycnonotus pseudosimplex Shakya, Lim, Moyle, Rahman, Lakim & Sheldon, 2019	OR	Shak	2019	138
Ashy-fronted bulbul	Pycnonotus cinereifrons (Tweeddale, 1878)	OR	Twee	1878	139
White-browed bulbul	Pycnonotus luteolus (Lesson, RP, 1841)	OR	Less	1841	140
Ayeyarwady bulbul	Pycnonotus blanfordi Jerdon, 1862	OR	Jerd	1862	141
Streak-eared bulbul	Pycnonotus conradi (Finsch, 1873)	OR	Fins	1873	142
Pale-eyed bulbul	Pycnonotus davisoni (Hume, 1875)	OR	Hume	1875	143
Stripe-throated bulbul	Pycnonotus finlaysoni Strickland, 1844	OR	Stri	1844	144
Flavescent bulbul	Pycnonotus flavescens Blyth, 1845	OR	Blyt	1845	145
Aceh bulbul	Pycnonotus snouckaerti Siebers, 1928	OR	Sieb	1928	146
Orange-spotted bulbul	Pycnonotus bimaculatus (Horsfield, 1821)	OR	Hors	1821	147
Pale-faced bulbul	Pycnonotus leucops (Sharpe, 1888)	OR	Shar	1888	148
Yellow-throated bulbul	Pycnonotus xantholaemus (Jerdon, 1845)	OR	Jerd	1845	149
Yellow-eared bulbul	Pycnonotus penicillatus Blyth, 1851	OR	Blyt	1851	150
Brown-breasted bulbul	Pycnonotus xanthorrhous Anderson, 1869	OR	Ande	1869	151
Light-vented bulbul	Pycnonotus sinensis (Gmelin, JF, 1789)	OR	Gmel	1789	152
Styan's bulbul	Pycnonotus taivanus Styan, 1893	OR	Stya	1893	153
Red-whiskered bulbul	Pycnonotus jocosus (Linnaeus, 1758)	OR	Linn	1758	154
Yellow-vented bulbul	Pycnonotus goiavier (Scopoli, 1786)	OR	Scop	1786	155
Red-vented bulbul	Pycnonotus cafer (Linnaeus, 1766)	OR	Linn	1766	156
Sooty-headed bulbul	Pycnonotus aurigaster (Vieillot, 1818)	OR	Viei	1818	157
White-eared bulbul	Pycnonotus leucotis (Gould, 1836)	PAL	Goul	1836	158
Himalayan bulbul	Pycnonotus leucogenys (Gray, JE, 1835)	PAL	Gray	1835	159
White-spectacled bulbul	Pycnonotus xanthopygos (Hemprich & Ehrenberg, 1833)	PAL	Hemp	1833	160
African red-eyed bulbul	Pycnonotus nigricans (Vieillot, 1818)	AF	Viei	1818	161
Common bulbul	Pycnonotus barbatus (Desfontaines, 1789)	AF	Desf	1789	162
Dodson's bulbul	Pycnonotus dodsoni Sharpe, 1895	AF	Shar	1895	163
Somali bulbul	Pycnonotus somaliensis Reichenow, 1905	AF	Reic	1905	164
Dark-capped bulbul	Pycnonotus tricolor (Hartlaub, 1862)	AF	Hart	1862	165
Cape bulbul	Pycnonotus capensis (Linnaeus, 1766)	AF	Linn	1766	166

Pycnonotidae genera

Genus + authority	spp	Br. range	au	yr	id	Comment
Andropadus Swainson, 1832	1	AF	Swai	1832	1
Stelgidillas Oberholser, 1899	1	AF	Ober	1899	2	Stelgidillas bulbuls separated from Andropadus following Johansson et al. (2007).
Calyptocichla Oberholser, 1905	1	AF	Ober	1905	3
Neolestes Cabanis, 1875	1	AF	Caba	1875	4	Neolestes is a member of the Afrotropical clade of bulbuls (Johansson et al. 2008; Zuccon & Ericson 2010).
Bleda Bonaparte, 1857	5	AF	Bona	1857	5	Sequence of Bleda follows Huntley & Voelker (2016).
Atimastillas Oberholser, 1905	1	AF	Ober	1905	6
Ixonotus Verreaux, J & Verreaux, É, 1851	1	AF	Verr	1851	7
Thescelocichla Oberholser, 1905	1	AF	Ober	1905	8
Chlorocichla Sharpe, 1882	5	AF	Shar	1882	9
Baeopogon Heine, 1860	2	AF	Hein	1860	10
Arizelocichla Oberholser, 1905	12	AF	Ober	1905	11	Arizelocichla bulbuls separated from Andropadus following Johansson et al. (2007).
Criniger Temminck, 1820	5	AF	Temm	1820	12
Eurillas Oberholser, 1899	5	AF	Ober	1899	13	Eurillas bulbuls separated from Andropadus following Johansson et al. (2007).
Phyllastrephus Swainson, 1832	20	AF	Swai	1832	14
Tricholestes Salvadori, 1874	1	OR	Salv	1874	15
Setornis Lesson, RP, 1839	1	OR	Less	1839	16
Alophoixus Oates, 1889	8	OR	Oate	1889	17	Separate the Oriental species of Criniger bulbuls to a different genus Alophoixus (Fishpool & Tobias 2005; BLI). Sequence in Alophoixus is based on Shakya et al. (2020).
Alcurus Blyth, 1843	1	OR	Blyt	1843	18	Genus resurrected for Pycnonotus striatus, which is sister to a clade of species deeply genetically divergent from members of core Pycnonotus (Shakya & Sheldon 2017).
Iole Blyth, 1844	7	OR	Blyt	1844	19
Hemixos Blyth, 1845	4	OR	Blyt	1845	20
Acritillas Oberholser, 1905	1	OR	Ober	1905	21
Ixos Temminck, 1825	5	OR	Temm	1825	22
Hypsipetes Vigors, 1831	25	OR AU AF IO PAL	Vigo	1831	23
Euptilotus Reichenbach, 1850	1	OR	Reic	1850	24	Pycnonotus s.l. Clade A of Shakya & Sheldon (2017) is deeply divergent from Pycnonotus s.s. and Clade B (formerly Pycnonotus). Relatively long branch lengths and disparate morphology support treatment as four genera, including the monotypic Euptilotus.
Microtarsus Eyton, 1839	1	OR	Eyto	1839	25	Pycnonotus s.l. Clade A of Shakya & Sheldon (2017) is deeply divergent from Pycnonotus s.s. and Clade B (formerly Pycnonotus). Relatively long branch lengths and disparate morphology support treatment as four genera, including Microtarsus.
Poliolophus Sharpe, 1877	1	OR	Shar	1877	26	Pycnonotus s.l. Clade A of Shakya & Sheldon (2017) is deeply divergent from Pycnonotus s.s. and Clade B (formerly Pycnonotus). Relatively long branch lengths and disparate morphology support treatment as four genera, including the monospecific Poliolophus.
Brachypodius Blyth, 1845	4	OR	Blyt	1845	27	Pycnonotus s.l. Clade A of Shakya & Sheldon (2017) is deeply divergent from Pycnonotus s.s. and Clade B (formerly Pycnonotus). Relatively long branch lengths and disparate morphology support treatment as four genera, including Brachypodius.
Ixodia Blyth, 1845	3	OR	Blyt	1845	28	Ixodia is resurrected here for a subclade within Clade B of Shakya & Sheldon (2017; where misspelled Ixodea) which is fairly deeply diverged genetically from other members of Clade B (now Rubigula) and deeply diverged from species in Pycnonotus s.s. The genus name Ixodia Blyth, 1845 is available and is not preoccupied by Ixodia Robert Brown, 1812 (Asteraceae) (R. Schodde, pers. comm.), hence there was no need for the replacement name Ixidia Blyth, 1846, which has nevertheless been used a few times after 1899.
Rubigula Blyth, 1845	5	OR	Blyt	1845	29	The resurrected genus Rubigula is adopted here for a subclade within Clade B of Shakya & Sheldon (2017) which is fairly deeply divergent genetically from other members of Clade B (now Ixodia) and deeply divergent from species in Pycnonotus s.s. (Shakya & Sheldon 2017).
Nok Fuchs, Pasquet, Stuart, B, Woxvold, Duckworth & Bowie, 2018	1	OR	Fuch	2018	30	Sister to Spizixos; moved from Pycnonotus to the monotypic genus Nok Fuchs et al., 2018.
Spizixos Blyth, 1845	2	OR	Blyt	1845	31
Pycnonotus Boie, F, 1826	34	OR AF PAL	Boie	1826	32