User:Cougroyalty/crocodiles

Collection of useful crocodilian cladograms and references

Cladograms

Crocodylidae cladogram (from 2021 Azzara et al.)

Crocodylidae	Osteolaeminae Rimasuchus lloydi† Voay robustus† Osteolaemus osborni Osborn’s dwarf crocodile Osteolaemus tetraspis Dwarf crocodile "Crocodylus" gariepensis† Brochuchus parvidens† Brochuchus pigotti† Euthecodon arambourgii† Euthecodon brumpti† Crocodylinae Mecistops cataphractus West African slender-snouted crocodile Crocodylus thorbjarnarsoni† Crocodylus anthropophagus† Crocodylus niloticus Nile crocodile Crocodylus checchiai† Crocodylus moreletii Morelet's crocodile Crocodylus intermedius Orinoco crocodile Crocodylus acutus American crocodile Crocodylus rhombifer Cuban crocodile Crocodylus palaeindicus† Crocodylus palustris Mugger crocodile Crocodylus ossifragus† Crocodylus siamensis Siamese crocodile Crocodylus mindorensis Philippine crocodile Crocodylus johnstoni Freshwater crocodile Crocodylus porosus Saltwater crocodile Crocodylus raninus Borneo crocodile Crocodylus novaeguineae New Guinea crocodile	Paleoafrican Crocodylus Neotropical Crocodylus Indo-Pacific Crocodylus

Crocodylinae cladogram (from 2021 Azzara et al.)

Crocodylinae	Mecistops cataphractus West African slender-snouted crocodile Crocodylus thorbjarnarsoni† Crocodylus anthropophagus† Crocodylus niloticus Nile crocodile Crocodylus checchiai† Crocodylus moreletii Morelet's crocodile Crocodylus intermedius Orinoco crocodile Crocodylus acutus American crocodile Crocodylus rhombifer Cuban crocodile Crocodylus palaeindicus† Crocodylus palustris Mugger crocodile Crocodylus ossifragus† Crocodylus siamensis Siamese crocodile Crocodylus mindorensis Philippine crocodile Crocodylus johnstoni Freshwater crocodile Crocodylus porosus Saltwater crocodile Crocodylus raninus Borneo crocodile Crocodylus novaeguineae New Guinea crocodile	Paleoafrican Crocodylus Neotropical Crocodylus Indo-Pacific Crocodylus

Crocodylidae cladogram (from 2021 Hekkala et al.)

Crocodylidae

Osteolaeminae Mecistops cataphractus West African slender-snouted crocodile Euthecodon† Brochuchus† Rimasuchus† Osteolaemus osborni Osborn’s dwarf crocodile Osteolaemus tetraspis Dwarf crocodile Crocodylinae Voay† Crocodylus Crocodylus anthropophagus† Crocodylus thorbjarnarsoni† Crocodylus palaeindicus† Crocodylus Tirari Desert† Asia+Australia Crocodylus johnstoni Freshwater crocodile Crocodylus novaeguineae New Guinea crocodile Crocodylus mindorensis Philippine crocodile Crocodylus porosus Saltwater crocodile Crocodylus siamensis Siamese crocodile Crocodylus palustris Mugger crocodile Africa+New World Crocodylus checchiai† Crocodylus falconensis† Crocodylus suchus West African crocodile Crocodylus niloticus Nile crocodile New World Crocodylus moreletii Morelet's crocodile Crocodylus rhombifer Cuban crocodile Crocodylus intermedius Orinoco crocodile Crocodylus acutus American crocodile

(crown group)

Crocodylinae cladogram (from 2021 Hekkala et al.)

Crocodylinae

Voay† Crocodylus Crocodylus anthropophagus† Crocodylus thorbjarnarsoni† Crocodylus palaeindicus† Crocodylus Tirari Desert† Asia+Australia Crocodylus johnstoni Freshwater crocodile Crocodylus novaeguineae New Guinea crocodile Crocodylus mindorensis Philippine crocodile Crocodylus porosus Saltwater crocodile Crocodylus siamensis Siamese crocodile Crocodylus palustris Mugger crocodile Africa+New World Crocodylus checchiai† Crocodylus falconensis† Crocodylus suchus West African crocodile Crocodylus niloticus Nile crocodile New World Crocodylus moreletii Morelet's crocodile Crocodylus rhombifer Cuban crocodile Crocodylus intermedius Orinoco crocodile Crocodylus acutus American crocodile

Alligatoroidea cladogram (from 2018 Bona et al.)

Alligatoroidea

Leidyosuchus† Diplocynodon† Alligatoridae Alligatorinae Ceratosuchus† Allognathosuchus† Navajosuchus† Arambourgia† Procaimanoidea† Wannaganosuchus† Alligator prenasalis† Alligator mcgrewi† Alligator olseni† Alligator sinensis Chinese alligator Culebrasuchus† Alligator mississippiensis American alligator Alligator mefferdi† Alligator thomsoni† (stem group) Caimaninae Stangerochampsa† Albertochampsa† Brachychampsa† Protocaiman† Gnatusuchus† Globidentosuchus† Eocaiman† Notocaiman† Kuttanacaiman† Purussaurus† Mourasuchus† Necrosuchus† Tsoabichi† Paleosuchus trigonatus Smooth-fronted caiman Paleosuchus palpebrosus Cuvier's dwarf caiman Centenariosuchus† Caiman latirostris Broad-snouted caiman Melanosuchus niger Black caiman Caiman yacare Yacare caiman Caiman crocodilus Spectacled caiman Caiman brevirostris† La Venta Caiman† Caiman wannlangstoni† (stem group) (crown group)

(stem group)

Gavialoidea cladogram (from 2022 Iijima et al.) with little continent pictures

Crocodyloidea Gavialoidea †Maroccosuchus zennaroi †Megadontosuchus arduini †Dollosuchoides densmorei †Xaymacachampsa kugleri †Kentisuchus astrei †Kentisuchus spenceri Gavialidae †Tomistoma cairense †Tomistoma coppensi †Maomingosuchus petrolica Tomistominae Tomistoma schlegelii, false gharial †Tomistoma lusitanica †Gavialosuchus eggenburgensis †Melitosaurus champsoides †Tomistoma calaritanum †Tomistoma gaudense †Thecachampsa carolinensis †Thecachampsa antiqua Gavialinae †Paratomistoma courti †Penghusuchus pani †Toyotamaphimeia machikanensis †Hanyusuchus sinensis †Eosuchus lerichei †Eosuchus minor †Ocepesuchus eoafricanus †Eothoracosaurus mississippiensis †Thoracosaurus macrorhynchus †Thoracosaurus neocesariensis †Aktiogavialis puertoricensis †Eogavialis africanum †Argochampsa krebsi †Piscogavialis jugaliperforatus †Ikanogavialis gameroi †Siquisiquesuchus venezuelensis †Dadagavialis gunai †Gryposuchus neogaeus †Gryposuchus croizati †Aktiogavialis caribesi †Gryposuchus pachakamue †Gryposuchus colombianus †Rhamphosuchus crassidens †Myanmar gavialid †Gavialis lewisi †Gavialis bengawanicus Gavialis gangeticus, gharial †Siwalik Gavialis

Recent Crocodilia studies

Studies supporting Longirostres

• 2022 Darlim et al. tip dating study of crown crocodilians simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data:

• 2021 Rio & Mannion really in-depth morphological study with revised data set to align with molecular data.

• 2018 Lee & Yates tip dating study simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data:

• 2003 Harshman et al. molecular study placing the False gharial in Gavialidae, and first naming Longirostres.

• 2007 Willis et al. molecular study placing the False gharial in Gavialidae.

• 2008 Gatesy & Amato molecular study placing the False gharial in Gavialidae.

• 2012 Erickson et al. bite force study referencing 2008 Gatesy study.

• 2021 Hekkala et al. study using DNA of extinct Voay crocodile species and referencing 2018 Lee & Yates study.

Fossil morphological studies showing Brevirostres

• 2009 Delfino & Smith study for Asiatosuchus depressifrons.

• 2011 Brochu study for Necrosuchus.

• 2011 Puértolas et al. study for Arenysuchus.

• 2012 Holliday & Gardner study of Aegisuchus.

• 2015 Narváez et al. study for Lohuecosuchus.

• 2016 Narváez et al. study for Agaresuchus.

• 2016 Wang et al. study of Eoalligator and Protoalligator.

• 2017 Wu et al. study of "Crocodylus" megarhinus.

• 2019 Massonne et al. study of Alligatoroidea and specifically Orientalosuchus.

• 2020 Cossette & Brochu study of Deinosuchus, a basal member of Alligatoroidea.

• 2021 Blanco study of Allodaposuchidae referencing 2016 Narvaez et al. study.

• 2021 Ristevski et al. study of Gunggamarandu.

Other studies

• 2013 Hastings et al. study of Panama caimans & alligators naming Culebrasuchus.

• 2018 Bona et al. study covering just alligators and caimans.

• 2021 Azzara et al. study of Crocodylus anthropophagus.

• 2021 Stocker et al. study of Alligatoridae, Caimaninae, and specifically of new species Chinatichampsus wilsonorum.