Trigonochlamydidae

Trigonochlamydidae; Temporal range: Recent PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Ghost slug, Selenochlamys ysbryda
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Infraorder:	Limacoidei
Family:	Trigonochlamydidae; Hesse, 1882
Subfamilies and genera
	Parmacellillinae P. Hesse, 1926 Parmacellilla Simroth, 1910; ; Trigonochlamydinae P. Hesse, 1882 Boreolestes Schileyko & Kijashko, 1999; Drilolestes Lindholm, 1925; Hyrcanolestes Simroth, 1901; Khostalestes Suvorov, 2003; Lesticulus Schileyko, 1988; Pseudomilax O. Boettger, 1881; Trigonochlamys O. Boettger, 1881; Troglolestes Liovushkin & Matiokin, 1965; ;
Diversity
	9 genera, 11 species

Trigonochlamydidae is a family of air-breathing land slugs, terrestrial gastropod molluscs in the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

Distribution

The native distribution of Trigonochlamydidae includes Caucasus,^[1] Iran and Turkey.^[3]

Taxonomy

Previously, Trigonochlamydidae was placed in the superfamily Trigonochlamydoidea Hesse, 1882, in the subinfraorder Sigmurethra. This was the only family within that superfamily.^[4]

The family Trigonochlamydidae is now classified within the limacoid clade, which itself belongs to the clade Stylommatophora, within the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).

The following two subfamilies have been recognized in the taxonomy of Bouchet & Rocroi (2005):

subfamily Trigonochlamydinae Hesse, 1882 – synonyms: Selenochlamydinae I. M. Likharev & Wiktor, 1980
subfamily Parmacellillinae Hesse, 1926^[5]

Genera

There are nine^[3] genera in the family Trigonochlamydidae with a total of 11 species:

Subfamily Trigonochlamydinae

Boreolestes Schileyko & Kijashko, 1999
- Boreolestes likharevi Schileyko & Kijashko, 1999 – type species^[6]
- Boreolestes sylvestris Kijashko in Schileyko & Kijashko, 1999^[6]
Drilolestes Lindholm, 1925 – with only one species Drilolestes retowskii (O. Boettger, 1884)^[6]
Hyrcanolestes Simroth, 1901 – with only one species Hyrcanolestes velitaris (Martens, 1880)^[6]
Khostalestes Suvorov, 2003^[3] – with only one species Khostalestes kochetkovi Suvorov, 2003^[3]^[6]
Lesticulus Schileyko, 1988 – with only one species Lesticulus nocturnus Schileyko, 1988^[6]
Trigonochlamys O. Boettger, 1881 – type genus of the family Trigonochlamydidae, with only one species Trigonochlamys imitatrix O. Boettger, 1881^[6]
Troglolestes Ljovushkin & Matiokin, 1965 – with only one species Troglolestes sokolovi Liovushkin & Matiokin, 1965^[6]

Subfamily Parmacellillinae

Parmacellilla Simroth, 1910 – type genus of the subfamily Parmacellillinae, with only one species Parmacellilla filipowitschi Simroth, 1910^[7]

Synonym:

Pseudomilax O. Boettger, 1881 is a synonym for various genera in Trigonochlamydidae.

Selenochlamys O. Boettger, 1883 has been removed from the Trigonochlamydidae to the Oxychilidae based on DNA evidence.^[8]

Cladogram

The following cladogram shows the phylogenic relationships of this family with the other families in the limacoid clade:^[9]

References

^ ^a ^b "Family summary for Trigonochlamydidae". AnimalBase, last modified 26 August 2005, accessed 1 September 2010.
^ MolluscaBase eds. (2024). MolluscaBase. Trigonochlamydidae Hesse, 1882. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=867826 on 21 August 2024
^ ^a ^b ^c ^d Suvorov A. N. (2003). "A new species and genus of carnivorous slugs (Pulmonata Trigonochlamydidae) from West Transcaucasia". Ruthenica 13: 149–152. abstract.
^ Sistemica (Spanish list of genera), cited 7 October 2008.
^ Hesse P. (1926). "Die Nacktschnecken der palearktischen Region". Abhandlungen des Archiv für Molluskenkunde 2(1): 1–152. page 47 and page 54. plates 1–2.
^ ^a ^b ^c ^d ^e ^f ^g ^h Kantor Yu I., Vinarski M. V., Schileyko A. A. & Sysoev A. V.(published online on 22 December 2009). "Catalogue of the continental mollusks of Russia and adjacent territories". Version 2.3.
^ Schileyko A. A. (2003). "Treatise on Recent Terrestrial Pulmonate Molluscs. Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." Ruthenica, Supplement 2(11): 1467–1626, Moscow. page 1467.
^ Neiber, Marco T.; Walther, Frank; Hausdorf, Bernhard (October 2020). "Phylogenetic relationships of ghost slugs (Selenochlamys) and overlooked instances of limacization in Western Palaearctic Limacoidei (Gastropoda: Stylommatophora)". Molecular Phylogenetics and Evolution. 151: 106897. Bibcode:2020MolPE.15106897N. doi:10.1016/j.ympev.2020.106897. PMID 32585288.
^ Hausdorf B. (March 2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379–390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.

[AB-1] "Family summary for Trigonochlamydidae". AnimalBase, last modified 26 August 2005, accessed 1 September 2010.

[2] MolluscaBase eds. (2024). MolluscaBase. Trigonochlamydidae Hesse, 1882. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetails&id=867826 on 21 August 2024

[Suvorov_2003-3] Suvorov A. N. (2003). "A new species and genus of carnivorous slugs (Pulmonata Trigonochlamydidae) from West Transcaucasia". Ruthenica 13: 149–152. abstract.

[4] Sistemica (Spanish list of genera), cited 7 October 2008.

[5] Hesse P. (1926). "Die Nacktschnecken der palearktischen Region". Abhandlungen des Archiv für Molluskenkunde 2(1): 1–152. page 47 and page 54. plates 1–2.

[Kantor_2009-6] ^ ^a ^b ^c ^d ^e ^f ^g ^h Kantor Yu I., Vinarski M. V., Schileyko A. A. & Sysoev A. V.(published online on 22 December 2009). "Catalogue of the continental mollusks of Russia and adjacent territories". Version 2.3.

[7] Schileyko A. A. (2003). "Treatise on Recent Terrestrial Pulmonate Molluscs. Part 11. Trigonochlamydidae, Papillodermidae, Vitrinidae, Limacidae, Bielziidae, Agriolimacidae, Boettgerillidae, Camaenidae." Ruthenica, Supplement 2(11): 1467–1626, Moscow. page 1467.

[Neiber2020-8] Neiber, Marco T.; Walther, Frank; Hausdorf, Bernhard (October 2020). "Phylogenetic relationships of ghost slugs (Selenochlamys) and overlooked instances of limacization in Western Palaearctic Limacoidei (Gastropoda: Stylommatophora)". Molecular Phylogenetics and Evolution. 151: 106897. Bibcode:2020MolPE.15106897N. doi:10.1016/j.ympev.2020.106897. PMID 32585288.

[9] Hausdorf B. (March 2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379–390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.

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