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Zerynthia Ochsenheimer , 1816

Ackery (1975) pointed out that Zerynthia is a junior synonym of Parnalius Rafinesque, 1815 published as a replacement name for the preoccupied Thais Fabricius and moreover correctly listed by Sherborne (1929) Neave (1940) and Cowan (1970). Parnalius has subsequently been suppressed cf. Bull. Zool. Nomencl. 36 (1979): 102)

The Generic generic synonymy is as follows:

Thais Fabricius, 1807 (Syst. Gloss: XI): typus mancat.; junior homonym of Thais Roding, 1789.

Thais Fabricius, 1807 (Mag. Insekt 6: 283) [Genotype|type]] Papilio hypsiphyle Fabricius by monotypy.

Parnalius Rafinesque, 1815 (Analyse Nat.: 128) type by Art. 67 (1): Papilio hypsiphyle Fabricius.

Zerynthia Ochsenheimer, 1816 (Schmett. Europ. 4: 128 type polyxena Schiff, selected by Scudder; n.n. pro Thais Fabricius. subspecies polyxena, medisicaste, rumina.

Eugraphis (Dalman i.l.) Billberg, 1820 (Ennum. Ins.: 75) type hypsiphyle Fabricius by monotypy.

Zerinthia Sodovsky, 1837 (Bull. Soc. Imp. Nat. Mosc. 1834 (10): 82: nom. emend. pro Zerynthia Ochs.

Cerynthia Staudinger, 1861 (Staudinger and Woche: Cat. Lep. Europ. (1): 1: nom. emend.

Zerynthyia Zerny, 1927 (Eos, Madrid 3: 313): nom. emend.

Thays Rocci, 1928 (Boll. Soc. Ent. Ital. 10: 55): nom. emend.

Allancastria Bryk, 1934 (Parnassiana 2: 104) type cerisyi Godart by monotypy: synonymised by Ackery, 1975.

Some considerable taxonomic changes have been introduced for this interesting group in recent years, e.g. Ackery (1975), Larsen (1973), Kuhna (1977), Kocak (1975, 1977), Freina (1979). To the present author those of Kuhna and Freina are most acceptable.

Species synonymy

polyxena Denis and Schiffermuller 1775 ( hypermnestra Scopoli, 1763)

hypsiphyle Fabricus, 1777

hypsipyle Schultzen, 1776

Bricuox (1975) has all South France populations under creusa Meigen described from Italy, with four sub-races that might or might not be comparable to local (ecological) races or forms:- creusa trans cassandra (Gard): creusa (Nice): microcreusa (Roquebrune) thus following the example set by Verity. The need for a strict revision is much in evidence.

Subspecies

(a) cassandra Hubner and Geyer, 1823-3 (1828) South France, North Italy.

forms

ochracea Staudinger, 1861

vitrina Rothschild, 1918

inornata Pionneau

bella Neuburger, 1903

reducta Zelezny, 1915

quincunx Eisner

microcreusa Verity, 1947

deminuta Verity, 1947

(b) padana Rocci, 1929 (North Italy: Piedmonte, Lombardy, Turin)

(c) aemiliae Rocci, 1929 (North Italy; Lombardy: Massalombarde, Modena, Livorne)

(d) reverdini Fruhstorfer, 1908 (West and North Italy: Liguria)

(e) nemorensis Verity, 1919 (Italy:Toscana, Firenze)

(f) linnea Bryk, 1932 (Elba)

(g) creusa Meigen, 1829 (Central Italy: Ventimiglia (Verity, 1950), cf. Eisner, 1974!)

(h) latiaris Stichel, 1907 (south Italy: Calabria, Rome, Monti Albani)

(i) vipsania Hemming, 1941 (:latevittata Verity, 1919, a secondary homonym of latevittata Schultz, 1908: creusaMann, nec Meigen, Dahl MS: polymnia Ragusa, 1906), (Sicily)

(j) polyxena Denis and Schiffermuller, 1775 ( cassandra Menetries (Austria,Hungary, Roumania)

punctata Schultz, 1908

marpha Schultz, 1908

rubra Hoffmann, 1916

bella Neuburger, 1903

bipunctata Cosmovici, 1892

meridionalis Hoffmann, 1916

tripunctata Zelezny, 1916

nora Schultz, 1908 (kreusa Domala)

muelleri Bryk, 1921 (mulleri Bryk, recte muelleri)

springeri Ronnicke, 1906

reducta Zelezny, 1915

nigromaculata Zelezny, 1915

unimaculata Zelezny, 1915

demaculata Schultz, 1908

confluens Schultz, 1908

lateviltata Schultz, 1908

fasciata Begrer, 1919

skalae Zelezny, 1917

derubescens Zullich, 1928

alba Esper, 1805

subalba Schultz, 1908

rufescens Oberthur, 1879

meta Meigen

ochracea Staudinger, 1861

cellopura Eisner

basinigra Eisner

quincunx Eisner

divisa Schultz, 1908

ornata Eisner

vitrina Rothschild

irregularis Holland

(k) petrii Bryk (South Ukrainskaja, SSR: Cherson, Kiev, lower Juzinyi Bug, Krim)

(l) thesto Fruhstorfer, 1908 (South-West U.S.S.R.: lowerVolga, Saratow, lower Don

rufescens Oberthür , 1879

muelleri Bryk, 1991

cellopura Eisner

(m) demnosia Freyer, 1833 (Dahl MS) ( albanica Riemel , thusnelda Schultz) (= ssp. macedonia Eisner, 1974 (Macedonia , Dalmatia Albania , Greece Macedonia, Thessalia)

chrysocoma Niepelt

quincunx Eisner

vitrina Rothschild

ochracea Staudinger, 1861

(n) bryki Eisner, 1954 (Montenegro, Herzegovina border)

(o) bosniensis Eisner, 1974 (Bosnia)

ochracea Staudinger, 1861

reducta Zelzny, 1915

(p) polymnia Milliere, 1880 (Greece: Euboea)

(q) taygetana von Rosen, 1929 (Greece: Taygetus,Peleponesos)

(r) gracilis Schultz, 1908 (North-east Turkey: Brusa, Karagja Dagh)

(s) idaensis Eisner, 1974 (Crete)

rumina Linne, 1758

(a) australis Esper, 1780 ( medesicaste Hoffmannsegg) (South France: Provence - East Pyrenees)

forms

alicea Neuburger, 1903

paucipunctata Neuburger

tristis Verity

divisa Schultez

hartmanni Staudfuss, 1896

unipunctata Eisner, 1954

minusculus Eisner

vitrina Rothschild sensu Eisner

aperta Eisner, 1954

quincunx Eisner

honnerathii Boisduval , 1832

albescens Eisner

binaria Eisner

(b) castiliana Ruhl, 1892 (transcastilia Mon) (Central Spain: Castilia, Albaracin)

forms

derubescens Schultz

honnorathii Boisduval, 1832 (henrietta Timmins)

tristis Verity, 1906

semitristis de Sagarra, 1930

ornatissima Blachier, 1908

nigricans Holland

alicea Neuburger, 1903

(c) catalonica de Sagarra 1930 (Spain: Catalonia)

semitristis de Sagarra, 1930

(d) petheri Romei, 1927 (South Spain: Sierra Nevada)

(e) minima Gerhardinger, 1951 Spain: Toledo)

(g)cantabrica Gomez-Bustille (Spain: Cantabrica)

(h)rumina Linnaeus (andalusica Ribbe 1910) (South Spain: Andalusia, South Huelva, Cadiz, Gibraltar, Malaga, Seville)

forms

tristis Verity

semitristis de Sagorra, 1930

xanthe Schultz, 1908 (mackeri Holland, 1910)

rubistriga Bryk

honorathii Boisduval, 1832

ochracea Staudinger, 1861

medicaste Illigner,

andalusica Ribbe, 1910

paucipunctata Neuburger

minusculus Eisner

ornatior Blachier, 1905

(i) lusitanica Bryk, 1932 (Portugal:Lissabon, Algarve, Cintra, Belem , Oporto, Leca)

semitristis de Sagarra, 1930

paucipunctata Neuburger

ornatissima Blachier, 1908

nigricans Eisner

minusculus Eisner

aperta Eisner

divisa Schultz, 1908

canteneri Staudinger, 1861

(j) ornatior Blachier, 1905 ( mauretanica Schultz, 1908, africana Stichel, 1907), (N Africa: Algeria, Morocco: Tangier)

The name ornatior is not the correct one for the N African representative. canteneri (Heydenrich i.l.) Staudinger, 1861 (= canteneri Heyd. 1851, was originally applied to African populations and is the valid name , cf Felder and Felder (1864), Rothschild (1917), canteneri is generally considered a European form but the type locality was fixed 'Algeria' by the Felder brothers (l.c.) whereas Staudinger (1861) only, and apparently erroneously, referred the name to the Iberian form ochracea. Manley and Allcard (1970) following Bryk (1934) referred to canteneri Staudinger as an orange aberration of both sexes which is frequent in Morocco but occasionally appears in warmer parts of South Spain.

forms

irregularis Holland, 1912

distorta Rothschild, 1918

ornatissima Blachier, 1908

nebulosa Holland, 1912

xanthe Schultze, 1908

honorathii Boisduval, 1832

poujadei Thierry-Meg, 1910

canteneri Staudinger, 1861

nigricans Holland, 1912

minusculus Eisner

posteriorrubromarginalis Eisner

paucipunctata Neuburger

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