Portal:Devonian

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The Devonian Portal

Map of Earth as it appeared 400 million years ago during the Early Devonian, Emsian stage

The Devonian (/dəˈvoʊni.ən, dɛ-/ də-VOH-nee-ən, deh-) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at 419.2 million years ago (Ma), to the beginning of the succeeding Carboniferous period at 358.9 Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied.

The first significant evolutionary radiation of life on land occurred during the Devonian, as free-sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants (pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolution. The earliest land animals, predominantly arthropods such as myriapods, arachnids and hexapods, also became well-established early in this period, after beginning their colonization of land at least from the Ordovician period.

Fishes, especially jawed fish, reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The armored placoderms began dominating almost every known aquatic environment. In the oceans, cartilaginous fishes such as primitive sharks became more numerous than in the Silurian and Late Ordovician. Tetrapodomorphs, which include the ancestors of all four-limbed vertebrates (i.e. tetrapods), began diverging from freshwater lobe-finned fish as their more robust and muscled pectoral and pelvic fins gradually evolved into forelimbs and hindlimbs, though they were not fully established for life on land until the Late Carboniferous. (Full article...)

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Selected Devonian Article

Marcellus shale exposure above Marcellus, N.Y. The vertical joints create sheer cliff faces.

The Marcellus Formation or the Marcellus Shale is a Middle Devonian age unit of sedimentary rock found in eastern North America. Named for a distinctive outcrop near the village of Marcellus, New York, in the United States, it extends throughout much of the Appalachian Basin.

The unit name usage by the U.S. Geological Survey (USGS) includes Marcellus Shale and Marcellus Formation. The term "Marcellus Shale" is the preferred name throughout most of the Appalachian region, although the term "Marcellus Formation" is also acceptable within the State of Pennsylvania. The unit was first described and named as the "Marcellus shales" by J. Hall in 1839. (Full article...)

List of selected Devonian articles
Sarcopterygii Trilobites Late Devonian extinction Acadian Orogeny Bactritida List of Early Devonian land plants Dunkleosteus

Selected Devonian land plant article

Archaeopteris fossil leaves

The progymnosperms are an extinct group of woody, spore-bearing plants that is presumed to have evolved from the trimerophytes, and eventually gave rise to the gymnosperms, ancestral to acrogymnosperms and angiosperms (flowering plants). They have been treated formally at the rank of division Progymnospermophyta or class Progymnospermopsida (as opposite). The stratigraphically oldest known examples belong to the Middle Devonian order the Aneurophytales, with forms such as Protopteridium, in which the vegetative organs consisted of relatively loose clusters of axes. Tetraxylopteris is another example of a genus lacking leaves. In more advanced aneurophytaleans such as Aneurophyton these vegetative organs started to look rather more like fronds, and eventually during Late Devonian times the aneurophytaleans are presumed to have given rise to the pteridosperm order, the Lyginopteridales. In Late Devonian times, another group of progymnosperms gave rise to the first really large trees known as Archaeopteris. The latest surviving group of progymnosperms is the Noeggerathiales, which persisted until the end of the Permian.

Other characteristics:

Vascular cambium with unlimited growth potential is present as well as xylem and phloem.
Ancestors of the earliest seed plants as well as the first true trees.

Strong monopodial growth is exhibited. (Full article...)

List of selected Devonian articles
pteridophytes Sigillaria Pteridospermatophyta Lycophyte Equisetidae Rhyniophyte Trimerophytopsida Lycopodiopsida

Selected Devonian formation

The Hamilton Group is a Devonian-age geological group which is located in the Appalachian region of the United States. It is present in New York, Pennsylvania, Maryland, Ohio, West Virginia, northwestern Virginia and Ontario, Canada, and is mainly composed of marine shale with some sandstone.

There are two main formations encompassed by the group: the Mahantango Formation and the Marcellus Shale. In southwestern Virginia, where the two sub-units are not easily distinguishable, the Hamilton Group is broadly equivalent to the Millboro Shale or Millboro Formation.

The group is named for the village of Hamilton, New York. These rocks are the oldest strata of the Devonian gas shale sequence. (Full article...)

List of selected Devonian formation articles
Tully Formation Tioga Bentonites Windom shale Rhinestreet Formation Oriskany Formation Bakken formation Traverse Group New Albany Shale Old Red Sandstone Grey Downtonian

Selected Devonian fish article

Fossil of Bothriolepis panderi showing its caliper-like pectoral fins

Placoderms (from Greek πλάξ (plax, plakos) 'plate' and δέρμα (derma) 'skin') are vertebrate animals of the class Placodermi, an extinct group of prehistoric fish known from Paleozoic fossils during the Silurian and the Devonian periods. While their endoskeletons are mainly cartilaginous, their head and thorax were covered by articulated armoured plates (hence the name), and the rest of the body was scaled or naked depending on the species.

Placoderms were among the first jawed fish (their jaws likely evolved from the first pair of gill arches), as well as the first vertebrates to have true teeth. They were also the first fish clade to develop pelvic fins, the second set of paired fins and the homologous precursor to hindlimbs in tetrapods. 380-million-year-old fossils of three other genera, Incisoscutum, Materpiscis and Austroptyctodus, represent the oldest known examples of live birth.

Placoderms are thought to be paraphyletic, consisting of several distinct outgroups or sister taxa to all living jawed vertebrates, which originated among their ranks. In contrast, one 2016 analysis concluded that Placodermi is likely monophyletic. (Full article...)

List of selected Devonian fish articles
Ctenaspis Brochoadmones Cheirolepis Holoptychius Hyneria Lepidaspis Scaumenacia Tinirau clackae Diplocercides Dipterus Elpistostegalia Gooloogongia Ventastega Tristichopteridae Phoebodus Stethacanthidae Xenacanthus

Selected Devonian invertebrate

Top view of a stromatoporoid with prominent mamelons. From the Columbus Limestone (Middle Devonian) of Ohio.

Stromatoporoidea is an extinct clade of sea sponges common in the fossil record from the Middle Ordovician to the Late Devonian. They can be characterized by their densely layered calcite skeletons lacking spicules. Stromatoporoids were among the most abundant and important reef-builders of their time, living close together in flat biostromes or elevated bioherms on soft tropical carbonate platforms.

Externally, some species have raised bumps (mamelons) and star-shaped crevices (astrorhizae), which together help vent exhalant water away from the living surface. Internally, stromatoporoids have a mesh-like skeletal system combining extensive horizontal layers (laminae), vertical rods (pillars), and boxy spaces (galleries), along with other features. The most common growth forms range from laminar (flattened) to domical (dome-shaped). Spheroidal, finger-like, or tree-like species also occur, though they are rare in most environments.

Stromatoporoids competed and coexisted with other reef-builders such as tabulate and rugose corals. Some stromatoporoid species are useful as environmental proxies, since their form and distribution can help approximate the depositional environment of sedimentary strata. They hosted a diverse fauna of encrusting symbionts both within and outside their skeletons. Some studies have argued that stromatoporoids were mixotrophs (engaged in a mutualistic relationship with photosynthetic algae), similar to modern scleractinian corals. Though this hypothesis is plausible, circumstantial evidence is inconclusive. (Full article...)