Paoliida

Paoliida Temporal range:
Fossils of Paoliida. A: Zdenekia silesiensis; B: Darekia sanguinea; C, D1, D2: Paoliidae gen. et sp. indet.
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Cohort:	Polyneoptera
Order:	†Paoliida Handlirsch, 1906
Families
†Anthracoptilidae †Blattinopsidae †Paoliidae
Synonyms
Protoptera Rasnitsyn, 1977

Paoliida is an extinct order of winged insects that lived in the late Paleozoic. Historically, both their systematic position and composition were controversial – for instance they had been considered as palaeodictyopterans, as basal Neoptera, or as stem-group of Pterygota – but recent studies have resolved them as the sister group of Dictyoptera.

Families and genera

After Prokop et al. (2014), Guan et al. (2016) and Nel & Poschmann (2021):

• †Anthracoptilidae Handlirsch, 1922 (Synonyms: Strephocladidae Martynov, 1938; Strephoneuridae Martynov, 1940; ?Adiphlebiidae Handlirsch, 1906)
  • ?†Adiphlebia Scudder, 1885
  • †Afrocladus Nel, Garrouste & Prokop, 2015
  • †Anthracoptilus Lameere, 1917
  • †Carrizocladus Rasnitsyn in Rasnitsyn et al., 2004
  • †Graticladus Novokshonov and Aristov, 2004
  • †Homocladus Carpenter, 1966
  • †Jarmilacladus Rasnitsyn and Aristov, 2004
  • †Lodevocladus Prokop et al., 2015
  • †Mesoptilus Lameere, 1917
  • †Mycteroptila Rasnitsyn, 1977
  • †Paracladus Carpenter, 1966
  • †Pruvostia Bolton, 1921
  • †Pseudomesoptilus Guan et al., 2016
  • †Rhinomaloptila Rasnitsyn, 1977
  • †Spargoptilon Kukalová, 1965
  • †Strephocladus Scudder, 1885
  • †Strephoneura Martynov, 1940
  • †Strephoptilus Rasnitsyn and Aristov, 2013
  • †Westphaloptilus Guan et al., 2016
• †Blattinopsidae Bolton, 1925 (Synonym: Klebsiellidae Handlirsch, 1906)
  • †Alienus Handlirsch, 1906
  • †Avionblattinopsis Quispe et al., 2021
  • †Balduriella Meunier, 1925
  • †Blattinopsis Giebel, 1867
  • †Glaphyrophlebia Handlirsch, 1906
  • †Klebsiella Meunier, 1908
  • †Microblattina Scudder, 1895
  • †Protoblattiniella Meunier, 1912
  • †Rhipidioptera Brongniart, 1894
  • †Stephanopsis Kukalová, 1958
  • †Westphaloblattinopsis Béthoux and Jarzembowski, 2010
• †Paoliidae Handlirsch, 1906 (Synonyms: Ideliidae Zalessky, 1928; Herbstialidae Schmidt, 1953; Permulidae Aristov & Storozhenko, 2011; Permotermopsidae Martynov, 1937)
  • †Acropermula Aristov and Storozhenko, 2011
  • †Aenigmidelia Sharov, 1961
  • †Archidelia Sharov, 1961
  • †Carbonidelia Nel, Garrouste & Jouault, 2023
  • †Darekia Prokop et al., 2012
  • †Herbstiala Schmidt, 1952
  • †Holasicia Kukalová, 1958
  • †Idelina Storozhenko, 1992
  • †Kemperala Brauckmann, 1984
  • †Khosaridelia Storozhenko, 1992
  • †Kortshakolia Sharov, 1961
  • †Kochopteron Brauckmann, 1984
  • †Mertovia Prokop and Nel, 2007
  • †Micaidelia Aristov 2004
  • †Mongoloidelia Storozhenko, 1992
  • †Olinka Kukalová, 1958
  • ?†Paolekia Riek, 1976
  • †Paolia Smith, 1871
  • †Paoliola Handlirsch, 1919
  • †Permotermopsis Martynov, 1937
  • †Permula Handlirsch, 1919
  • †Protoblattina Meunier, 1909
  • †Pseudofouquea Handlirsch, 1906
  • †Silesiapteron Prokop et al., 2013
  • †Simplexpaolia Santos et al., 2023
  • †Sojanidelia Storozhenko, 1992
  • †Stenaropodites Martynov, 1928
  • †Sustaia Kukalová, 1958
  • †Sylvidelia Martynov, 1940
  • †Zdenekia Kukalová, 1958