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Indian leopard

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Indian leopard
Male In Nagarhole National Park
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Pantherinae
Genus: Panthera
Species:
P. pardus
Subspecies:
P. p. fusca
Trinomial name
Panthera pardus fusca
(Meyer, 1794)
Synonyms

The Indian leopard (Panthera pardus fusca) is a subspecies of the leopard (P. pardus) widely distributed on the Indian subcontinent. It is threatened by illegal trade of skins and body parts, and persecution due to human-leopard conflict and retaliation for livestock depredation.

Taxonomy

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Felis fusca was the scientific name proposed by Friedrich Albrecht Anton Meyer in 1794 who described a black leopard from Bengal that was on display at the Tower of London.[2] Leopardus perniger proposed by Brian Houghton Hodgson in 1863 were five leopard skins from Nepal, out of which three were black. He mentioned Sikkim and Nepal as habitat.[3] Panthera pardus millardi proposed by Reginald Innes Pocock in 1930 was a single leopard skin and skull from Kashmir. It differed from typical P. p. fusca skins by longer hair and a more greyish colour.[4]

Since leopard populations in Nepal, Sikkim and Kashmir are not geographically isolated from leopard populations in the Indian subcontinent, they were subsumed to P. p. fusca in 1996. The Indus River in the west and the Himalayas in the north form topographical barriers to the dispersal of this subspecies.[5][6] In the east, the Ganges Delta and the lower course of the Brahmaputra River are thought to form natural barriers to the range of the Indochinese leopard.[6]

A genetic analysis of 49 leopard skin samples collected in Azad Jammu Kashmir and Galyat regions of northern Pakistan revealed haplotypes of both Persian and Indian leopards, indicating intergradation of both subspecies in this region.[7]

Characteristics

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Leopard in Nagarhole National Park

The Indian leopard has strong legs and a long, well-formed tail, broad muzzle, short ears, small, yellowish-grey eyes, and light-grey ocular bulbs.[2] Its coat is spotted and rosetted on a pale yellow to yellowish-brown or golden background, except for the melanistic forms; the spots fade toward the white underbelly and the insides and lower parts of the legs. Rosettes are most prominent on the back, flanks and hindquarters. The pattern of the rosettes is unique to each individual.[8][9] Juveniles have woolly fur, and appear dark due to the densely arranged spots. The white-tipped tail is 60–100 cm (24–39 in) long, white underneath, and displays rosettes, which form incomplete bands toward the end. The rosettes are larger in other leopard subspecies in Asia. Fur colour tends to be more pale and cream in arid habitats, more grey in colder climates, and of a darker golden hue in rainforest habitats.[10]

The clouded leopard can be told apart by its diffuse "clouds" of spots compared to the smaller and distinct rosettes of the leopard, longer legs and thinner tail.[11]

Skull

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The largest skull for an Indian leopard was recorded in 1920, and belonged to a large, melanistic cat in the area of Ootacamund, Tamil Nadu. The black panther was said to have bigger forelimbs and forequarters than hind-limbs and hind-quarters, with a skull and claws nearly as large as those of a tigress. The skull measured 11.2 in (28 cm) in basal length, and 7.9 in (20 cm) in breadth, and weighed 2 lb 4 oz (1,000 g). By comparison, the skull of one western African leopard measured 11.25 in (28.6 cm) in basal length, and 7.125 in (18.10 cm) in breadth, and weighed 1 lb 12 oz (790 g).[12]

Size

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Male Indian leopards grow to between 127 cm (4 ft 2 in) and 142 cm (4 ft 8 in) in body size with a 76 cm (2 ft 6 in) to 91 cm (3 ft) long tail and weigh between 50 and 77 kg (110 and 170 lb). Females are smaller, growing to between 104 cm (3 ft 5 in) and 117 cm (3 ft 10 in) in body size with a 76 cm (2 ft 6 in) to 87.6 cm (2 ft 10.5 in) long tail, and weigh between 29 and 34 kg (64 and 75 lb). Sexually dimorphic, males are larger and heavier than females.[8]

The largest wild individual appears to have been a male man-eater that was shot in the Dhadhol area of Bilaspur district, Himachal Pradesh in 2016. It reportedly measured 262 cm (8 ft 7 in) from head to tail, 86 cm (34 in) at the shoulder, and weighed 71 kg (157 lb).[13]

Distribution and habitat

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A leopard in Satpura National Park, India
In Gir National Park

The Indian leopard is distributed in India, Nepal, Bhutan and parts of Pakistan.[1] Bangladesh has no viable leopard population but there are occasional sightings in the forests of Sylhet, Chittagong Hill Tracts and Cox's Bazar.[14][15] It inhabits tropical rainforests, dry deciduous forests, temperate forests and northern coniferous forests but does not occur in the mangrove forests of the Sundarbans.[10]

In southern Tibet, it was recorded in Qomolangma National Nature Preserve.[16]

In Nepal's Kanchenjunga Conservation Area, a melanistic leopard was photographed at an elevation of 4,300 m (14,100 ft) by a camera trap in May 2012.[17]

Population in India

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Main article: List of Indian states by wildlife population
Male leopard in Nagarhole National Park

In 2015, 7,910 leopards were estimated to live in and around tiger habitat in India; about 12,000 to 14,000 leopards were speculated to live in the entire country. The following table gives the major leopard populations in the Indian states.[18] As of 2020, the leopard population within forested habitats in India's tiger range landscapes was estimated at 12,172 to 13,535 individuals. Surveyed landscapes included elevations below 2,600 m (8,500 ft) in the Shivalik Hills and Gangetic plains, Central India and Eastern Ghats, Western Ghats, as well as the Brahmaputra River basin and hills in Northeast India.[19]

Leopard population by state
State Leopards (2015)
Andhra Pradesh 343
Bihar 98
Chhattisgarh 846
Goa 86
Gujarat 1,395
Jharkhand 46
Karnataka 1,783
Kerala 650
Madhya Pradesh 3,421
Maharashtra 1,690
Odisha 760
Tamil Nadu 868
Uttar Pradesh 316
Uttarakhand 839
West Bengal 90

Behaviour and ecology

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Leopard with a killed langur
Leopard at Masanakudi

The leopard is elusive, solitary, and largely nocturnal. It is known for its ability in climbing, and has been observed resting on tree branches during the day, dragging its kills up trees and hanging them there, and descending from trees headfirst.[20] It is a powerful swimmer, although is not as disposed to swimming as the tiger. It is very agile, and can run at over 58 kilometres per hour (36 mph), leap over 6 m (20 ft) horizontally, and jump up to 3 m (9.8 ft) vertically.[21] It produces a number of vocalizations, including grunts, roars, growls, meows, and purrs.[22]

In Nepal's Bardia National Park, home ranges of male leopards comprised about 48 km2 (19 sq mi), and of females about 17 km2 (6.6 sq mi); female home ranges decreased to 5 to 7 km2 (1.9 to 2.7 sq mi) when they had young cubs.[23] In Gir National Park, the home range of a male radio-collared leopard was estimated at 28.15 km2 (10.87 sq mi). It killed prey once in 3.7 days.[24]

The leopard is a versatile, opportunistic hunter, and has a very broad diet.[10] It is able to take large prey due to its massive skull and powerful jaw muscles.[25][26] In Sariska Tiger Reserve, the dietary spectrum of the Indian leopard includes axis deer, sambar deer, nilgai, wild boar, common langur, Indian hare and peafowl.[27] In Periyar Tiger Reserve, primates make up a large proportion of its diet.[28]

Reproduction

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Depending on the region, the leopard mates all year round. The estrous cycle lasts about 46 days and the female usually is in heat for 6–7 days.[29] Gestation lasts for 90 to 105 days.[30] Cubs are usually born in a litter of 2–4 cubs.[31] Mortality of cubs is estimated at 41–50% during the first year. Females give birth in a cave, crevice among boulders, hollow tree, or thicket to make a den. Cubs are born with closed eyes, which open four to nine days after birth.[32] The fur of the young tends to be longer and thicker than that of adults. Their pelage is also more grey in colour with less defined spots. Around three months of age, the young begin to follow the mother on hunts. At one year of age, leopard young can probably fend for themselves, but remain with the mother for 18–24 months. The average typical life span of a leopard is between 12 and 17 years.[33]

Sympatric carnivores

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Indian leopards are not common in habitats where tiger density is high, and are wedged between prime tiger habitat on the one side, and cultivated village land on the other.[34] Where the tiger population is high or increasing, tigers drive leopards off to areas located closer to human settlements, like in Nepal's Bardia National Park and Sariska Tiger Reserve.[35][36] Resource partitioning occurs where leopards share their range with tigers. Leopards tend to take smaller prey, usually less than 75 kg (165 lb), where tigers are present.[37] In areas where leopard and tiger are sympatric, coexistence is reportedly not the general rule, with leopards being few where tigers are numerous.[38] The mean leopard density decreased significantly (from 9.76 to 2.07 animals per 100 km2 (39 sq mi)) while the mean density of tigers increased (from 3.31 to 5.81 animals/100 km2) from 2004–2005 to 2008 in Rajaji National Park following the relocation of pastoralists out of the park. There, the two species have high dietary overlap, and an increase in the tiger population resulted in a sharp decrease in the leopard population and a shift in the leopard diet to small prey (from 9% to 36%) and domestic prey (from 6.8% to 31.8%).[39] In Chitwan National Park, leopards killed prey ranging from less than 25–100 kg (55–220 lb) in weight with most kills in the 25–50 kg (55–110 lb) range. Tigers killed more prey in the 50–100 kg (110–220 lb) range. There were also differences in the microhabitat preferences of the individual tiger and leopard followed over five months; the tiger used roads and forested areas more frequently, while the leopard used recently burned areas and open areas more frequently. When a tiger killed baits at sites formerly frequented by leopards, the leopards did not hunt there for some time.[38]

In the tropical forests of India's Nagarhole National Park, tigers selected prey weighing more than 176 kg (388 lb), whereas leopards selected prey in the 30–175 kg (66–386 lb) range.[40] In tropical forests, they do not always avoid the larger cats by hunting at different times. With relatively abundant prey and differences in the size of prey selected, tigers and leopards seem to successfully coexist without competitive exclusion or interspecies dominance hierarchies that may be more common to the leopard's co-existence with the lion in savanna habitats.[41] In areas with high tiger populations, such as in the central parts of India's Kanha National Park, leopards are not permanent residents, but transients. They were common near villages at the periphery of the park and outside the park.[38] In a reserved forest of southern India, species preyed upon by leopard, dhole and striped hyena overlapped considerably.[42]

The leopard and snow leopard both hunt Himalayan tahr and musk deer, but the leopard usually prefers forested habitats located at lower altitudes.[43] Leopard may conflict with sloth bears and can follow them up trees.[44] Bear cubs are probably far more vulnerable and healthy adult bears may be avoided by leopards. One leopard killed a three-quarters grown female sloth bear in an apparently lengthy fight that culminated in the trees. Apparently, a sloth bear killed a leopard in a confrontation in Yala National Park, Sri Lanka but was itself badly injured in the fight and was subsequently put down by park rangers.[45][46]

Threats

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Leopard skins

Hunting of Indian leopards for the illegal wildlife trade is the biggest threat to their survival. They are also threatened by loss of habitat and fragmentation of formerly connected populations, and various levels of human–leopard conflict in human–dominated landscapes.[10]

Several newspapers reported of leopards falling into open wells and being rescued with the help of Forest Department officials.[47][48][49]

Poaching

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A significant immediate threat to wild leopard populations is the illegal trade in poached skins and body parts between India, Nepal and China. The governments of these countries have failed to implement adequate enforcement response, and wildlife crime remained a low priority in terms of political commitment and investment for years. There are well-organised gangs of professional poachers, who move from place to place and set up camp in vulnerable areas. Skins are rough-cured in the field and handed over to dealers, who send them for further treatment to Indian tanning centres. Buyers choose the skins from dealers or tanneries and smuggle them through a complex interlinking network to markets outside India, mainly in China.[50] Seized skins in Kathmandu confirm the city's role as a key staging point for illegal skins smuggled from India bound for Tibet and China.[51]

It is likely that seizures represent a tiny fraction of the total illegal trade, with the majority of smuggled skins reaching their intended end market.[50] Seizures revealed:

  • in India: more than 200 leopards killed by humans every year,[52][53] leopards in India is 7 times more likely to be killed than Indian tigers.[53] WPSI reported that during 1994-2010 at least 3,189 leopards were killed,[53][54][55][56][57][58] then again in 2002-2010 period at least 200 or four leopards per week were reportedly killed by poachers for illegal trade.[52] For every tiger skin, there are at least seven leopard skins in the haul.[53]
  • in Nepal: more than 40 leopards were reported killed by humans every year, e.g. 243 poached leopards between May 2002 and May 2008;[50][51][59][60][61]
  • in China and Tibet: nearly 130 leopards were killed every year, e.g. more than 774 poached leopards between July 1999 and September 2005.[50][51]

Human–leopard conflict

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Causes of conflict

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Expansion of agriculturally used land, encroachment by humans and their livestock into protected areas are main factors contributing to habitat loss and decrease of wild prey. As a result, leopards approach human settlements, where they are tempted to prey on dogs, pigs and goats – domestic livestock, which constitutes an important part of their diet, if they live on the periphery of human habitations. Human–leopard conflict situations ensue, and have increased in recent years. In retaliation for attacks on livestock, leopards are shot, poisoned and trapped in snares. The leopards are considered to be unwanted trespassers by villagers. Conservationists criticize these actions, claiming that people are encroaching on the leopard's native habitat.[62][63] India's Forest Department is entitled to set up traps only in cases of a leopard having attacked humans. If only the presence of a crowd of people prevents the leopard from escaping, then the crowd has to be dispersed and the animal allowed to escape.[64]

As urban areas expanded, the natural habitats of leopards shrunk resulting in leopards venturing into urbanized areas due to easy access of domestic food sources.[65] Karnataka has a high number of such conflicts.[66][67] In recent years, leopards were sighted in Bangalore, and the forest department captured six leopards in the city's outskirts, relocated four of them to various other locations.[68]

Man-eater leopards

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Main article: Leopard attack
The Panar Leopard killed by Jim Corbett

Every year more leopards are killed by humans than the humans killed by leopards. On average nearly 400 leopards are reported killed yearly in India, Nepal and China combined based on the leopard skins caught from the poachers,[52][53][51][59][50] though the actual number of leopards killed by humans is likely to be several times higher.[50] In and around the Shivalik hills of Himachal Pradesh alone, 68 leopards were killed by people between 2001 and 2013, of which only 10 were man-eaters.[69]

The frequency of Leopard attacks on humans varies by geographical region and historical period. Since India and Nepal have the majority of Indian leopards population, consequently attacks are regularly reported only from India and Nepal.[70][71] Among the five "big cats", leopards are less likely to become man-eaters—only jaguars and snow leopards have a less fearsome reputation.[72][73] While leopards generally avoid humans, they tolerate proximity to humans better than lions and tigers and often come into conflict with humans when raiding livestock.[74]

Attacks in India are still reported, since leopards population in India outnumber population of all other large carnivores combined, consequently the number of humans killed by leopards is also more than those killed by all other large carnivores combined.[75][76]

In Nepal, where most attacks occur in the midland regions, i.e. in the Terai, midhills, and lesser Himalaya, the rate of leopard predation on humans results in approximately 1.9 human deaths annually per million inhabitants. .[71]

Historically, with rapid urbanization in late 19th and early 20th centuries, leopard attacks may have peaked in India during those times.[72] Notable man-eaters of that era include Leopard of Central Provinces, Rudraprayag, Gummalapur, Yellagiri Hills, Golis Range and Panar.[77]

Ways to minimise conflict

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Key to avoiding conflict or leopard's predation of humans is to shift the focus on human's behavioral change to minimise the chances of a leopard encounter or attack, which can be achieved by "clearing bushes and overgrowth around homes to minimise hiding spaces for leopards, leaving a light on at night to deter them, and ensuring people, especially children, did not go out alone at night." Leopards are shy and avoid humans and are more active at night, during encounter with leopards "give way to the leopard and move away calmly" and alert the forest department immediately.[78]

Conservation

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A captive leopard

Panthera pardus is listed in CITES Appendix I.[1] Despite India and Nepal being contracting parties to CITES, national legislation of both countries does not incorporate and address the spirit and concerns of CITES. Trained human resources, basic facilities and effective networks for control of poaching and trade in wildlife are lacking.[59] The Indian leopard is considered Vulnerable in India,[79] Bhutan,[80] and Nepal[81] but Critically Endangered in Pakistan.[82]

Frederick Walter Champion was one of the first in India who after World War I advocated for the conservation of leopards, condemned sport hunting and recognised their key role in the ecosystem.[83] Billy Arjan Singh championed their cause since the early 1970s.[84]

There are a few leopard rescue centres in India, such as the Manikdoh Leopard Rescue Centre in Junnar,[85] but more rescue and rehabilitation centres are being planned.[86] Some wildlife experts think that such centres are not an ideal solution, but that conflict resolution by way of changing human behaviour, land use or grazing patterns and implementing responsible forest management to lessen human-animal conflict would be far more effective to conserve leopards.[87]

In culture and literature

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An Indian leopard used for hunting, probably early 20th century
Cajetan Lobo with two pet Indian leopards
See also: Black panther § Culture and literature

See also

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References

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  1. ^ a b c Shivakumar, S.; Khettry, A.; Surve, N.; Rahman, H.; Ghimirey, Y.; Tharchen, L.; Zaw, T.; Waseem, M. & Jhala, Y. (2023). "Panthera pardus ssp. fusca". IUCN Red List of Threatened Species. 2023: e.T215195524A215195533.
  2. ^ a b Meyer, F. A. A. (1794). "Über de la Metheries schwarzen Panther". Zoologische Annalen. Erster Band. Weimar: Im Verlage des Industrie-Comptoirs. pp. 394–396.
  3. ^ Hodgson, B. H. (1863). "Leopardus perniger, Hodgson". Catalogue of the Specimens and Drawings of Mammalia, Birds, Reptiles and Fishes of Nepal and Tibet. London: British Museum.
  4. ^ Pocock, R. I. (1930). "The Panthers and Ounces of Asia". Journal of the Bombay Natural History Society. 34 (2): 307–336.
  5. ^ Miththapala, S.; Seidensticker, J.; O'Brien, S. J. (1996). "Phylogeographic Subspecies Recognition in Leopards (Panthera pardus): Molecular Genetic Variation". Conservation Biology. 10 (4): 1115–1132. doi:10.1046/j.1523-1739.1996.10041115.x.
  6. ^ a b Uphyrkina, O.; Johnson, E.W.; Quigley, H.; Miquelle, D.; Marker, L.; Bush, M. & O'Brien, S. J. (2001). "Phylogenetics, genome diversity and origin of modern leopard, Panthera pardus" (PDF). Molecular Ecology. 10 (11): 2617–2633. doi:10.1046/j.0962-1083.2001.01350.x. PMID 11883877. S2CID 304770. Archived from the original (PDF) on 28 April 2020. Retrieved 2 December 2012.
  7. ^ Asad, M.; Martoni, F.; Ross, J.G.; Waseem, M. & Paterson, A.M. (2019). "Assessing subspecies status of leopards (Panthera pardus) of northern Pakistan using mitochondrial DNA". PeerJ. 7: e7243. doi:10.7717/peerj.7243. hdl:10182/10848.
  8. ^ a b Pocock, R. I. (1939). "Panthera pardus". The fauna of British India, including Ceylon and Burma. Mammalia. Vol. 1. London: Taylor and Francis. pp. 222–239.
  9. ^ Menon, V. (2014). Indian Mammals: A Field Guide. Gurgaon, India: Hachette India. ISBN 978-93-5009-761-8.
  10. ^ a b c d Nowell, K.; Jackson, P. (1996). "Leopard Panthera pardus (Linnaeus, 1758)". Wild Cats: status survey and conservation action plan. Gland, Switzerland: IUCN/SSC Cat Specialist Group. pp. 24–30. Archived from the original on 22 February 2014.
  11. ^ Francis, C.M. (2008). A Field Guide to the Mammals of Southeast Asia. London, UK: New Holland. p. 296. ISBN 978-1-84537-735-9.
  12. ^ Prater, S. H. (1921). "Record Panther Skull (P. p. pardus)". The Journal of the Bombay Natural History Society. XXVII (IV): 933–935. Archived from the original on 2 May 2019. Retrieved 24 August 2019.
  13. ^ Chauhan, P. (2016). "Leopard shot in Bilaspur turns out to be a record breaker". The Tribune. Archived from the original on 26 March 2019. Retrieved 24 March 2018.
  14. ^ Khan, M. A. R. (2009). "Endangered mammals of Bangladesh". Oryx. 18 (3): 152–156. doi:10.1017/S0030605300019001.
  15. ^ Kabir, M.T.; Ahsan, M.F.; Khatoon, A. (2017). "Occurrence and conservation of the Indian Leopard (Mammalia: Carnivora: Felidae: Panthera pardus) in Cox's Bazar District of Bangladesh". Journal of Threatened Taxa. 9 (6): 10320–10324. doi:10.11609/jott.1898.9.6.10320-10324.
  16. ^ Laguardia, A.; Kamler, J. F.; Li, S.; Zhang, C.; Zhou, Z. & Shi, K. (2017). "The current distribution and status of leopards Panthera pardus in China". Oryx. 51 (1): 153−159. doi:10.1017/S0030605315000988.
  17. ^ Thapa, K.; Pradhan, N. M. B.; Berker, J.; Dhakal, M.; Bhandari, A. R.; Gurung, G. S.; Rai, D. P.; Thapa, G. J.; Shrestha, S. & Singh, G. R. (2013). "High elevation record of a leopard cat in the Kangchenjunga Conservation Area, Nepal". Cat News (58): 26–27.
  18. ^ Bhattacharya, A. (2015). "Finally, India gets a count of its leopard numbers: 12,000-14,000". The Times of India. Archived from the original on 27 November 2015. Retrieved 20 February 2016.
  19. ^ Jhala, Y.V.; Qureshi, Q. & Yadav, S.P. (2020). Status of leopards in India, 2018. Technical Report TR/2020/16 (Report). New Delhi and Dehradun: National Tiger Conservation Authority, Government of India and Wildlife Institute of India.
  20. ^ Jerdon, T.C. (1874). Mammals of India: a natural history of the animals known to inhabit continental India. John Wheldon, London.
  21. ^ "Animal bytes – Panthera pardus". Sea World. Archived from the original on 24 June 2008. Retrieved 6 June 2008.
  22. ^ Estes, R. (1991). The Behavior Guide to African Mammals, Including Hoofed Mammals, Carnivores, Primates. Los Angeles: The University of California Press. ISBN 978-0520080850.
  23. ^ Odden, M.; Wegge, P. (2005). "Spacing and activity patterns of leopards Panthera pardus in the Royal Bardia National Park, Nepal" (PDF). Wildlife Biology. 11 (2): 145–152. doi:10.2981/0909-6396(2005)11[145:SAAPOL]2.0.CO;2. S2CID 86140708. Archived from the original (PDF) on 28 July 2011. Retrieved 24 February 2011.
  24. ^ Zehra, N.; Chaudhary, R.; Khan, J. A. "Ecology of Leopard (Panthera pardus fusca Meyer) in Dry Tropical Forests of Gir National Park and Sanctuary, Gujarat, India". International Journal of Ecology and Environmental Sciences. 45 (3): 241–255.
  25. ^ Burnie, D; Wilson, D. E. (2001). Animal: The Definitive Visual Guide to the World's Wildlife'. DK Adult. ISBN 978-0789477644.
  26. ^ Boitani, L. (1984). Simon & Schuster's Guide to Mammals. Simon & Schuster/Touchstone Books. ISBN 978-0-671-42805-1.
  27. ^ Mondal, K.; Gupta, S.; Bhattacharjee, S.; Qureshi, Q. & K. Sankar (2012). "Prey selection, food habits and dietary overlap between leopard Panthera pardus (Mammalia: Carnivora) and re-introduced tiger Panthera tigris (Mammalia: Carnivora) in a semi-arid forest of Sariska Tiger Reserve, Western India". Italian Journal of Zoology. 79 (4): 607–616. doi:10.1080/11250003.2012.687402.
  28. ^ Srivastava, K. K.; Bhardwaj, A. K.; Abraham, C. J.; Zacharias, V. J. (1996). "Food habits of mammalian predators in Periyar Tiger Reserve, South India". The Indian Forester. 122 (10): 877–883. Archived from the original on 20 January 2015. Retrieved 22 March 2013.
  29. ^ Sadleir, R. (1966). "Notes on the Reproduction of the larger Felidae". International Zoo Yearbook. 6: 184–187. doi:10.1111/j.1748-1090.1966.tb01746.x.
  30. ^ Hemmer, H. (1976). "Gestation period and postnatal development in felids". In R.L. Eaton (ed.). The World's Cats. Vol. 3. Carnivore Research Institute. pp. 143–165.
  31. ^ Eaton, R.L. (1977). "Reproductive biology of the leopard". Zoologischer Garten. 47 (5): 329–351.
  32. ^ Sunquist, M. E.; Sunquist, F. (2002). "LeopardPanthera pardus". Wild Cats of the World. Chicago: University of Chicago Press. pp. 318–342. ISBN 978-0-226-77999-7.
  33. ^ "Leopard (Panthera pardus); Physical characteristics and distribution". Comparative Mammalian Brain Collections. Archived from the original on 3 January 2011. Retrieved 15 May 2016.
  34. ^ McDougal, C. (1988). "Leopard and Tiger Interactions at Royal Chitwan National Park, Nepal". Journal of the Bombay Natural History Society. 85: 609–610. Archived from the original on 26 March 2019. Retrieved 21 September 2018.
  35. ^ Odden, M.; Wegge, P.; Fredriksen, T. (2010). "Do tigers displace leopards? If so, why?". Ecological Research. 25 (4): 875–881. doi:10.1007/s11284-010-0723-1. S2CID 19799372.
  36. ^ Rajvi, A. S. (2016). Tigress kills leopard at Sariska Reserve in India (Motion picture). Barcroft TV. Archived from the original on 8 April 2019. Retrieved 8 April 2019.
  37. ^ Nowell, K. & Jackson, P. (1996). "Leopard Panthera pardus (Linnaeus, 1758)". Wild Cats: status survey and conservation action plan. Gland, Switzerland: IUCN/SSC Cat Specialist Group. Archived from the original on 22 February 2014.
  38. ^ a b c Seidensticker, J. (1976). "On the ecological separation between tigers and leopards" (PDF). Biotropica. 8 (4): 225–234. doi:10.2307/2989714. JSTOR 2989714. Archived (PDF) from the original on 19 July 2021. Retrieved 19 July 2021.
  39. ^ Harihar, A.; Pandav, B.; Goyal, S. P. (2011). "Responses of leopard Panthera pardus to the recovery of a tiger Panthera tigris population". Journal of Applied Ecology. 48 (3): 806–814. doi:10.1111/j.1365-2664.2011.01981.x.
  40. ^ Karanth, K. U.; Sunquist, M. E. (1995). "Prey selection by tiger, leopard and dhole in tropical forests". Journal of Animal Ecology. 64 (4): 439–450. doi:10.2307/5647. JSTOR 5647.
  41. ^ Karanth, U. K.; Sunquist, M. E. (2000). "Behavioural correlates of predation by tiger (Panthera tigris), leopard (Panthera pardus) and dhole (Cuon alpinus) in Nagarahole, India". Journal of Zoology. 250 (2): 255–265. doi:10.1111/j.1469-7998.2000.tb01076.x.
  42. ^ Arivazhagan, C.; Arumugam, R.; Thiyagesan, K. (2007). "Food habits of leopard (Panthera pardus fusca), dhole (Cuon alpinus) and striped hyena (Hyaena hyaena) in a tropical dry thorn forest of southern India" (PDF). Journal of the Bombay National Historical Society. 104 (2). Archived from the original (PDF) on 4 March 2009.
  43. ^ Lovari, S.; Minder, I.; Ferretti, F.; Mucci, N.; Randi, E.; Pellizzi, B. (2013). "Common and snow leopards share prey, but not habitats: competition avoidance by large predators?". Journal of Zoology. 291 (2): 127–135. doi:10.1111/jzo.12053.
  44. ^ Hadley, B. (21 December 2008), The Sloth Bear (PDF), Bear Specialist Group, archived from the original (PDF) on 21 December 2008
  45. ^ Baskaran, N., Sivaganesan, N., & Krishnamoorthy, J. (1997). Food habits of sloth bear in Mudumalai wildlife sanctuary, Tamil Nadu, southern India. JOURNAL-BOMBAY NATURAL HISTORY SOCIETY, 94, 1–9.
  46. ^ Kurt, F., & Jayasuriya, A. (1968). Notes on a dead bear. Loris, 11, 182–183.
  47. ^ "Watch Villagers Save Drowning Leopard in Dramatic Rescue". 4 August 2016. Archived from the original on 5 August 2016. Retrieved 12 August 2018.
  48. ^ "Leopard rescued from well in eastern India". USA TODAY. Archived from the original on 12 August 2018. Retrieved 12 August 2018.
  49. ^ "Leopard rescued from well, released". The Hindu. 11 August 2018. ISSN 0971-751X. Archived from the original on 25 April 2020. Retrieved 12 August 2018.
  50. ^ a b c d e f Banks, D., Lawson, S., Wright, B. (eds.) (2006). Skinning the Cat: Crime and Politics of the Big Cat Skin Trade Archived 2 January 2021 at the Wayback Machine. Environmental Investigation Agency, Wildlife Protection Society of India
  51. ^ a b c d Banks, D. (2004). The Tiger Skin Trail. Environmental Investigation Agency.
  52. ^ a b c Raza, R.H.; Chauhan, D.S.; Pasha, M.K.S. & Sinha, S. (2012). Illuminating the blind spot: A study on illegal trade in Leopard parts in India (2001–2010) (PDF) (Report). New Delhi: TRAFFIC India, WWF India. Archived (PDF) from the original on 24 September 2020. Retrieved 9 August 2020.
  53. ^ a b c d e Wildlife Protection Society of India (2010). Leopards Battling For Survival In India Archived 22 November 2022 at the Wayback Machine. Wildlife Protection Society of India, 18 May 2010.
  54. ^ Wildlife Trust of India (2008). Leopard skin traders arrested in UP; eight skins recovered. Wildlife Trust of India, 29 July.
  55. ^ Ghosh, A. (2008). 27 leopard skins seized in 45 days Archived 26 March 2019 at the Wayback Machine. Wildlife Protection Society of India.
  56. ^ The Hindu (2008). Leopard skin, other wildlife products seized; five held Archived 26 March 2019 at the Wayback Machine.
  57. ^ Wildlife Protection Society of India (2009). Leopard Skins Seized in Dehradun Archived 26 March 2019 at the Wayback Machine, 18 March 2009.
  58. ^ The Indian Express Limited (2010). 4 with leopard hide held, role of politician to be probed, 12 October 2010.
  59. ^ a b c Aryal, R. S. (2009). CITES : Implementation in Nepal and India, Law, Policy and Practice (PDF) (2nd ed.). Kathmandu: Bhrikuti Academic Publications. ISBN 978-99933-673-3-8.[permanent dead link]
  60. ^ Yonzon, P. (2008). Conservation of Tigers in Nepal 2007. Wildlife Conservation Nepal
  61. ^ Wildlife Trust of India (2008). Cross-border wildlife traders arrested in Nepal with WTI’s help. Wildlife Trust of India, 12 May.
  62. ^ Sears, S. (2008). "Mumbai Leopards: Killers or Victims?" Archived 16 July 2011 at the Wayback Machine Wildlife Extra, 11 April 2008.
  63. ^ Sears, S. (2009). "The wild leopards of Oman and Nepal – And how to see them" Archived 17 October 2011 at the Wayback Machine. Wildlife Extra, April 2009.
  64. ^ Athreya, V., Belsare, A. (2007). Human – Leopard Conflict Management Guidelines. Kaati Trust, Pune, India.
  65. ^ Dollar, L. (2016). "Leopards of India's Silicon City". National Geographic (blogs). Archived from the original on 4 January 2016. Retrieved 7 February 2016.
  66. ^ "Almost half of Karnataka 'lives' with leopards". The Hindu. 2015. ISSN 0971-751X. Archived from the original on 27 April 2020. Retrieved 15 February 2016.
  67. ^ "Nature Conservation Foundation - The secret lives of leopards". ncf-india.org. Archived from the original on 29 April 2016. Retrieved 15 February 2016.
  68. ^ "Leopard Spotted Inside Bengaluru School". The New Indian Express. Archived from the original on 8 February 2016. Retrieved 7 February 2016.
  69. ^ Kumar, P.; Chandel, S.; Kumar, V.; Sankhyan, V. (2017). "Leopard–Human Conflict Led Casualties and Conservation Awareness Campaign in Shivalik Hills of Northern India". Proceedings of the National Academy of Sciences, India Section B: Biological Sciences. 87 (3): 893−898. doi:10.1007/s40011-015-0653-3. S2CID 25553974.
  70. ^ Athreya, V. (2012). Conflict resolution and leopard conservation in a human dominated landscape (PhD). Manipal University. hdl:10603/5431. Archived from the original on 15 November 2022. Retrieved 29 March 2013.
  71. ^ a b Maskey, T. M.; Bauer, J.; Cosgriff, K. (2001). Village children, leopards and conservation. Patterns of loss of human live through leopards (Panthera pardus) in Nepal (Report). Kathmandu, Nepal: Department of National Parks and Wildlife Conservation/Sustainable Tourism CRC.
  72. ^ a b Quigley, H.; Herrero, S. (2005). "Chapter 3: Characterization and prevention of attacks on humans". In Woodroffe, R.; Thirgood, S.; Rabinowitz, A. (eds.). People and wildlife: Conflict or co-existence?. Cambridge University Press. pp. 27–48. ISBN 9780521825054.
  73. ^ Inskip, C.; Zimmermann, A. (2009). "Human-felid conflict: A review of patterns and priorities worldwide". Oryx. 43 (1): 18–34. doi:10.1017/S003060530899030X.
  74. ^ Quammen, D. (2003). Monster of God: The Man-Eating Predator in the Jungles of History and the Mind. New York: W. W. Norton & Company. pp. 55–61. ISBN 9780393326093. Retrieved 20 March 2013.
  75. ^ Kimothi, P. (2011). "Losers on both sides as man-animal war rages". The Pioneer. Archived from the original on 2 March 2011. Retrieved 22 March 2013.
  76. ^ Athreya, V. R.; Thakur, S. S.; Chaudhuri, S.; Belsare, A. V. (2004). A study of the man-leopard conflict in the Junnar Forest Division, Pune District, Maharashtra (PDF) (Report). Submitted to the Office of the Chief Wildlife Warden, Maharashtra State Forest Department, and the Wildlife Protection Society of India, New Delhi, India. Archived (PDF) from the original on 5 February 2020. Retrieved 26 August 2019.
  77. ^ Corbett, E. J. (1954). The temple tiger and more man-eaters of Kumaon. London: Oxford University Press. pp. 64–86. OCLC 1862625. Retrieved 29 March 2013.
  78. ^ Spotted in India: Humans and leopards living in harmony Archived 11 July 2021 at the Wayback Machine, ourbetterworld.org, 9 Apr 2021.
  79. ^ Zoological Survey of India (1994). The Red Data Book on Indian Animals (PDF). Vol. Part. 1: Vertebrata (Mammalia, Aves, Reptilia and Amphibia) . Calcutta: White Lotus Press. Archived (PDF) from the original on 6 December 2022. Retrieved 11 October 2022.
  80. ^ Bhutan Biodiversity User List (PDF). National Environment Commission, Royal Government of Bhutan. 2021. ISBN 9789998046023. Archived (PDF) from the original on 12 October 2022. Retrieved 12 October 2022.
  81. ^ Jnawali, S.R.; Baral, H.S.; Lee, S.; Acharya, K.P.; Upadhyay, G.P.; Pandey, M.; Shrestha, R.; Joshi, D.; Laminchhane, B.R.; Griffiths, J.; Khatiwada, A.P.; Subedi, N. & Amin, R. (2011). "Panthera pardus (Schlegel, 1857)". The Status of Nepal Mammals (PDF). The National Red List Series. Kathmandu, Nepal: Department of National Parks and Wildlife Conservation. pp. 84–85. ISBN 9780900881602. Archived (PDF) from the original on 21 October 2022. Retrieved 12 October 2022.
  82. ^ Sheikh, K. M.; Molur, S., eds. (2004). "Panthera pardus (Linnaeus, 1758). Panther or Leopard". Status and Red List of Pakistan's Mammals. Based on the Conservation Assessment and Management Plan (PDF). Islamabad: IUCN Pakistan. p. 59. Archived (PDF) from the original on 9 October 2022. Retrieved 12 October 2022.
  83. ^ Champion, F. W. (1934). "What is the Use of Leopards?". The Jungle in Sunlight and Shadow (Reprinted 1996 ed.). Dehra Dun: Natraj Publishers. pp. 71–80. ISBN 81-85019-53-3.
  84. ^ Singh, A. (1982). Prince of Cats. London: Jonathan Cape. ISBN 978-0195654028.
  85. ^ "Manikdoh Leopard rescue centre to get facelift". dna. 2013. Archived from the original on 22 November 2022. Retrieved 12 August 2018.
  86. ^ "Kanpur zoo to propose leopard rescue centre - Times of India". The Times of India. Archived from the original on 23 November 2022. Retrieved 12 August 2018.
  87. ^ "Protected parks or change in human behaviour: what will save the threatened leopards of Gujarat?". india.mongabay.com. 2018. Archived from the original on 12 August 2018. Retrieved 12 August 2018.
  88. ^ "Ben Kingsley to Voice Bagheera in Disney's The Jungle Book". Deadline. 25 June 2014. Archived from the original on 29 June 2014. Retrieved 25 June 2014.
  89. ^ Man-Eaters of Kumaon (1944), Jim Corbett, Oxford University Press, Bombay.
  90. ^ "Marathi film 'Ajoba' based on a leopard's fascinating journey, Urmila plays a wildlife biologist". 13 June 2013. Archived from the original on 26 March 2019. Retrieved 26 August 2017.
  91. ^ Raman, Giji K. (13 June 2021). "Tale of a mystery cat echoes on Anamudi hills". The Hindu. Archived from the original on 31 July 2021. Retrieved 26 February 2022.

Further reading

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