List of Chthoniidae species

This is a list of the described species of the pseudoscorpion family Chthoniidae. The data is taken from Joel Hallan's Biology Catalog.

Afrochthonius Beier, 1930

Afrochthonius brincki Beier, 1955 — Lesotho
Afrochthonius ceylonicus Beier, 1973 — Sri Lanka
Afrochthonius godfreyi (Ellingsen, 1912) — Southern Africa
Afrochthonius inaequalis Beier, 1958 — Southern Africa
Afrochthonius natalensis Beier, 1931 — Southern Africa
Afrochthonius reductus Beier, 1973 — Sri Lanka
Afrochthonius similis Beier, 1930 — Namibia

Allochthonius J.C. Chamberlin, 1929

Allochthonius borealis Sato, 1984 — Japan
Allochthonius buanensis W.K.Lee, 1982 — Korea
Allochthonius kinkaiensis Sakayori, 2002 — Japan
Allochthonius montanus Sakayori, 2000 — Honshu, Tochigi, Mount Kohshin, Japan
Allochthonius opticus (Ellingsen, 1907) — Japan

Allochthonius opticus opticus (Ellingsen, 1907) — Japan
Allochthonius opticus coreanus Morikawa, 1970 — Korea
Allochthonius opticus troglophilus Morikawa, 1956 — Japan

Allochthonius shintoisticus J.C. Chamberlin, 1929 — Japan
Allochthonius tamurai Sakayori, 1999 — Japan
Allochthonius biocularis (Morikawa, 1956) — Japan
Allochthonius ishikawai (Morikawa, 1954) — Japan

Allochthonius ishikawai ishikawai (Morikawa, 1954) — Japan
Allochthonius ishikawai deciclavatus (Morikawa, 1956) — Japan
Allochthonius ishikawai kyushuensis (Morikawa, 1960) — Japan
Allochthonius ishikawai shiragatakiensis (Morikawa, 1954) — Japan
Allochthonius ishikawai uenoi (Morikawa, 1956) — Japan

Allochthonius uyamadensis (Morikawa, 1954) — Japan

Aphrastochthonius J.C. Chamberlin, 1962

Aphrastochthonius alteriae Muchmore, 1977 — Mexico
Aphrastochthonius cubanus Dumitresco & Orghidan, 1977 — Cuba
Aphrastochthonius grubbsi Muchmore, 1984 — California
Aphrastochthonius major Muchmore, 1973 — Mexico
Aphrastochthonius pachysetus Muchmore, 1976 — New Mexico
Aphrastochthonius palmitensis Muchmore, 1986 — Mexico
Aphrastochthonius parvus Muchmore, 1972 — Mexico
Aphrastochthonius patei Muchmore, 1982 — Mexico
Aphrastochthonius pecki Muchmore, 1968 — Alabama
Aphrastochthonius russelli Muchmore, 1972 — Mexico
Aphrastochthonius similis Muchmore, 1984 — California
Aphrastochthonius tenax J.C. Chamberlin, 1962
Aphrastochthonius verapazanus Muchmore, 1972 — Guatemala

Apochthonius J.C. Chamberlin, 1929

Apochthonius coecus (Packard, 1884) — Virginia
Apochthonius colecampi Muchmore, 1967 — Missouri
Apochthonius diabolus Muchmore, 1967 — Arkansas
Apochthonius forbesi Benedict, 1979 — Oregon
Apochthonius grubbsi Muchmore, 1980 — California
Apochthonius hobbsi Muchmore, 1994 — Ohio
Apochthonius holsingeri Muchmore, 1967 — Virginia
Apochthonius hypogeus Muchmore, 1976 — Virginia
Apochthonius indianensis Muchmore, 1967 — Indiana
Apochthonius intermedius J.C. Chamberlin, 1929 — Washington
Apochthonius irwini R.O. Schuster, 1966 — California
Apochthonius knowltoni Muchmore, 1980 — Wyoming
Apochthonius magnanimus Hoff, 1956 — New Mexico
Apochthonius malheuri Benedict & Malcolm, 1973 — Oregon
Apochthonius maximus R.O. Schuster, 1966 — California
Apochthonius minimus R.O. Schuster, 1966 — British Columbia, Oregon, Washington
Apochthonius minor Muchmore, 1976 — Georgia
Apochthonius moestus (Banks, 1891) — Eastern U.S.
Apochthonius mysterius Muchmore, 1976 — Missouri
Apochthonius occidentalis J.C. Chamberlin, 1929 — Oregon
Apochthonius paucispinosus Muchmore, 1967 — West Virginia
Apochthonius russelli Muchmore, 1976 — Alabama
Apochthonius titanicus Muchmore, 1976 — Arkansas
Apochthonius typhlus Muchmore, 1967 — Missouri

Austrochthonius J.C. Chamberlin, 1929

Austrochthonius argentinae Hoff, 1950 — Argentina, Brazil
Austrochthonius australis Hoff, 1951 — Australia
Austrochthonius bolivianus Beier, 1930 — Bolivia
Austrochthonius cavicola Beier, 1968 — Australia
Austrochthonius chilensis (J.C. Chamberlin, 1923)

Austrochthonius chilensis chilensis (J.C. Chamberlin, 1923) — Chile, Argentina
Austrochthonius chilensis magalhanicus Beier, 1964 — Chile
Austrochthonius chilensis transversus Beier, 1964 — Argentina

Austrochthonius easti Harvey, 1991 — western Australia
Austrochthonius iguazuensis Vitali-di Castri, 1975 — Brazil
Austrochthonius insularis Vitali-di Castri, 1968 — Crozet Islands
Austrochthonius mordax Beier, 1967 — New Zealand
Austrochthonius paraguayensis Vitali-di Castri, 1975 — Paraguay
Austrochthonius parvus (Mello-Leitão, 1939) — Argentina
Austrochthonius persimilis Beier, 1930 — Chile
Austrochthonius rapax Beier, 1976 — New Zealand
Austrochthonius semiserratus Beier, 1930 — Chile
Austrochthonius tullgerni (Beier, 1931) — Southern Africa
Austrochthonius zealandicus Beier, 1966 — New Zealand

Austrochthonius zealandicus zealandicus Beier, 1966 — New Zealand
Austrochthonius zealandicus obscurus Beier, 1966 — New Zealand

Caribchthonius Muchmore, 1976

Caribchthonius butleri Muchmore, 1976 — Virgin Islands
Caribchthonius orthodentatus Muchmore, 1976 — Belize

Centrochthonius Beier, 1931

Centrochthonius koslovi (Redikorzev, 1918) — Tibet, India, Nepal
Centrochthonius schnitnikovi (Redikorzev, 1934) — Russia
Centrochthonius sichuanensis Schawaller, 1995 — Sichuan

Chiliochthonius Vitali-di Castri, 1975

Chiliochthonius centralis Vitali-di Castri, 1975 — Chile
Chiliochthonius montanus Vitali-di Castri, 1975 — Chile
Chiliochthonius usuriensis Beier, 1979 — Russia (?)

Chthonius C.L. Koch, 1843

Chthonius absoloni Beier, 1938 — Yugoslavia
Chthonius agazzii Beier, 1966 — Italy
Chthonius alpicola Beier, 1951 — Austria, Italy
Chthonius apollinis Mahnert, 1978 — Greece
Chthonius azerbaidzhanus Schawaller & Dashdamirov, 1988 — Azerbaijan
Chthonius baccettii Callaini, 1980 — Italy
Chthonius balazuci Vachon, 1963 — France
Chthonius bogovinae Curcic, 1972 — Yugoslavia

Chthonius bogovinae bogovinae Curcic, 1972 — Yugoslavia
Chthonius bogovinae latidentatus Curcic, 1972 — Yugoslavia

Chthonius brandmayri Callaini, 1986 — Yugoslavia
Chthonius caprai Gardini, 1977 — Italy
Chthonius caoduroi Callaini, 1987 — Italy
Chthonius cavernarum Ellingsen, 1909 — Romania, Yugoslavia
Chthonius cavophilus Hadzi, 1939 — Bulgaria
Chthonius cebenicus Leclerc, 1981 — France
Chthonius cephalotes (Simon, 1875) — France
Chthonius chamberlini (Leclerc, 1983) — Algeria, France
Chthonius comottii Inzaghi, 1987 — Italy
Chthonius cryptus J.C. Chamberlin, 1962 — Greece
Chthonius dacnodes Navás, 1918 — Spain, Azores, Italy
Chthonius dalmatinus Hadzi, 1930 — Yugoslavia
Chthonius densedentatus Beier, 1938 — Italy, Yugoslavia
Chthonius diophthalmus Daday, 1888 — eastern Europe
Chthonius doderoi Beier, 1930 — France, Italy

Chthonius doderoi doderoi Beier, 1930 — France, Italy
Chthonius doderoi horridus Beier, 1934 — Italy

Chthonius ellingseni Beier, 1928 — Italy, Yugoslavia
Chthonius elongatus Lazzeroni, 1970 — Italy
Chthonius euganeus Gardini, 1991 — Italy
Chthonius exarmatus Beier, 1939 — Yugoslavia
Chthonius graecus Beier, 1963 — Greece
Chthonius guglielmii Callaini, 1986 — Italy
Chthonius halberti Kew, 1916 — France, Great Britain
Chthonius herbarii Mahnert, 1980 — Crete
Chthonius heterodactylus Tomosvary, 1882 — central Europe
Chthonius heutaultae Leclerc, 1981 — France
Chthonius hungaricus Mahnert, 1981 — Hungary
Chthonius ilvensis Beier, 1963 — Italy
Chthonius imperator Mahnert, 1978 — Greece
Chthonius ischnocheles (Hermann, 1804) — Europe, U.S.

Chthonius ischnocheles ischnocheles (Hermann, 1804) — Europe, U.S.
Chthonius ischnocheles reductus Beier, 1939 — southern Europe
Chthonius ischnocheles ruffoi Caporiacco, 1951 — Italy
Chthonius ischnocheles stammeri Beier, 1942 — Italy

Chthonius ischnocheloides Beier, 1974 — Italy
Chthonius italicus Beier, 1930 — Italy
Chthonius iugoslavicus Curcic, 1972 — Yugoslavia
Chthonius jalzici Curcic, 1988 — Yugoslavia
Chthonius jonicus Beier, 1931 — southern Europe
Chthonius jugorum Beier, 1952 — Italy
Chthonius karamanianus Hadzi, 1937 — Yugoslavia
Chthonius lanzai Caporiacco, 1947 — Italy

Chthonius lanzai lanzai Caporiacco, 1947 — Italy
Chthonius lanzai vannii Callaini, 1986 — Italy

Chthonius leoi (Callaini, 1988) — Sardinia
Chthonius leruthi Beier, 1939 — Romania
Chthonius lesnik Curcic, 1994 — Yugoslavia
Chthonius lessiniensis Schawaller, 1982 — Italy
Chthonius ligusticus Beier, 1930 — Italy
Chthonius lindbergi Beier, 1956 — Crete
Chthonius litoralis Hadzi, 1933 — Italy, Yugoslavia
Chthonius lucifugus Mahnert, 1977 — France, Spain
Chthonius macedonicus Curcic, 1972 — Yugoslavia
Chthonius magnificus Beier, 1938 — Yugoslavia
Chthonius malatestai Callaini, 1980 — Italy
Chthonius mauritanicus (Callaini, 1988) — Morocco
Chthonius mayi Heurtault-Rossi, 1968 — France
Chthonius mazaurici Leclerc, 1981 — France

Chthonius mazaurici mazaurici Leclerc, 1981 — France
Chthonius mazaurici coironi Leclerc, 1981 — France

† Chthonius mengei Beier, 1937 — fossil: Baltic amber
Chthonius microphthalmus Simon, 1879 — France, Spain
Chthonius mingazzinii Callaini, 1991 — Italy
Chthonius minotaurus Henderickx, 1997 — Kournas Cave, Crete
Chthonius monicae Boghean, 1989 — Romania
Chthonius motasi Dumitresco & Orghidan, 1964 — Romania
Chthonius multidentatus Beier, 1963 — Sicily
Chthonius occultus Beier, 1939 — Yugoslavia
Chthonius orthodactyloides Beier, 1967 — Turkey
Chthonius orthodactylus (Leach, 1817) — Europe
Chthonius pacificus Muchmore, 1968 — California
Chthonius paganus (Hoff, 1961) — Colorado
Chthonius paludis (J.C. Chamberlin, 1929) — southern U.S.
Chthonius paolettii Beier, 1974 — Italy
Chthonius persimilis Beier, 1939 — Yugoslavia
Chthonius petrochilosi Heurtault, 1972 — Greece
Chthonius pivai Gardini, 1991 — Italy
Chthonius ponticoides Mahnert, 1975 — Greece
Chthonius ponticus Beier, 1965 — Turkey
Chthonius porevidi Curcic, Makarov & Lucic, 1998 — Krivosije, Knezlaz Pecina Cave, Montenegro
† Chthonius pristinus Schawaller, 1978 — fossil: Baltic amber
Chthonius prove Curcic, Dimitrijevic & Makarov, 1997 — Gornji Morinj, Serbia, Montenegro
Chthonius pusillus Beier, 1947 — Austria
Chthonius pygmaeus Beier, 1934 — central Europe

Chthonius pygmaeus pygmaeus Beier, 1934 — central Europe
Chthonius pygmaeus carinthiacus Beier, 1951 — central Europe

Chthonius radjai Curcic, 1988 — Yugoslavia
Chthonius raridentatus Hadzi, 1930 — Yugoslavia
Chthonius ressli Beier, 1956 — Austria, France, Italy
Chthonius satapliaensis Schawaller & Dashdamirov, 1988 — Russia
Chthonius sestasi Mahnert, 1980 — Greece
Chthonius shelkovnikovi Redikorzev, 1930 — western Asia
Chthonius shulovi Beier, 1963 — Israel
Chthonius stevanovici Curcic, 1986 — Yugoslavia
Chthonius strinatii Mahnert, 1975 — Greece
Chthonius submontanus Beier, 1963 — central Europe
Chthonius subterraneus Beier, 1931 — Hungary, Yugoslavia

Chthonius subterraneus subterraneus Beier, 1931 — Hungary, Yugoslavia
Chthonius subterraneus meuseli Beier, 1939 — Yugoslavia

Chthonius tadzhikistanicus Dashdamirov & Schawaller, 1992 — Tajikistan
Chthonius tenuis L. Koch, 1873 — Europe, Algeria
Chthonius thessalus Mahnert, 1980 — Greece
Chthonius trebinjensis Beier, 1938 — Yugoslavia
Chthonius troglobius Hadzi, 1937 — Yugoslavia
Chthonius troglodites Redikorzev, 1928 — Bulgaria
Chthonius tzanoudakisi Mahnert, 1975 — Greece
Chthonius aegatensis (Callaini, 1989) — Sicilly
Chthonius aguileraorum Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2000 — Cadiz, Villaluenga del Rosario, Cueva de la Hiedra, Spain
Chthonius amatei Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
Chthonius anatolicus (Beier, 1969) — Iran, Turkey
Chthonius anophthalmus (Ellingsen, 1908) — Algeria
Chthonius apulicus (Beier, 1958) — Italy
Chthonius asturiensis (Beier, 1955) — Spain
Chthonius atlantis (Mahnert, 1980) — Morocco
Chthonius balearicus (Mahnert, 1977) — Balearic Islands
Chthonius bartolii (Gardini, 1976) — Italy
Chthonius bauneensis (Callaini, 1983) — Sardinia
Chthonius beieri (Lazzeroni, 1966) — Italy
Chthonius bellesi (Mahnert, 1989) — Balearic Islands
Chthonius berninii (Callaini, 1983) — Sardinia
Chthonius bidentatus (Beier, 1938) — Yugoslavia
Chthonius boldorii (Beier, 1934) — Italy, Yugoslavia
Chthonius bolivari (Beier, 1930) — Spain
Chthonius cabreriensis Mahnert, 1993 — Balearic Islands
Chthonius cassolai (Beier, 1974) — Sardinia
Chthonius catalonicus (Beier, 1939) — Spain
Chthonius cavicola Gardini, 1990 — Italy
Chthonius chius Schawaller, 1990 — Greece
Chthonius concii (Beier, 1953) — Italy
Chthonius corcyraeus (Mahnert, 1976) — Greece
Chthonius corsicus (Callaini, 1981) — Corsica
Chthonius creticus (Mahnert, 1980) — Crete
Chthonius daedaleus (Mahnert, 1980) — Crete
Chthonius distinguendus (Beier, 1930) — Spain
Chthonius dubius Mahnert, 1993 — Canary Islands
Chthonius elbanus (Beier, 1963) — Italy, Morocco
Chthonius fuscimanus (Simon, 1900) — central Europe, Turkey
Chthonius gasparoi (Gardini, 1989) — Greece
Chthonius genuensis Gardini, 1990 — Italy
Chthonius gestroi (Simon, 1989) — Italy
Chthonius gibbus (Beier, 1953) — Spain, Italy
Chthonius girgentiensis (Mahnert, 1982) — Malta
Chthonius giustii (Callaini, 1981) — Corsica
Chthonius gracilimanus Mahnert, 1997 — La Palma, Mazo, Salto de Tigalate, Canary Islands
Chthonius grafittii (Gardini, 1981) — Sardinia
Chthonius hiberus (Beier, 1930) — Spain
Chthonius hispanus (Beier, 1930) — Spain, Portugal
Chthonius insularis (Beier, 1938) — Yugoslavia
Chthonius iranicus (Beier, 1971) — Iran
Chthonius kabylicus (Callaini, 1983) — Algeria
Chthonius kemza Curcic, Lee & Makarov, 1993 — Yugoslavia
Chthonius kewi (Gabbutt, 1966) — Great Britain
Chthonius longesetosus (Mahnert, 1976) — Morocco
Chthonius lucanus (Callaini, 1984) — Italy
Chthonius machadoi (Vachon, 1940) — Spain, Morocco

Chthonius machadoi machadoi (Vachon, 1940) — Spain, Morocco
Chthonius machadoi canariensis (Beier, 1965) — Canary Islands

Chthonius mahnerti (Zaragoza, 1984) — Spain
Chthonius maltensis (Mahnert, 1975) — Malta
Chthonius mariolae Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
Chthonius maroccanus (Mahnert, 1980) — Morocco
Chthonius mayorali Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
Chthonius microtuberculatus (Hadzi, 1937) — Bulgaria, Yugoslavia
Chthonius minous (Mahnert, 1980) — Crete

Chthonius minous minous (Mahnert, 1980) — Crete
Chthonius minous peramae (Mahnert, 1980) — Crete

Chthonius minutus (Vachon, 1940) — Portugal
Chthonius nanus (Beier, 1953) — Italy
Chthonius nerjaensis Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
Chthonius nidicola (Mahnert, 1979) — Switzerland
Chthonius parmensis (Beier, 1963) — Italy
Chthonius pieltaini (Beier, 1930) — Italy
Chthonius pinai (Zaragoza, 1985) — Spain
Chthonius platakisi (Mahnert, 1980) — Crete
Chthonius poeninus (Mahnert, 1979) — Switzerland
Chthonius ponsi Mahnert, 1993 — Balearic Islands
Chthonius poseidonis Gardini, 1993 — Sardinia
Chthonius pyrenaicus (Beier, 1934) — France, Spain
Chthonius remyi (Heurtault, 1975) — Corsica
Chthonius rimicola Mahnert, 1993 — Canary Islands
Chthonius romanicus (Beier, 1935) — Greece, Iran
Chthonius ruizporteroi Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain
Chthonius sacer (Beier, 1963) — Israel
Chthonius samius (Mahnert, 1982) — Greece
Chthonius schmalfussi Schawaller, 1990 — Greece
Chthonius scythicus Georgescu & Capuse, 1994 — Romania
Chthonius sendrai (Zaragoza, 1985) — Spain
Chthonius serbicus (Hadzi, 1937) — Bulgaria, Yugoslavia
Chthonius setosus Mahnert, 1993 — Canary Islands
Chthonius siculus (Beier, 1961) — Italy, Spain
Chthonius siscoensis (Heurtault, 1975) — Corsica
Chthonius tetrachelatus (Preyssler, 1790) — Cosmopolitan
Chthonius troglophilus (Beier, 1920) — Italy
Chthonius tuberculatus (Hadzi, 1937) — central Europe, Greece
Chthonius vachoni (Heurtault-Rossi, 1963) — France
Chthonius ventalloi (Beier, 1939) — Spain

Chthonius ventalloi ventalloi (Beier, 1939) — Spain
Chthonius ventalloi cazorlensis (Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001) — Spain

Chthonius verai (Zaragoza, 1987) — Spain
Chthonius virginicus (J.C. Chamberlin, 1929) — Eastern U.S.
Chthonius zoiai Gardini, 1990 — Italy
Chthonius abnormis (Beier, 1939) — Yugoslavia
Chthonius caligatus (Beier, 1938) — Yugoslavia
Chthonius cavernicola (Beier, 1938) — Yugoslavia
Chthonius cerberus (Beier, 1938) — Yugoslavia
Chthonius globifer (Simon, 1879) — France, Switzerland
Chthonius pancici (Curbic, 1972) — Yugoslavia
Chthonius polychaetus (Hadzi, 1937) — Yugoslavia
Chthonius purgo Curcic, Lee & Makarov, 1993 — Yugoslavia
Chthonius simplex (Beier, 1939) — Yugoslavia
Chthonius spelaeophilus (Hadzi, 1930) — Yugoslavia

Chthonius spelaeophilus spelaeophilus (Hadzi, 1930) — Yugoslavia
Chthonius spelaeophilus histricus (Beier, 1931) — Yugoslavia, Italy

Chthonius vandeli (Dumitresco & Orghidan, 1964) — Romania
Chthonius californicus (J.C. Chamberlin, 1929) — California
Chthonius oregonicus (Muchmore, 1968) — Oregon
Chthonius spingolus (R.O. Schuster, 1962) — California
Chthonius karamani (Hadzi, 1933) — Yugoslavia
Chthonius rogatus (Beier, 1938) — Yugoslavia

Congochthonius Beier, 1959

Congochthonius nanus Beier, 1959 — Democratic Republic of Congo

Drepanochthonius Beier, 1964

Drepanochthonius horridus Beier, 1964 — Chile

Francochthonius Vitali-di Castri, 1975

Francochthonius Vitali-di Castri, 1975 — Chile

Kleptochthonius J.C. Chamberlin, 1949

Kleptochthonius crosbyi (J.C. Chamberlin, 1929) — Kentucky, North Carolina
Kleptochthonius geophilus Malcolm & J.C. Chamberlin, 1961 — Oregon
Kleptochthonius griseomanus Muchmore, 2000 — Hoosier National Forest, Indian Cave, Indiana
Kleptochthonius inusitatus Muchmore, 1994 — Ohio
Kleptochthonius lewisorum Muchmore, 2000 — Harrison County, Baelz Cave, Indiana
Kleptochthonius magnus Muchmore, 1966 — Tennessee
Kleptochthonius multispinosus (Hoff, 1945) — Eastern U.S.
Kleptochthonius oregonus Malcolm & J.C. Chamberlin, 1961 — Oregon
Kleptochthonius polychaetus Muchmore, 1994 — Virginia
Kleptochthonius sheari Muchmore, 1994 — West Virginia
Kleptochthonius affinis (Muchmore, 1976) — Tennessee
Kleptochthonius anophthalmus (Muchmore, 1970) — Virginia
Kleptochthonius attenuatus (Malcolm & J.C. Chamberlin, 1961) — Kentucky
Kleptochthonius barri (Muchmore, 1965) — Tennessee
Kleptochthonius binoculatus (Muchmore, 1974) — Virginia
Kleptochthonius cerberus (Malcolm & J.C. Chamberlin, 1961) — Kentucky
Kleptochthonius charon (Muchmore, 1965) — Tennessee
Kleptochthonius daemonius (Muchmore, 1965) — Tennessee
Kleptochthonius erebicus (Muchmore, 1965) — Kentucky
Kleptochthonius gertschi (Malcolm & J.C. Chamberlin, 1961) — Virginia
Kleptochthonius hageni (Muchmore, 1963) — Kentucky
Kleptochthonius henroti (Vachon, 1952) West Virginia
Kleptochthonius hetricki (Muchmore, 1974) — West Virginia
Kleptochthonius hubrichti (Muchmore, 1965) — Kentucky
Kleptochthonius infernalis (Malcolm & J.C. Chamberlin, 1961) — Tennessee
Kleptochthonius krekeleri (Muchmore, 1965) — Kentucky
Kleptochthonius lutzi (Malcolm & J.C. Chamberlin, 1961); — Tennessee
Kleptochthonius microphthalmus (Malcolm & J.C. Chamberlin, 1961) — Kentucky
Kleptochthonius myoplus (Malcolm & J.C. Chamberlin, 1961) — Tennessee
Kleptochthonius orpheus (Muchmore, 1965) — West Virginia
Kleptochthonius packardi (Hagen, 1879)
Kleptochthonius pluto (Muchmore, 1965) — Tennessee
Kleptochthonius proserpinae (Muchmore, 1965) — West Virginia
Kleptochthonius proximosetus (Muchmore, 1976) — Virginia
Kleptochthonius regulus (Muchmore, 1970) — Virginia
Kleptochthonius rex (Malcolm & J.C. Chamberlin, 1961) — Tennessee
Kleptochthonius similis (Muchmore, 1976) — Virginia
Kleptochthonius stygius (Muchmore, 1965) — Tennessee
Kleptochthonius tantalus (Muchmore, 1966) — Tennessee

Lagynochthonius Beier, 1951

Lagynochthonius annamensis (Beier, 1951) — Vietnam
Lagynochthonius arctus (Beier, 1967) — New Guinea
Lagynochthonius australicus (Beier, 1966) — Australia
Lagynochthonius bakeri (J.C. Chamberlin, 1929) — Philippines
Lagynochthonius brincki (Beier, 1973) — Sri Lanka
Lagynochthonius curvidigitatus Mahnert, 1997 — Tenerife, Cueva Felipe Reventon, Canary Islands
Lagynochthonius dybasi (Beier, 1957) — Palau
Lagynochthonius exiguus (Beier, 1952) — Malaysia
Lagynochthonius flavus (Mahnert, 1986) — Kenya
Lagynochthonius guasirih (Mahnert, 1988) — Sarawak
Lagynochthonius hamatus Harvey, 1988 — Sumatra
Lagynochthonius himalayensis (Morikawa, 1968) — Nepal
Lagynochthonius hygricus Murthy & Ananthakrishnan, 1977 — India
Lagynochthonius indicus (Murthy & Ananthakrishnan, 1977) — India
Lagynochthonius johni (Redikorzev, 1922) — Indonesia, Philippines
Lagynochthonius kapi Harvey, 1988 — Indonesia
Lagynochthonius kenyensis (Mahnert, 1986) — Kenya
Lagynochthonius mordor Harvey, 1989 — Australia
Lagynochthonius nagaminei (Sato, 1983) — Japan
Lagynochthonius novaeguineae (Beier, 1965) — New Guinea
Lagynochthonius paucedentatus (Beier, 1955) — Malaysia
Lagynochthonius ponapensis (Beier, 1957) — Caroline Islands
Lagynochthonius roeweri (J.C. Chamberlin, 1930) — Java
Lagynochthonius salomonensis (Beier, 1966) — Solomon Islands
Lagynochthonius sinensis (Beier, 1967) — China
Lagynochthonius thorntoni Harvey, 1988 — Java
Lagynochthonius tonkinensis (Beier, 1951) — Vietnam

Malcolmochthonius Benedict, 1978

Malcolmochthonius malcolmi Benedict, 1978 — Oregon
Malcolmochthonius oregonus Benedict, 1978 — Oregon
Malcolmochthonius perplexus Benedict, 1978 — California

Maorichthonius J.C. Chamberlin, 1925

Maorichthonius mortenseni J.C. Chamberlin, 1925 — New Zealand

Mexichthonius Muchmore, 1975

Mexichthonius exoticus Muchmore, 1996 — Texas
Mexichthonius pacal Muchmore, 1978 — Mexico
Mexichthonius unicus Muchmore, 1975 — Mexico

Mundochthonius J.C. Chamberlin, 1929

Mundochthonius alpinus Beier, 1947 — Austria
Mundochthonius basarukini Schawaller, 1989 — Russia
Mundochthonius bifurcatus Kim & Hong, 1994 — South Korea
Mundochthonius carpaticus Rafalski, 1948 — Poland, Ukraine
Mundochthonius cavernicola Muchmore, 1968 — Illinois
Mundochthonius decoui Dumitresco & Orghidan, 1970 — Romania
Mundochthonius dominicanus Muchmore, 1996 — Dominican Republic
Mundochthonius erosidens J.C. Chamberlin, 1929 — California
Mundochthonius holsingeri Benedict & Malcolm, 1974 — Virginia
Mundochthonius japonicus J.C. Chamberlin, 1929 — Japan

Mundochthonius japonicus japonicus J.C. Chamberlin, 1929 — Japan
Mundochthonius japonicus imadatei Morikawa, 1956 — Japan
Mundochthonius japonicus scolytidis Morikawa, 1954 — Japan
Mundochthonius japonicus tripartitus Morikawa, 1956 — Japan

Mundochthonius kiyoshii Sakayori, 2002 — Japan
Mundochthonius magnus J.C. Chamberlin, 1929 — California
Mundochthonius mexicanus Muchmore, 1973 — Mexico
Mundochthonius minusculus Kim & Hong, 1994 — South Korea
Mundochthonius montanus J.C. Chamberlin, 1929 — Colorado, New Mexico
Mundochthonius pacificus (Banks, 1893) — western U.S.
Mundochthonius rossi Hoff, 1949 — central U.S.
Mundochthonius sandersoni Hoff, 1949 — Illinois
Mundochthonius singularis Muchmore, 2001 — Fremont County, Canon City, Fly Cave, Colorado
Mundochthonius styriacus Beier, 1971 — Austria
Mundochthonius ussuricus Beier, 1979 — Russia

Neochthonius J.C. Chamberlin, 1929

Neochthonius amplus (R.O. Schuster, 1962) — California
Neochthonius decoui (Georgescu & Capuse, 1994) — Romania
Neochthonius imperialis Muchmore, 1996 — California
Neochthonius stanfordianus J.C. Chamberlin, 1929 — California
Neochthonius troglodytes Muchmore, 1969 — California

Paraliochthonius Beier, 1956

Paraliochthonius azanius Mahnert, 1986 — Kenya
Paraliochthonius canariensis Vachon, 1961 — Canary Islands
Paraliochthonius carpenteri Muchmore, 1984 — Bahamas
Paraliochthonius hoestlandti Vachon, 1960 — Madeira Islands

Paraliochthonius hoestlandti hoestlandti Vachon, 1960 — Madeira Islands

Paraliochthonius insulae Hoff, 1963 — Jamaica
Paraliochthonius johnstoni (J.C. Chamberlin, 1923) — Mexico
Paraliochthonius martini Mahnert, 1989 — Canary Islands
Paraliochthonius mirus Mahnert, 2002 — Canary Islands
Paraliochthonius puertoricensis Muchmore, 1967 — Puerto Rico
Paraliochthonius singularis (Menozzi, 1924) — Europe
Paraliochthonius takashimai (Morikawa, 1958) — Japan
Paraliochthonius tenebrarum Mahnert, 1989 — Canary Islands
Paraliochthonius weygoldti Muchmore, 1967 — Florida

Pseudochthonius Balzan, 1892

Pseudochthonius arubensis Wagenaar-Hummelinck, 1948 — Aruba
Pseudochthonius beieri Mahnert, 1978 — Republic of Congo
Pseudochthonius billae Vachon, 1941 — Ivory Coast
Pseudochthonius biseriatus Mahnert, 2001 — Minas Gerais, Itacarambi, Gruta Olhos dAgua, Brazil
Pseudochthonius brasiliensis Beier, 1970 — Brazil
Pseudochthonius clarus Hoff, 1963 — Jamaica
Pseudochthonius congicus Beier, 1959 — Democratic Republic of Congo
Pseudochthonius doctus Hoff, 1963 — Jamaica
Pseudochthonius falcatus Muchmore, 1977 — Belize
Pseudochthonius galapagensis Beier, 1977 — Galapagos
Pseudochthonius gracilmanus Mahnert, 2001 — Bahia, Gruta Azul, Iraquara, Brazil
Pseudochthonius heterodentatus Hoff, 1946 — Trinidad
Pseudochthonius homodentatus J.C. Chamberlin, 1929 — Venezuela
Pseudochthonius insularis J.C. Chamberlin, 1929 — St. Vincent
Pseudochthonius leleupi Beier, 1959 — Democratic Republic of Congo
Pseudochthonius moralesi Muchmore, 1977 — Mexico
Pseudochthonius mundanus Hoff, 1963 — Jamaica
Pseudochthonius naranjitensis (Ellingsen, 1902) — Ecuador
Pseudochthonius orthodactylus Muchmore, 1970 — Brazil
Pseudochthonius perreti Mahnert, 1986 — Kenya
Pseudochthonius pulchellus (Ellingsen, 1902) — Ecuador
Pseudochthonius ricardoi Mahnert, 2001 — Sao Paulo, Vale do Betari, Caverna Agua Suja, Brazil
Pseudochthonius simoni (Balzan, 1892) — Venezuela
† Pseudochthonius squamosus Schawaller, 1980 — fossil: Dominican amber
Pseudochthonius strinatii Beier, 1969 — Brazil
Pseudochthonius thibaudi Vitali-di Castro, 1984 — Guadeloupe
Pseudochthonius troglobius Muchmore, 1986 — Mexico
Pseudochthonius tuxeni Mahnert, 1979 — Brazil
Pseudochthonius yucatanus Muchmore, 1977 — Mexico

Pseudotyrannochthonius Beier, 1930

Pseudotyrannochthonius australiensis Beier, 1966 — Australia
Pseudotyrannochthonius bornemisszai Beier, 1966 — Australia
Pseudotyrannochthonius dentifer (Morikawa, 1970) — Korea
Pseudotyrannochthonius giganteus Beier, 1971 — Australia
Pseudotyrannochthonius gigas Beier, 1979 — Australia
Pseudotyrannochthonius gracilis Benedict & Malcolm, 1970 — California, Washington
Pseudotyrannochthonius hamiltonsmithi Beier, 1968 — Australia
Pseudotyrannochthonius incognitus (R.O. Schuster, 1966) — western U.S.
Pseudotyrannochthonius jonesi (J.C. Chamberlin, 1962) — Australia
Pseudotyrannochthonius kobayashii (Morikawa, 1956) — Japan

Pseudotyrannochthonius kobayashii kobayashii (Morikawa, 1956) — Japan
Pseudotyrannochthonius kobayashii akiyoshiensis (Morikawa, 1956) — Japan
Pseudotyrannochthonius kobayashii dorogawaensis (Morikawa, 1960) — Japan

Pseudotyrannochthonius kubotai (Morkawa, 1954) — Japan
Pseudotyrannochthonius octospinosus Beier, 1930 — Chile
Pseudotyrannochthonius queenslandicus Beier, 1969 — Australia
Pseudotyrannochthonius rossi Beier, 1964 — Chile
Pseudotyrannochthonius silvestrii (Ellingsen, 1905) — Chile
Pseudotyrannochthonius solitarius (Hoff, 1951) — Australia
Pseudotyrannochthonius tasmanicus Dartnall, 1970 — Tasmania
Pseudotyrannochthonius typhlus Dartnall, 1970 — Tasmania
Pseudotyrannochthonius undecimclavatus (Morikawa, 1956) — Japan

Pseudotyrannochthonius undecimclavatus undecimclavatus (Morikawa, 1956) — Japan
Pseudotyrannochthonius undecimclavatus kishidai (Morikawa, 1960) — Japan

Pseudotyrannochthonius utahensis Muchmore, 1967 — Utah

Sathrochthoniella Beier, 1967

Sathrochthoniella zealandica Beier, 1967 — New Zealand

Sathrochthonius J.C. Chamberlin, 1962

Sathrochthonius crassidens Beier, 1966 — Australia
Sathrochthonius insulanus Beier, 1976 — Lord Howe Island
Sathrochthonius kaltenbachi Beier, 1966 — New Caledonia
Sathrochthonius maoricus Beier, 1976 — New Zealand
Sathrochthonius pefauri Vitali-di Castro, 1974 — Chile
Sathrochthonius tuena J. C. Chamberlin, 1962 — Australia
Sathrochthonius tullgreni J. C. Chamberlin, 1962 — Australia
Sathrochthonius venezuelanus Muchmore, 1989 — Venezuela
Sathrochthonius webbi Muchmore, 1982 — Australia

Selachochthonius J. C. Chamberlin, 1929

Selachochthonius cavernicola Lawrence, 1935 — Southern Africa
Selachochthonius heterodentata Beier, 1955 — Southern Africa
Selachochthonius serratidentaus (Ellingsen, 1912) — Southern Africa

Spelyngochthonius Beier, 1955

Spelyngochthonius beieri Gardini, 1994 — Sardinia
Spelyngochthonius grafittii Gardini, 1994 — Sardinia
Spelyngochthonius heurtaultae Vachon, 1967 — Spain, Sardinia
Spelyngochthonius provincialis Vachon & Heurtault-Rossi, 1964 — France
Spelyngochthonius sardous Beier, 1955 — Sardinia

Stygiochthonius Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001

Stygiochthonius barrancoi Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain

Troglochthonius Beier, 1939

Troglochthonius doratodactylus Helversen, 1968 — Italy
Troglochthonius mirabilis Beier, 1939 — Yugoslavia

Tyrannochthoniella Beier, 1966

Tyrannochthoniella zealandica Beier, 1966 — New Zealand

Tyrannochthoniella zealandica zealandica Beier, 1966 — New Zealand
Tyrannochthoniella zealandica foveauxana Beier, 1967 — New Zealand

Tyrannochthonius J.C. Chamberlin, 1929

Tyrannochthonius alabamensis Muchmore, 1996 — Alabama
Tyrannochthonius aladdinensis Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius albidus (Beier, 1977) — Galapagos
Tyrannochthonius amazonicus Mahnert, 1979 — Brazil
Tyrannochthonius aralu Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius archeri Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius assimilis Hong & Tae Heung Ki, 1993 — South Korea
Tyrannochthonius attenuatus Muchmore, 1996 — Alabama
Tyrannochthonius avernicolus Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius bagus Harvey, 1988 — Sumatra
Tyrannochthonius bahamensis Muchmore, 1984 — Bahamas
Tyrannochthonius barri Muchmore, 1996 — Alabama
Tyrannochthonius beieri Morikawa, 1963 — New Guinea, Solomons
Tyrannochthonius binoculatus Muchmore, 1996 — Alabama
Tyrannochthonius bispinosus (Beier, 1974) — India
Tyrannochthonius brasiliensis Mahnert, 1979 — Brazil
Tyrannochthonius brevimanus Beier, 1935 — central Africa
Tyrannochthonius brooksi Harvey, 1991 — western Australia
Tyrannochthonius butleri Harvey, 1991 — western Australia
Tyrannochthonius caecatus (Beier, 1976) — New Zealand
Tyrannochthonius callidus Hoff, 1959 — Jamaica
Tyrannochthonius cavernicola (Beier, 1976) — Lord Howe Island
Tyrannochthonius cavicola (Beier, 1967) — Australia
Tyrannochthonius centralis Beier, 1931 — Costa Rica, Ecuador
Tyrannochthonius chamarro J.C. Chamberlin, 1947 — Caroline Islands
Tyrannochthonius chamberlini Muchmore, 1996 — Alabama
Tyrannochthonius charon Muchmore, 1996 — Alabama
Tyrannochthonius chelatus Murthy & Ananthakrishnan, 1977 — India
Tyrannochthonius confusus Mahnert, 1986 — Kenya
Tyrannochthonius contractus (Tullgren, 1907) — Africa
Tyrannochthonius convivus Beier, 1974 — India
Tyrannochthonius curazavius Wagenaar-Hummelinck, 1948 — Curacao
Tyrannochthonius densedentatus (Beier, 1967) — New Zealand
Tyrannochthonius diabolus Muchmore, 1996 — Alabama
Tyrannochthonius ecuadoricus (Beier, 1977) — Ecuador, Peru
Tyrannochthonius elegans Beier, 1944 — central Africa
Tyrannochthonius erebicus Muchmore, 1996 — Alabama
Tyrannochthonius felix Muchmore, 1996 — Alabama
Tyrannochthonius ferox Mahnert, 1978 — Republic of Congo
Tyrannochthonius fiskei Muchmore, 1996 — Tennessee
Tyrannochthonius floridensis Malcolm & Muchmore, 1985 — Florida
Tyrannochthonius gezei Vachon, 1941 — Cameroon
Tyrannochthonius gigas Beier, 1954 — Venezuela
Tyrannochthonius gnomus Muchmore, 1996 — Alabama
Tyrannochthonius gomyi Mahnert, 1975 — Reunion
Tyrannochthonius grimmeti J.C. Chamberlin, 1929 — New Zealand
Tyrannochthonius guadeloupensis Vitali-di Castri, 1984 — Guadeloupe
Tyrannochthonius halopotamus Muchmore, 1996 — Alabama
Tyrannochthonius helenae (Beier, 1977) — St. Helena
Tyrannochthonius heterodentatus Beier, 1930 — India
Tyrannochthonius horridus (Beier, 1976) — New Zealand
Tyrannochthonius hypogeus Muchmore, 1996 — Kentucky
Tyrannochthonius imitatus Hoff, 1959 — Jamaica, Dominican Republic
Tyrannochthonius infernalis Muchmore, 1996 — Alabama
Tyrannochthonius innominatus Muchmore, 1996 — Florida
Tyrannochthonius innoxius Hoff, 1959 — Jamaica
Tyrannochthonius insulae Hoff, 1946 — Trinidad
Tyrannochthonius intermedius Muchmore, 1986 — Mexico
Tyrannochthonius irmleri Mahnert, 1979 — Brazil
Tyrannochthonius japonicus (Ellingsen, 1907) — Japan, Taiwan

Tyrannochthonius japonicus japonicus (Ellingsen, 1907) — Japan, Taiwan
Tyrannochthonius japonicus dogoensis Morikawa, 1954 — Japan

Tyrannochthonius jonesi Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius kermadecensis (Beier, 1976) — New Zealand
Tyrannochthonius krakatau Harvey, 1988 — Indonesia
Tyrannochthonius laevis Beier, 1966 — Australia
Tyrannochthonius luxtoni (Beier, 1967) — New Zealand
Tyrannochthonius mahunkai Mahnert, 1978 — Republic of Congo
Tyrannochthonius meneghettii (Caporiacco, 1949) — central Africa
Tyrannochthonius meruensis Beier, 1962 — Eastern Africa
Tyrannochthonius migrans Mahnert, 1979 — Brazil
Tyrannochthonius minor Mahnert, 1979 — Brazil
Tyrannochthonius monodi Vachon, 1941 — Cameroon
Tyrannochthonius nanus (Beier, 1966) — New Guinea
Tyrannochthonius nergal Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius noaensis Moyle, 1989 — New Zealand
Tyrannochthonius norfolkensis (Beier, 1976) — Norfolk Islands
Tyrannochthonius oahuanus Muchmore, 2000 — Hawaii, Oahu, Koolau Range, Puu Konahuanui, Pacific Islands
Tyrannochthonius orpheus Muchmore, 1996 — Alabama
Tyrannochthonius osiris Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius ovatus Vitali-di Castro, 1984 — Martinique
Tyrannochthonius pachythorax Redikorzev, 1938 — Vietnam
Tyrannochthonius palauanus Beier, 1957 — Caroline Islands
Tyrannochthonius pallidus Muchmore, 1973 — Mexico
Tyrannochthonius parvus Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius pecki Muchmore, 1996 — Alabama
Tyrannochthonius perpusillus Beier, 1951 — Vietnam
Tyrannochthonius philippinus (Beier, 1966) — Philippines
Tyrannochthonius pholeter Muchmore, 1996 — Alabama
Tyrannochthonius pluto Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius procerus Mahnert, 1978 — Republic of Congo
Tyrannochthonius proximus Hoff, 1959 — Jamaica, Dominican Republic
Tyrannochthonius pugnax Mahnert, 1978 — Republic of Congo
Tyrannochthonius pupukeanus Muchmore, 1983 — Hawaii
Tyrannochthonius pusillimus Beier, 1951 — Vietnam
Tyrannochthonius pusillus Beier, 1955 — Peru
Tyrannochthonius queenslandicus (Beier, 1969) — Australia
Tyrannochthonius rahmi Beier, 1976 — Nepal, Bhutan
Tyrannochthonius rex Harvey, 1989 — Australia
Tyrannochthonius riberai Mahnert, 1984 — Peru
Tyrannochthonius robustus Beier, 1951 — Vietnam
Tyrannochthonius rotundimanus Mahnert, 1985 — Brazil
Tyrannochthonius satan Muchmore, 1996 — Alabama
Tyrannochthonius semidentatus (Redikorzev, 1924) — Kenya
Tyrannochthonius semihorridus (Beier, 1969) — Australia
Tyrannochthonius setiger Mahnert, 1997 — Tenerife, Cueva del Sobrado, Canary Islands
Tyrannochthonius sheltae Muchmore, 1996 — Alabama
Tyrannochthonius similidentatus Sato, 1984 — Japan
Tyrannochthonius simillimus Beier, 1951 — Cambodia
Tyrannochthonius simulans Mahnert, 1986 — Kenya
Tyrannochthonius skeletonis Muchmore, 1996 — Alabama
Tyrannochthonius sokolovi (Redikorzev, 1924) — central Africa
Tyrannochthonius sparsedentatus Beier, 1959 — Democratic Republic of Congo
Tyrannochthonius spinatus Hong, in Yong Hong, Tae Heung Kim & Won Koo Le 1996 — South Korea
Tyrannochthonius steevesi Muchmore, 1996 — Tennessee
Tyrannochthonius stonei Muchmore, 1989 — Hawaii
Tyrannochthonius strinatii (Beier, 1974) — Guatemala
Tyrannochthonius stygius Muchmore, 1996 — Alabama
Tyrannochthonius superstes Mahnert, 1986 — Canary Islands
Tyrannochthonius suppressalis Hong, in Yong Hong, Tae Heung Kim & Won Koo Le 1996 — South Korea
Tyrannochthonius swiftae Muchmore, 1993 — Hawaii
Tyrannochthonius tartarus Muchmore, 1996 — Alabama
Tyrannochthonius tekauriensis Moyle, 1989 — New Zealand
Tyrannochthonius tenuis Chamberlin, in Muchmore & Chamberlin 1995 — Alabama
Tyrannochthonius terribilis (With, 1906) — southeast Asia

Tyrannochthonius terribilis terribilis (With, 1906) — southeast Asia
Tyrannochthonius terribilis malaccensis Beier, 1952 — Malaysia

Tyrannochthonius tlilapanensis Muchmore, 1986 — Mexico
Tyrannochthonius torodei Muchmore, 1996 — Alabama
Tyrannochthonius troglobius Muchmore, 1969 — Mexico
Tyrannochthonius troglodytes Muchmore, 1986 — Texas
Tyrannochthonius troglophilus (Beier, 1968) — New Caledonia
Tyrannochthonius vampirorum Muchmore, 1986 — Mexico
Tyrannochthonius volcancillo Muchmore, 1986 — Mexico
Tyrannochthonius volcanus Muchmore, 1977 — Mexico
Tyrannochthonius wittei Beier, 1955
Tyrannochthonius wlassicsi (Daday, 1897) — New Guinea
Tyrannochthonius zicsii Mahnert, 1978 — Republic of Congo
Tyrannochthonius zonatus (Beier, 1964) — New Caledonia

Vulcanochthonius Muchmore, 2000

Vulcanochthonius aa Muchmore, 2000 — Hawaii, Manuka Natural Area Reserve, Visor Cave, Pacific Islands
Vulcanochthonius howarthi (Muchmore, 1979) — Hawaii
Vulcanochthonius pohakuloae Muchmore, 2000 — Hawaii, Mauna Kea, Pohakuloa Training Area, Bobcat Trail, Pacific Islands

References

Joel Hallan's Biology Catalog: Chthoniidae