Drymaeus

Drymaeus
	Drymaeus multilineatus hanging on a tree branch.
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Suborder:	Helicina
Superfamily:	Orthalicoidea
Family:	Bulimulidae
Genus:	Drymaeus; Albers, 1850
Type species
	Helix hygrohylaea d'Orbigny, 1835;
Diversity
	more than 750 species names
Synonyms
	Antidrymaeus Germain, 1907; Buliminus (Mesembrinus) Albers, 1850; Bulimulus (Drymaeus) Albers, 1850; Bulimulus (Liostracus); Bulimulus (Mesembrinus) Albers, 1850 (superseded combination); Bulimus (Drymaeus) Albers, 1850 (original rank); Bulimus (Liostracus); Bulimus (Mesembrinus) Albers, 1850 (original combination); Bulimus (Semiclausaria) L. Pfeiffer, 1856 (junior synonym); Diaphanomormus Weyrauch, 1964; Drymaeus (Antidrymaeus); Drymaeus (Drymaeus) Albers, 1850· accepted, alternate representation; Drymaeus (Leptodrymaeus)· accepted, alternate representation; Drymaeus (Mesembrinus) Albers, 1850· accepted, alternate representation; Drymaeus (Metadrymaeus) Pilsbry, 1926· accepted, alternate representation; Drymaeus (Ornatimormus) Weyrauch, 1958· accepted, alternate representation; Drymaeus (Orodrymaeus) Pilsbry, 1926· accepted, alternate representation; Goniognathmus Crosse & P. Fischer, 1875; Hamadryas Albers, 1850; Leptodrymaeus Pilsbry, 1946; Leptomormus Weyrauch, 1958; Mesembrinus Albers, 1850; Mesembrinus (Ornatimormus) Weyrauch, 1958; Mormus E. von Martens, 1860; Otostomus (Drymaeus) Albers, 1850; Semiclausaria L. Pfeiffer, 1856 (junior synonym);

Drymaeus is a large genus of medium-sized air-breathing, tropical land snails, terrestrial pulmonate gastropod molluscs in the subfamily Peltellinae of the family Bulimulidae.^[3]^[4]

Distribution

Distribution of genus Drymaeus include South and Central America. For example in Mexico live about 65 species of Drymaeus.^[2]

Species

There are two subgenera: Drymaeus sensus stricto and subgenus Mesembrinus. They are accepted as alternate representations.

Species within the genus Drymaeus include:

Drymaeus abruptus (Rolle, 1904)
Drymaeus abyssorum (d'Orbigny, 1835)
Drymaeus acervatus Pfeiffer, 1857
Drymaeus acobambensis Weyrauch, 1967
Drymaeus acuminatus Da Costa, 1906
Drymaeus aequatorianus (E.A. Smith, 1877)
Drymaeus aestivus (L. Pfeiffer, 1857)
Drymaeus alabastinus (Scott, 1952)
Drymaeus alabastrinus Da Costa, 1906
Drymaeus albolabiatus (E.A. Smith, 1877)
Drymaeus albostriatus (Strebel, 1882)
Drymaeus alsophilus (Phillipi, 1867)
Drymaeus altenai Breure, 1976
Drymaeus amandus (L. Pfeiffer, 1855)
Drymaeus ambustus (Reeve, 1849)
Drymaeus amoenus (L. Pfeiffer, 1847)
Drymaeus anceps (Albers, 1854)
Drymaeus andai Jousseaume, 1898
Drymaeus angulobasis Pilsbry, 1944
Drymaeus angustus da Costa, 1906
Drymaeus annulatus (Reeve, 1849)
Drymaeus apicepunctatus (Preston, 1914)
Drymaeus arcuatostriatus (L. Pfeiffer, 1855)
Drymaeus albostriatus (Strebel, 1882)^[2]
Drymaeus attenuatus (Pfeiffer, 1851)^[2]
- Drymaeus attenuatus pittieri (Von Martens, 1893)^[2]
- Drymaeus attenuatus varicosus (Pfeiffer, 1851)^[2]
Drymaeus aurantiostomus Thompson & Deisler, 1982
Drymaeus aureolus (Guppy, 1866)
Drymaeus aurifluus (Pfeiffer, 1856)^[2]
Drymaeus auris (L. Pfeiffer, 1866)
Drymaeus aurisratti (Philippi, 1867)
Drymaeus baezensis (Hidalgo, 1869)
Drymaeus beyerleanus (Hupé, 1857)
Drymaeus botterii (Crosse & Fischer, 1875)^[2]
Drymaeus branneri F. Baker, 1914^[5]
Drymaeus bugabensis (Von Martens, 1893)^[2]
Drymaeus castilhensis Simone & Amaral, 2018
Drymaeus castus (Pfeiffer, 1846)^[2]
- Drymaeus castus xantholeucus (Von Martens, 1893)^[2]
- Drymaeus castus porrectus (Von Martens, 1893)^[2]
Drymaeus cecileae (Moricand, 1858)^[5]
Drymaeus championi (Von Martens, 1893)^[2]
Drymaeus chiapensis (Pfeiffer, 1866)^[2]
- Drymaeus chiapensis quadrifasciatus (Von Martens, 1893)^[2]
- Drymaeus chiapensis nebulosus (Von Martens, 1893)^[2]
Drymaeus chiriquiensis DaCosta, 1901^[2]
Drymaeus colimensis (Rolle, 1895)^[2]
Drymaeus costaricensis (Pfeiffer, 1862)^[2]
Drymaeus cozumelensis Richards, 1937^[2]
Drymaeus cucullus (Morelet, 1851)^[2]
Drymaeus dakryodes Salvador, Cavallari & Simone, 2015^[6]
Drymaeus denticulus Breure & Borrerro, 2019
Drymaeus discrepans (Sowerby I, 1833)^[2]
Drymaeus dombeyanus (Férussac, 1842)^[2]
Drymaeus dominicus (Reeve, 1850)^[2]^[7]
Drymaeus droueti (Pfeiffer, 1856)^[2]
- Drymaeus droueti deletus Solem, 1955^[2]
- Drymaeus droueti sporlederi (Pfeiffer, 1866)^[2]
Drymaeus dunkeri (Pfeiffer, 1846)^[2]
- Drymnaeus dunkeri forreri (Mousson, 1883)^[2]
Drymaeus duplexannulus Breure & Borrerro, 2019
Drymaeus elongatus (Röding, 1789)^[4]
Drymaeus emeus (Say, 1829)^[2]
Drymaeus eurystomus (Philippi, 1867)^[5]
Drymaeus expansus (Pfeiffer, 1848)^[4]
- Drymaeus expansus balboa Pilsbry, 1926^[2]
Drymaeus fenestratus (Pfeiffer, 1846)^[2]
Drymaeus fenestrellus (Von Martens, 1863)^[2]
Drymaeus fusoides (d’Orbigny, 1835)
Drymaeus gabbi (Angas, 1879)^[2]
Drymaeus ghiesbreghti (Pfeiffer, 1866)^[2]
- Drymaeus ghiesbreghti interstitialis (Von Martens, 1893)^[2]
- Drymaeus ghiesbreghti iodostylus (Pfeiffer, 1861)^[2]
- Drymaeus ghiesbreghti stolli (Von Martens, 1887)^[2]
Drymaeus hegewischi (Pfeiffer, 1842)^[2]
Drymaeus hepatostomus (Pfeiffer, 1861)^[2]
Drymaeus heterogeneus (Pfeiffer, 1866)^[2]
Drymaeus hondurasanus (Pfeiffer, 1846)^[2]
Drymaeus inconspicuus (Haas, 1949)
Drymaeus iniurius Breure & Borrerro, 2019
Drymaeus inglorius (Reeve, 1848)^[2]
- Drymaeus inglorius heynemanni (Pfeiffer, 1866)^[2]
Drymaeus intermissus Breure & Borrerro, 2019
Drymaeus intrapictus Pilsbry, 1930^[2]
Drymaeus iracema (Simone, 2015)^[8]
Drymaeus irazuensis (Angas, 1878)^[2]
Drymaeus jonasi (Pfeiffer, 1846)^[2]
Drymaeus josephus (Angas, 1878)^[2]
- Drymaeus josephus concolor (Von Martens, 1893)^[2]
- Drymaeus josephus errans Pilsbry, 1926^[2]
- Drymaeus josephus maculosus (Von Martens, 1893)^[2]
Drymaeus joubini Germain, 1907
Drymaeus jousseaumei Dautzenberg, 1901
Drymaeus lattrei (Pfeiffer, 1846)^[2]
- Drymaeus lattrei hiabundus (Von Martens, 1893)^[2]
Drymaeus liliaceus (Férussac, 1821)
Drymaeus lilacinus (Reeve, 1949)^[2]
- Drymaeus lilacinus crossei (Von Martens, 1893)^[2]
- Drymaeus lilacinus ictericus (Von Martens, 1893)^[2]
- Drymaeus lilacinus jansoni (Von Martens, 1893)^[2]
- Drymaeus lilacinus undulosus (Von Martens, 1893)^[2]
- Drymaeus lilacinus unicolor (Von Martens, 1893)^[2]
Drymaeus lineolatus (Conrad, 1855)^[2]
Drymaeus linostoma (d'Orbigny, 1835)
Drymaeus lirinus (Morelet, 1851)^[2]
Drymaeus livescens (Pfeiffer, 1842)^[2]
Drymaeus lophoicus (D'Orbigny, 1835)
Drymaeus marmarinus (D'Orbigny, 1835)
Drymaeus mayaorum Rehder, 1966^[2]
Drymaeus megastomus Parodiz, 1962^[2]
Drymaeus mexicanus (Lamarck, 1822)
Drymaeus micropyrus Simone & Amaral, 2018
Drymaeus moricandi (Pfeiffer, 1846)^[2]
- Drymaeus moricandi hyalinoalbidus (Fischer & Crosse, 1875)^[2]
Drymaeus moritinctus (Von Martens, 1893)^[2]
Drymaeus multifasciatus (Lamarck, 1822)^[9]
Drymaeus multilineatus (Say, 1825)^[2]
Drymaeus necaxanus Solem, 1955^[2]
Drymaeus pamplonensis Pilsbry, 1939^[4]
Drymaeus perductorum Rehder, 1943^[2]
Drymaeus pilsbryi Zetek, 1933^[2]
Drymaeus pluvialis (Pfeiffer, 1862)^[2]
Drymaeus poecilus (d'Orbigny, 1835)
Drymaeus polygramma (S. Moricand, 1836)
Drymaeus ponsonbyi DaCosta, 1907
Drymaeus praetextus (Reeve, 1850)
Drymaeus protractus (L. Pfeiffer, 1855)
Drymaeus pseudelatus Haas, 1951
Drymaeus pseudobesus Breure, 1979
Drymaeus pseudofusoides da Costa, 1906
Drymaeus puellaris (Reeve, 1850)
Drymaeus pulchellus (Broderip, 1832)
Drymaeus pulcherrimus (H. Adams, 1867)
Drymaeus punctatus Da Costa, 1907
Drymaeus puncticulatus (L. Pfeiffer, 1857)
Drymaeus quadrifasciatus (Angas, 1878)
Drymaeus rabuti (Jousseaume, 1898)
Drymaeus rawsoni (Guppy, 1871)
Drymaeus recedens (L. Pfeiffer, 1864)
Drymaeus recluzianus (Pfeiffer, 1847)^[2]
- Drymaeus recluzianus martensianus Pilsbry, 1899^[2]
Drymaeus rudis (Anton, 1839)^[2]
Drymaeus sargi (Crosse & Fischer, 1875)^[2]
- Drymaeus sargi motaguae (Von Martens, 1893)^[2]
Drymaeus schadei Quintana & Magaldi, 1985
Drymaeus schmidti (L. Pfeiffer, 1854)
Drymaeus schunkei Haas, 1949
Drymaeus scitulus (Reeve, 1849)
Drymaeus scoliodes Dautzenberg, 1902
Drymaeus selli (Preston, 1909)
Drymaeus semifasciatus (Mousson, 1869)
Drymaeus semimaculatus Pilsbry, 1898^[2]
Drymaeus semipellucidus (Tristram, 1861)^[2]
Drymaeus serperastrus (Say, 1829)^[2]
Drymaeus serratus (Pfeiffer, 1855)^[4]
Drymaeus shattucki Bequaert & Clench, 1931^[2]
Drymaeus stramineus (Guilding, 1824)^[4]
Drymaeus strigatus (Sowerby, 1838)^[5]
Drymaeus sulcosus (Pfeiffer, 1841)^[2]
Drymaeus suprapunctatus F. Baker, 1913
Drymaeus surinamensis Vernhout, 1914
Drymaeus sykesi Da Costa, 1906
Drymaeus tenuilabris (L. Pfeiffer, 1866)
Drymaeus terreus (Simone, 2015)
Drymaeus totonacus (Strebel, 1882)^[2]
Drymaeus translucens (Broderip, 1832)^[2]
- Drymaeus translucens alternans (Beck, 1837)^[2]
- Drymaeus translucens juquilensis (Von Martens, 1893)^[2]
- Drymaeus translucens misellus Pilsbry, 1926^[2]
- Drymaeus translucens pachecensis Pilsbry, 1930^[2]
- Drymaeus translucens panamensis (Broderip, 1833)^[2]
- Drymaeus translucens sororcula Pilsbry, 1926^[2]
- Drymaeus translucens subfloccosus Pilsbry, 1899^[2]
- Drymaeus translucens tonosiesis Pilsbry, 1930^[2]
Drymaeus trimarianus (Von Martens, 1893)^[2]
Drymaeus tripictus (Albers, 1857)^[2]
- Drymaeus tripictus hoffmanni (Von Martens, 1893)^[2]
Drymaeus tropicalis (Morelet, 1849)^[2]
Drymaeus trujillensis (Philippi, 1867)
Drymaeus tryoni (Fischer & Crosse, 1875)^[2]
- Drymaeus tryoni pochutlensis (Crosse & Fischer, 1875)^[2]
Drymaeus tzubi Dourson, Caldwell & Dourson, 2018
Drymaeus uhdeanus (Von Martens, 1893)^[2]
- Drymaeus uhdeanus borealis (Von Martens, 1893)^[2]
- Drymaeus uhdeanus cuernovacensis (Crosse & Fischer, 1874)^[2]
- Drymaeus uhdeanus tepicensis (Von Martens, 1893)^[2]
Drymaeus verecundus Breure & Mogollón, 2019
Drymaeus vesperus Jousseaume, 1887
Drymaeus vexillum (Broderip, 1832)^[4] (synonym: Bulinus vexillum Broderip, 1832)
Drymaeus vicinus (Preston, 1907)
Drymaeus villavicioensis Breure, 1977
Drymaeus vincentinus (L. Pfeiffer, 1846)
Drymaeus virginalis (L. Pfeiffer, 1857)
Drymaeus virgulatus (Férussac, 1821)
Drymaeus volsus Fulton, 1907
Drymaeus waldoschmidti Parodiz, 1962
Drymaeus weeksi Pilsbry, 1926
Drymaeus yapacanensis Breure, 1981
Drymaeus zhorquinensis (Angas, 1879)^[2]
Drymaeus ziczac (Da Costa, 1898)
Drymaeus ziegleri (Pfeiffer, 1846)^[2]
Drymaeus zilchi Haas, 1955
Drymaeus zingarensis Restrepo & Breure, 1987
Drymaeus zoographica (d'Orbigny, 1835)

Synonyms

Drymaeus henseli is a synonym of Anthinus henselii

Previous species

These species were previously in Drymaeus and moved to Antidrymaeus:^[10]

Antidrymaeus dormani (W. G. Binney, 1857)
Antidrymaeus immaculatus (C. B. Adams, 1850)
Antidrymaeus inusitatus (Fulton, 1900)
Antidrymaeus laticinctus (Guppy, 1868)
Antidrymaeus multifasciatus (Lamarck, 1822)
Antidrymaeus rufescens (J. E. Gray, 1825)
Antidrymaeus stramineus (Guilding, 1824)
Antidrymaeus sulphureus (L. Pfeiffer, 1857)

References

^ (in German) Albers J. C. (1850). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet von Joh. Christ. Albers: 1-262. Berlin. Drymaeus is on the page 155. Mesembrinus is on the page 157.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad ^ae ^af ^ag ^ah ^ai ^aj ^ak ^al ^am ^an ^ao ^ap ^aq ^ar ^as ^at ^au ^av ^aw ^ax ^ay ^az ^ba ^bb ^bc ^bd ^be ^bf ^bg ^bh ^bi ^bj ^bk ^bl ^bm ^bn ^bo ^bp ^bq ^br ^bs ^bt ^bu ^bv ^bw ^bx ^by ^bz ^ca ^cb ^cc ^cd ^ce ^cf ^cg ^ch ^ci ^cj ^ck ^cl ^cm ^cn ^co ^cp ^cq ^cr ^cs ^ct ^cu ^cv ^cw ^cx ^cy ^cz Thompson F. G. (16 June 2008). "AN ANNOTATED CHECKLIST AND BIBLIOGRAPHY OF THE LAND AND FRESHWATER SNAILS OF MEXICO AND CENTRAL AMERICA" Archived 2012-10-12 at the Wayback Machine. "PART 3 PART 3 (ORTHALICOIDEA)". accessed 17 May 2011.
^ ^a ^b MolluscaBase eds. (2020). MolluscaBase. Drymaeus Albers, 1850. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=879692 on 2020-10-26
^ ^a ^b ^c ^d ^e ^f ^g Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. doi:10.1127/arch.moll/1869-0963/141/001-020.
^ ^a ^b ^c ^d Breure A. S. H. & Mogollón Avila V. (2010). "Well-known and little-known: miscellaneous notes on Peruvian Orthalicidae (Gastropoda, Stylommatophora)". Zoologische Mededelingen 84. HTM Archived 2018-09-26 at the Wayback Machine.
^ Salvador, R.B.; Cavallari, D.C.; Simone, L.R.L. (2015). "Taxonomical study on a sample of land snails from southeastern Tocantins State, Brazil, with description of a new species". Journal of Conchology. 42 (1): 67–78.
^ "Mollusca" Archived July 18, 2011, at the Wayback Machine. Diversidad Biológica Cubana, accessed 23 March 2011.
^ Salvador R. B. & Simone L. R. L. (2016). "A new species of Kora from Bahia, Brazil (Gastropoda: Pulmonata: Orthalicoidea), with an emended diagnosis of the genus". Stuttgarter Beiträge zur Naturkunde A 9: 1–7. doi:10.18476/sbna.v9.a1.
^ Robinson D. G., Hovestadt A., Fields A. & Breure A. S. H. (July 2009). "The land Mollusca of Dominica (Lesser Antilles), with notes on some enigmatic or rare species". Zoologische Mededelingen 83 http://www.zoologischemededelingen.nl/83/nr03/a13 Archived 2011-10-07 at the Wayback Machine
^ Salvador, Rodrigo B.; Silva, Fernanda S.; Cavallari, Daniel C.; Köhler, Frank; Slapcinsky, John; Breure, Abraham S. H. (2023-07-26). "Molecular phylogeny of the Orthalicoidea land snails: Further support and surprises". PLOS ONE. 18 (7): e0288533. Bibcode:2023PLoSO..1888533S. doi:10.1371/journal.pone.0288533. ISSN 1932-6203. PMC 10370776. PMID 37494326.

External links

[Albers_1850-1] (in German) Albers J. C. (1850). Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet von Joh. Christ. Albers: 1-262. Berlin. Drymaeus is on the page 155. Mesembrinus is on the page 157.

[Thompson_2008-2] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y ^z ^aa ^ab ^ac ^ad ^ae ^af ^ag ^ah ^ai ^aj ^ak ^al ^am ^an ^ao ^ap ^aq ^ar ^as ^at ^au ^av ^aw ^ax ^ay ^az ^ba ^bb ^bc ^bd ^be ^bf ^bg ^bh ^bi ^bj ^bk ^bl ^bm ^bn ^bo ^bp ^bq ^br ^bs ^bt ^bu ^bv ^bw ^bx ^by ^bz ^ca ^cb ^cc ^cd ^ce ^cf ^cg ^ch ^ci ^cj ^ck ^cl ^cm ^cn ^co ^cp ^cq ^cr ^cs ^ct ^cu ^cv ^cw ^cx ^cy ^cz Thompson F. G. (16 June 2008). "AN ANNOTATED CHECKLIST AND BIBLIOGRAPHY OF THE LAND AND FRESHWATER SNAILS OF MEXICO AND CENTRAL AMERICA" Archived 2012-10-12 at the Wayback Machine. "PART 3 PART 3 (ORTHALICOIDEA)". accessed 17 May 2011.

[WoRMS-3] MolluscaBase eds. (2020). MolluscaBase. Drymaeus Albers, 1850. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=879692 on 2020-10-26

[Breure_2012-4] ^ ^a ^b ^c ^d ^e ^f ^g Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. doi:10.1127/arch.moll/1869-0963/141/001-020.

[Breure_2010-5] Breure A. S. H. & Mogollón Avila V. (2010). "Well-known and little-known: miscellaneous notes on Peruvian Orthalicidae (Gastropoda, Stylommatophora)". Zoologische Mededelingen 84. HTM Archived 2018-09-26 at the Wayback Machine.

[SalvadorCavallSimone42-6] Salvador, R.B.; Cavallari, D.C.; Simone, L.R.L. (2015). "Taxonomical study on a sample of land snails from southeastern Tocantins State, Brazil, with description of a new species". Journal of Conchology. 42 (1): 67–78.

[ecosis.cu-7] "Mollusca" Archived July 18, 2011, at the Wayback Machine. Diversidad Biológica Cubana, accessed 23 March 2011.

[Salvador_2016-8] Salvador R. B. & Simone L. R. L. (2016). "A new species of Kora from Bahia, Brazil (Gastropoda: Pulmonata: Orthalicoidea), with an emended diagnosis of the genus". Stuttgarter Beiträge zur Naturkunde A 9: 1–7. doi:10.18476/sbna.v9.a1.

[Robinson_2009-9] Robinson D. G., Hovestadt A., Fields A. & Breure A. S. H. (July 2009). "The land Mollusca of Dominica (Lesser Antilles), with notes on some enigmatic or rare species". Zoologische Mededelingen 83 http://www.zoologischemededelingen.nl/83/nr03/a13 Archived 2011-10-07 at the Wayback Machine

[10] Salvador, Rodrigo B.; Silva, Fernanda S.; Cavallari, Daniel C.; Köhler, Frank; Slapcinsky, John; Breure, Abraham S. H. (2023-07-26). "Molecular phylogeny of the Orthalicoidea land snails: Further support and surprises". PLOS ONE. 18 (7): e0288533. Bibcode:2023PLoSO..1888533S. doi:10.1371/journal.pone.0288533. ISSN 1932-6203. PMC 10370776. PMID 37494326.

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