Columella (genus)

Columella
	Six shells of Columella edentula, scale bar in mm
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Family:	Truncatellinidae
Genus:	Columella; Westerlund, 1878
Type species
	Pupa inornata Michaud, 1831;
Synonyms
	Edentulina Clessin, 1876; Paludinella R. T. Lowe, 1852;

Columella is a genus of very small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Truncatellinidae, the whorl snails and allies. ^[2]

Taxonomy

Columella is commonly classified within the Vertiginidae. However, as shown by Nekola & Coles (2016), the genus is not that closely related to the Vertiginidae, but may rather be affiliated with the Chondrinidae or form a pupilloid family of its own. A systematic revision is pending.^[2]

Species

Species within the genus Columella include:

Columella acicularis Almuhambetova, 1979
Columella aspera Waldén, 1966
Columella columella (G. v. Martens, 1830)
Columella edentula (Draparnaud, 1805) - toothless column
†Columella hartmutnordsiecki Schlickum & Geissert, 1980
Columella hasta (Hanna, 1911)
Columella intermedia Schileyko & Almuhambetova, 1984
Columella microspora (R.T. Lowe, 1852)
Columella nymphaepratensis Hlaváč & Pokryszko, 2009
Columella polvonense (Pilsbry, 1894)
Columella simplex (Gould, 1840)
Columella talgarica Schileyko & Rymzhanov, 2010
†Columella wyciski Schlickum & Strauch, 1972

Species brought into synonymy

Columella inornata (Michaud, 1831): synonym of Columella edentula (Draparnaud, 1805) (junior synonym)
Columella ninagongonis Pilsbry, 1935: synonym of Truncatellina ninagongonis (Pilsbry, 1935) (original combination)
Columella tridentata A. B. Leonard, 1946: synonym of Gastrocopta ruidosensis (Cockerell, 1899)

References

^ Westerlund C. A. (1876-1878). Fauna Europæa molluscorum extramarinorum. Prodromus. Sistens descriptiones systematicas et criticas omnium generum et specierum horum animalium in Europa viventium et hodie cognitarum. pp. 1-320. Lundae. (Berlingia).
^ ^a ^b ^c MolluscaBase eds. (2020). "Columella". WoRMS. World Register of Marine Species. Retrieved 22 September 2021. {{cite web}}: |author= has generic name (help)

[1] Westerlund C. A. (1876-1878). Fauna Europæa molluscorum extramarinorum. Prodromus. Sistens descriptiones systematicas et criticas omnium generum et specierum horum animalium in Europa viventium et hodie cognitarum. pp. 1-320. Lundae. (Berlingia).

[WoRMS-2] MolluscaBase eds. (2020). "Columella". WoRMS. World Register of Marine Species. Retrieved 22 September 2021. {{cite web}}: |author= has generic name (help)

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[2]