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Colhuehuapian

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The Colhuehuapian age is a period of geologic time (21.0–17.5 Ma) within the Early Miocene epoch of the Neogene, used more specifically within the SALMA classification in South America. It follows the Deseadan and precedes the Santacrucian age.[1]

Etymology

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The age is named after the Colhué Huapí Member of the Sarmiento Formation in the Golfo San Jorge Basin, Patagonia, Argentina.

Formations

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Colhuehuapian is located in South America
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Colhuehuapian
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Locations of Colhuehuapian formations
Formation
bold is type 		Country Basin Notes
Colhué Huapí Member  Argentina Golfo San Jorge Basin
[2][3]
Abanico Formation  Chile Abanico Basin
[4][5]
Biblián Formation  Ecuador Cuenca Basin
[6]
Castillo Formation  Venezuela Falcón Basin
[7]
Cerro Bandera Formation  Argentina Neuquén Basin
[8]
Chichinales Formation  Argentina North Patagonian Massif
[9]
Chilcatay Formation  Peru Pisco Basin
[10][11]
Cura-Mallín Group  Chile Cura-Mallín Basin
[12]
Gaiman Formation  Argentina Valdés Basin
[13]
Gran Bajo del Gualicho Formation  Argentina Colorado Basin
[14]
Monte León Formation  Argentina Austral Basin
[15][16]
Montera Formation  Peru Sechura Basin
[17]
Pirabas Formation  Brazil Amazon Basin
[18]
Río Jeinemeni Formation  Argentina Austral Basin
[19]
Uitpa Formation  Colombia Cocinetas Basin
[20]

Fossils

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Group Fossils Formation Notes
Mammals Abderites meridionalis, Acarechimys cf. minutissimus, Acaremys cf. murinus, A. preminutus, Acrocyon riggsi, Acyon tricuspidatus, Anargyrolagus primus, Arctodictis sinclairi, Argyrohippus cf. boulei, ?Astrapotherium ruderarium, Australoprocta fleaglei, Borhyaena macrodonta, Branisamyopsis australis, Caviocricetus lucasi, Cephalomyopsis hipselodontus, Cladosictis centralis, Clenia minuscula, Colpodon distinctus, Cramauchenia normalis, Dolichocebus gaimanensis, Dudumus ruigomezi, Eomicrobiotherium mykerum, Eosteiromys homogenidens, E. segregatus, Eoviscaccia australis, Euphractus patagonicus, Gaimanomys alwinea, Galileomys antelucanus, Hypsosteiromys axiculus, H. nectus, Licaphrops festinus, Luanthus initialis, Luantus propheticus, Mazzonicebus almendrae, Mormopterus barrancae, Necrolestes mirabilis, Neoreomys pinturensis, Oligobiotherium divisum, Pachybiotherium acclinum, Palaeothentes ?minutus, Parabderites bicrispatus, Paradelphomys fissus, Parasteiromys friantae, P. uniformis, Parastrapotherium symmetrum, Patagonia peregrina, Peltephilus pumilus, Perimys dissimilis, P. incavatus, P. transversus, Proargyrolagus argentinus, Proheptaconus patagonicus, Prolicaphrium spectabile, Prospaniomys priscus, Prostichomys bowni, Protacaremys avunculus, P. prior, P. pulchellus, Protadelphomys latus, Prozaedyus humilis, Pseudostegotherium glangeaudi, Pseudothylacynus rectus, Pternoconius bondi, Sipalocyon externus, Soriamys gaimanensis, S. ganganensis, Stegotherium tesselatum, S. variegatum, Tremacebus harringtoni, Willidewu estaparius, Acarechimys sp., Interatherium sp., Microbiotherium sp., Nematherium sp., Pachyrukhos sp., Proeutatus sp., Protypotherium sp., Prozaedyus sp., Soriamys sp., Tetramerorhinus sp., Titanothentes sp., Argyrolagoidea indet., Caluromyinae indet., ?Cephalomyidae indet., Chinchilloidea indet., Dasypodidae indet., Didelphidae indet., Didelphimorphia indet., Didelphoidea indet., Glyptodontidae indet., Hathlyacininae indet., Leontiniidae indet., Marsupialia indet., Megalonychidae indet., Neoepiblemidae indet., Phyllostominae indet., Propalaehoplophorinae indet., Thylacosmilidae indet. Colhué Huapí
[21][22]
[23][24]
[25][26]
Chilecebus carrascoensis, Notoungulata indet., Rodentia indet. Abanico
[27]
Xenastrapotherium aequatorialis Biblián
[28]
Banderomys leanzai, Caviocricetus lucasi, Comahuetherium coccaorum, Cramauchenia normalis, Doryperimys olsacheri, Eoviscaccia australis, Garridomys curunuquem, Leucokephalos maior, Proadinotherium cf. muensteri, Protypotherium sinclairi, Abderites sp., Astrapothericulus sp., Cladosictis sp., Eosteiromys sp., Palaeothentes sp., Parastrapotherium sp., Perimys sp., Pitheculites sp., Proeutatus sp., Protypotherium sp., Prozaedyus sp., Sipalocyon sp., Stenotatus sp., Astegotheriini indet., Borhyaenidae indet., Hegetotheriidae indet., Macraucheniidae indet., Peltephilidae indet., Platyrrhini indet., Propalaehoplophorinae indet., Proterotheriinae indet. Cerro
Bandera
[29][30]
[31][32]
[33][34]
Australoprocta fleaglei, Caviocricetus lucasi, Cladosictis cf. patagonica, Eoviscaccia australis, Willidewu esteparius, cf. Colpodon sp., Hegetotherium sp., Octodontoidea sp., Proeutatus sp., cf. Protypotherium sp., cf. Stenotatus sp., Interatheriidae indet., Mesotheriinae indet., Nesodontinae indet., Propalaehoplophorinae indet., Proterotheriinae indet. Chichinales
[35]
Ensidelphis riveroi, Inticetus vertizi, Notocetus vanbenedeni, Pinnipedia indet., Platanidelphidi indet. Chilcatay [36][37]
[38][39]
[40][41]
[42]
Colpodon antucoensis, Acarechimys sp., Luantus sp., Protacaremys sp., Protypotherium sp., Sipalocyon sp., Echimyidae indet. Cura-Mallín
[43][44]
Aglaocetus moreni, Aondelphis talen, Argyrocetus patagonicus, Idiorophus patagonicus, Morenocetus parvus, Phoberodon arctirostris, Plesiocetus dyticus, Prosqualodon australis, Kentriodon sp., Eurhinodelphinidae indet., Squalodontidae indet. Gaiman
[45]
[46][47]
Metaxytherium crataegense Montera
[48]
Dioplotherium allisoni, Sirenotherium pirabense, Metaxytherium sp., Rytiodus sp., Dugongidae indet. Pirabas
[49][50]
[51][52]
[53][54]
Phanomys mixtus Río
Jeinemení
[55]
Birds Falconidae indet. Cerro
Bandera
[34]
Opisthodactylus horacioperezi, Patagorhacos terrificus, Psilopteridae indet. Chichinales
[35][56]
Palaeospheniscus sp. Chilcatay [57]
Apterodytes ictus, Eretiscus tonnii, Palaeospheniscus affinis, P. bergi, P. bilocular, P. gracilis, P. intermedius, P. interruptus, P. patagonicus, P. planus, P. rothi, Paraptenodytes antarcticus, P. robustus, Treleudytes crassa, Arthrodytes sp., Anseriformes indet., Sphenisciformes indet. Gaiman
[46][47]
[58]
Reptiles Gaimanophis tenuis, Waincophis sp., Colubridae indet. Colhué Huapí
[24]
Geochelone sp., Pleurodira indet. Chichinales
[35]
Crocodylia indet., Gavialoidea indet., Pleurodira indet. Pirabas
[49][50]
[51][52]
Fish Alopias superciliosus, Alopias cf. vulpinus, Anotodus agassizii, Carcharhinus brachyurus, Carcharhinus cf. leucas, Carcharocles chubutensis, Carcharodon hastalis, Cosmopolitodus plicatilis, Galeocerdo aduncus, Hemipristis serra, Isurus desori, Isurus oxyrinchus, Megachasma cf. applegatei, Megalolamna paradoxodon, Negaprion brevirostris, Parotodus benedeni, Physogaleus contortus, Sphyrna zygaena, cf. Anoxypristis sp., Carcharias sp., Squatina sp., Dasyatidae indet., Myliobatoidea indet. Chilcatay [59]
Carcharhinus ackermanii, C. egertoni, C. priscus, Carcharodon subauriculatus, Diodon ferreirai, Galeocerdo paulinoi, Ginglymostoma obliquum, G. serra, Hemipristis serra, Isurus novus, Nebrius obliquus, Rhinoptera studeri, Scoliodon taxandriae, Sphyraena cunhai, S. egleri, Sphyrna magna, S. prisca, Arius sp., Hypoprion sp., Myliobatis sp. Pirabas
[49][50]
[51][52]
[53]
Heptranchias cf. howellii, Centrophorus sp., Dalatias cf. licha, Pristiophorus sp., Isurus cf. oxyrinchus, Carcharocles sp., Alopias cf. superciliosus, Hemipristis serra, Carcharhinus gibbesii, Sphyrna laevissima, Mobula sp., Lamniformes indet., Teleostei indet. Uitpa
[60]
Decapods Acanthocarpus obscurus, Arenaeus cribarius, Calappilia broksi, Callinectes cf. declivis, C. ferreirai, C. paraensis, C. pirabensis, C. reticulatus, Cyclocancer tuberculatus, Euphylax septendentatus, Hepatella amazonica, Necronectes tajinensis, Paratumidocarcinus marajoarus, Panopeus capanemensis, Parthenope tuberculata, Portunus atecuicitilis, P. haitensis, P. pirabensis, P. spimanus, P. spinimanus, Scylla costata, Sesarma paraensis, Tasadia tuberculatus, Tetraxanthus rathbunae, Typilobus unispinatus, Randallia sp. Pirabas
[49][50]
[52]

References

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  1. ^ Paleo Database: Colhuehuapian
  2. ^ Dozo et al., 2014, p.241
  3. ^ Acosta & Tambussi, 2005, p.128
  4. ^ Godoy et al., 2012
  5. ^ Muñoz et al., 2006
  6. ^ Veintimilla González, 2015, p.47
  7. ^ Johnson, 2009, p.60
  8. ^ Kramarz et al., 2005
  9. ^ Kramarz et al., 2005, p.288
  10. ^ De la Cruz, 2008, p.53
  11. ^ Solíz Mundaca, 2018, p.13
  12. ^ Utgé et al., 2009
  13. ^ Acosta Hospitaleche, 2007, p.419
  14. ^ Reichler, 2010, p.181
  15. ^ Náñez et al., 2009
  16. ^ Pérez Panera, 2010, p.52
  17. ^ Córdova Aguilar, 2009, p.24
  18. ^ De Araújo et al., 2010, p.207
  19. ^ Dal Molín & Colombo, 2003, p.140
  20. ^ Moreno et al., 2015, p.7
  21. ^ Cheme et al., 2016
  22. ^ Arnal et al., 2014
  23. ^ Gran Barranca Upper Fossil Zone at Fossilworks.org
  24. ^ a b Southern Cliff of lower Chubut River Valley at Fossilworks.org
  25. ^ Gran Barranca - southern cliff of lake Colhué Huapí at Fossilworks.org
  26. ^ Sacanana at Fossilworks.org
  27. ^ Río Las Lenas at Fossilworks.org
  28. ^ Point 354989 Anzogues at Fossilworks.org
  29. ^ Vucetich et al., 2015
  30. ^ Kramarz et al., 2015
  31. ^ Cerro Bandera Type at Fossilworks.org
  32. ^ Cerro Bandera Barda Negra at Fossilworks.org
  33. ^ Cerro Bandera Sierra del Portezuelo Norte at Fossilworks.org
  34. ^ a b Cerro Bandera Bayo Mesa at Fossilworks.org
  35. ^ a b c Paso Córdoba at Fossilworks.org
  36. ^ Lambert et al., 2017
  37. ^ Zamaca in the Paleobiology Database
  38. ^ Santa Rosa in the Paleobiology Database
  39. ^ Cerro Buque in the Paleobiology Database
  40. ^ Bianucci et al., 2020
  41. ^ Landini et al., 2019
  42. ^ Varas Malca et al., 2011
  43. ^ Shockey et al., 2012
  44. ^ Flynn et al., 2008, p.414
  45. ^ Viglino et al., 2018, p.264
  46. ^ a b Cerro del Castillo, Trelew at Fossilworks.org
  47. ^ a b Bryn Gwyn at Fossilworks.org
  48. ^ Cerro Chocan at Fossilworks.org
  49. ^ a b c d Jazida B17 at Fossilworks.org
  50. ^ a b c d Atalaia at Fossilworks.org
  51. ^ a b c Praia de Fortalezinha at Fossilworks.org
  52. ^ a b c d Ilha de Fortaleza; São João de Pirabas at Fossilworks.org
  53. ^ a b Colonia Pedro Teixeira at Fossilworks.org
  54. ^ Tauari at Fossilworks.org
  55. ^ Río Jeinemení at Fossilworks.org
  56. ^ Agnolin & Chafrat, 2015
  57. ^ Acosta Hospitaleche & Stucchi, 2005
  58. ^ Patagonian at Fossilworks.org
  59. ^ Alván et al., 2004
  60. ^ Carrillo Briceño et al., 2016, p.91

Bibliography

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Colhué Huapí Member
  • Arnal, Michelle; Kramarz, Alejandro G.; Vucetich, M. Guiomar; Vieytes, E. Carolina (2014), "A new early Miocene octodontoid rodent (Hystricognathi, Caviomorpha) from Patagonia (Argentina) and a reassessment of the early evolution of Octodontoidea", Journal of Vertebrate Paleontology, 34 (2): 397–406, Bibcode:2014JVPal..34..397A, doi:10.1080/02724634.2013.808203, hdl:11336/29748, retrieved 2019-02-12
  • Cheme Arriaga, Lucas; Dozo, María Teresa; Gelfo, Javier N. (2016), "A new Cramaucheniinae (Litopterna, Macraucheniidae) from the early Miocene of Patagonia, Argentina", Journal of Vertebrate Paleontology, 36 (6): e1229672, Bibcode:2016JVPal..36E9672C, doi:10.1080/02724634.2017.1229672, retrieved 2019-02-12
Abanico Formation
Biblián Formation
Castillo Formation
Cerro Bandera Formation
Chichinales Formation
Chilcatay Formation
Cura-Mallín Group
Gaiman Formation
Gran Bajo del Gualicho Formation
Monte León Formation
Montera Formation
Pirabas Formation
Río Jeinemení Formation
Uitpa Formation
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