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List of arthropods of the Cambrian Period

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This list contains many extinct arthropod[1] genera from the Cambrian Period of the Paleozoic Era. Some trilobites, bradoriids and phosphatocopines may not be included due to the lack of literature on these clades and inaccessibility of many papers describing their genera.

Genus Year of Discovery Location of Fossils
Anomalocaris[1][2] 1892 Emu Bay Shale

Maotianshan Shales

Burgess Shale

Pioche Shale

Cucumericrus[3] 1995 Maotianshan Shales
Paranomalocaris[4] 2013 Wulongqing Formation
Ramskoeldia[5] 2018 Maotianshan Shales
Cambroraster[6] 2019 Burgess Shale

Maotianshan Shales

Hurdia[7] 1912 Burgess Shale
Peytoia[8] 1911 Burgess Shale

Zawiszyn Formation

Ursulinacaris[9] 2019 Mount Cap Formation

Carrara Formation

Titanokorys[10] 2018 Burgess Shale
Cordaticaris[11] 2020 Zhangxia Formation
Pahvantia[12] 1981 Wheeler Shale

Marjum Formation

Amplectobelua [3] 1995 Maotianshan Shales

Burgess Shale

Houcaris[13] 2021 Maotianshan Shales

Pioche Shale

Pyramid Shale

Lyrarapax[14] 2014 Maotianshan Shales
Caryosyntrips[15] 2010 Burgess Shale

Wheeler Shale

Marjum Formation

Valdemiedes Formation

Tamisiocaris[16] 2010 Sirius Passet
Opabinia[17] 1912 Burgess Shale
Utaurora[18] 2022 Wheeler Formation
Kerygmachela[19] 1993 Sirius Passet
Laminacaris[20] 2018 Maotianshan Shales

Kinzers Formation

Pambdelurion[21] 1997 Sirius Passet
Mobulavermis[22] 2023 Pioche Shale
Echidnacaris[2] 2023 Emu Bay Shale
Utahnax[23] 2022 Wheeler Shale?
Buccaspinea[24] 2021 Marjum Formation
Guanshancaris[25] 2023 Guanshan biota
Omnidens[26] 2006 Maotianshan Shales

Xiaoshiba Lagerstatte

Innovatiocaris[27] 2022 Maotianshan Shales
Shucaris[28] 2024 Maotianshan Shales
Stanleycaris[29][30] 2018 Burgess Shale

Qingjiang biota

Youti[31] 2024 Maotianshan Shales
Siberion[32] 2011 Sinsk Formation
Jianshanopodia[33] 2006 Maotianshan Shales
Megadictyon[34] 1999 Maotianshan Shales
Parvibellus[35] (dubious[22]) 2022 Maotianshan Shales
Hadranax[36] 1998 Sirius Passet
Genus Year of Discovery Location of Fossils
Liangshanella 1997 Maotianshan Shales

Burgess Shale

Kunmingella 1912 Maotianshan Shales
Comptaluta 1968 Maotianshan Shales
Jiucunella 1991 Maotianshan Shales

Wirrapowie Limestone

Kunyangella 1965 Maotianshan Shales
Alutella[37] 1951 Scania
Albrunnicola[37] 1979 Borgholm Formation

Bastion Formation

Wirrapowie Limestone

Amphikeropsis[37] 2007 South Australia
Anabarochilina 1960 Weeks Formation

Marjum Formation

Beyrichona[37] 1886 Hanford Brook Formation

Maclean Brook Formation

Annge[37] 2014 Giles Creek Dolostone
Aluta[37] 1896 Niutitang Formation

Chamberlains Brook Formation

Bradoria[38] 1899 Dugald Formation

Hanford Brook Formation

Cambria[37] 1956 Pardailhan Formation
Cordubiella[39] 2008 Pedroche Formation
Dictyocharion[37] 2009 Thorntonia Limestone
Emeiella[37] 1975 Jince Formation
Escasona[38] 1902 Dugald Formation
Euzepaera[37] 1990 Wirrapowie Limestone
Haoia[37] 1990 Mernmerna Formation
Hipponicharion[37] 1886 Hanford Brook Formation
Indiana[37] 1902 Comley Limestone

Bright Angel Shale

Hanford Brook Formation

Dugald Formation

Kinzers Formation?

Indianites[37] 1931 Saint-jean-de-la-Riviere Formation
Indota[38] 1968 Murray Shale
Jinxinlingella[37] 1975 Wirrapowie Limestone
Konicekion[37] 1975 Jince Formation
Matthoria[37] 1997 Pardailhan Formation
Monceretia[37] 2005 Pardailhan Formation
Mononotella[38] 1931 Hanford Brook Formation
Navarana[37] 2017 Henson Gletscher Formation
Neokunmingella[37] 1974 Wirrapowie Limestone
Parahoulongdongella[37] 1990 Wirrapowie Limestone
Petrianna[40] 1975 Buen Formation
Pseudobeyrichona[37] 1990 Tannenknock Formation
Quadricona[37] 2011 Wirrapowie Limestone
Sanlangella[37] 2008 Yanwangbian Formation
Sinskolutella[37] 1998 Sinsk Formation
Spinospitella[37] 2006 Mernmerna Formation
Tscholponaella[37] 1997 Beshtash Formation
Uskutchiella[37] 1997 Tandoshka Formation
Walcottella 1931 Bright Angel Shale

Riley Formation?

Wimanicharion[37] 1993 Valdemiedes Formation
Zepaera[37] 1973 Ajax Limestone
Cambroarchilocus[37] 2020 Puerto Blanco Formation
Fuxianhuia[41][42] 1987 Maotianshan Shales
Alacaris[43] 2018 Xiaoshiba Lagerstatte
Chengjiangocaris[42] 1991 Maotianshan Shales
Guangweicaris[44] 2007 Guanshan biota
Tauricornicaris[45] 2017 Maotianshan Shales
Isoxys[46] 1890 Burgess Shale

Maotianshan Shales

Sirius Passet

Balang Formation

Spence Shale

Sinsk Formation

Emu Bay Shale

Shujintuo Formation

Rockslide Formation

Kaili Biota

Mantou Formation

Pedrouche Formation

Surusicaris[47] 2015 Burgess Shale
Kylinxia[48] 2020 Maotianshan Shales
Erratus[49] 2022 Helinpu Formation
Fengzhengia[50] 2022 Maotianshan Shales
Xiaocaris[51] 2020 Maotianshan Shales
Shankouia[52] 2005 Maotianshan Shales
Liangwangshania[53] 2005 Maotianshan Shales
Genus Year of Discovery Location of Fossils
Sanctacaris[54][55] 1988 Burgess Shale
Habelia[56] 1912 Burgess Shale
Mollisonia[57][58] 1912 Maotianshan Shales

Burgess Shale

Langston Formation

Wheeler Shale

Thelxiope[59][60] 1975 Burgess Shale

Wheeler Shale

Fezouata Formation

Linyi Lagerstätte

Cambropycnogon[61] 2002 Orsten Fauna
Wisangocaris[62] 2016 Emu Bay Shale
Utahcaris[63] 1988 Spence Shale
Actaeus 1970 Burgess Shale
Alalcomenaeus[64] 1970 Burgess Shale

Chengjiang Biota

Wheeler Shale

Leanchoilia[65] 1912 Burgess Shale

Chengjiang Biota

Spence Shale

Fortiforceps[66] 1997 Maotianshan Shales
Yawunik[67] 2015 Burgess Shale
Parapeytoia [3] 1995 Maotianshan Shales
Jianfengia[68] 1987 Maotianshan Shales
Haikoucaris[69] 2004 Maotianshan Shales
Kanoshoia[70] 2020 Wheeler Formation
Sklerolibyon[71] 2020 Maotianshan Shales
Tanglangia[72][73] 1999 Maotianshan Shales

Emu Bay Shale

Oestokerkus[74] 2011 Emu Bay Shale
Yohoia[75] 1912 Burgess Shale
Genus Year of Discovery Location of Fossils
Branchiocaris[76] 1929 Wheeler Shale

Burgess Shale

Tokummia[77] 2017 Burgess Shale
Canadaspis[78] 1978 Burgess Shale

Pioche Shale

Maotianshan Shales

Waptia[79] 1912 Burgess Shale
Pectocaris[80] 1999 Maotianshan Shales
Clypecaris[80] 1999 Maotianshan Shales
Pauloterminus[81] 2002 Sirius Passet
Perspicaris[82] 1977 Burgess Shale
Nereocaris[83] 2012 Burgess Shale
Plenocaris 1912 Burgess Shale

Maotianshan Shales

Fibulacaris[84] 2019 Burgess Shale
Synophalos[85] 2009 Maotianshan Shales
Odaraia[86] 1912 Burgess Shale
Protocaris[87] 1884 Parker Formation
Hymenocaris 1853 Burgess Shale
Loricicaris[88] 2014 Burgess Shale
Jugatacaris[89] 2011 Maotianshan Shales
Erjiecaris[90] 2014 Maotianshan Shales
Balhuticaris[91] 2022 Burgess Shale
Xiazhuangocaris[92] 2021 Maotianshan Shales
Vermontcaris[93] 2022 Parker Quarry
Pseudoarctolepis[94] 1956 Wheeler Shale
Pakucaris[95] 2021 Burgess Shale
Duplapex[96] 2021 Qingjiang biota
Tuzoia[97][98] 1912 Burgess Shale

Emu Bay Shale

Kaili Biota

Spence Shale

Wheeler Formation

Marjum Formation

Kinzers Formation

Sinsk Formation

Parker Shale

Wulongqing Formation

Tangshih Formation

Eager Formation

Tsinghsutung Formation

Balang Formation

Heymonsicambria[99] 1994 Orsten Fauna
Skara[100] 1983 Orsten Fauna
Goticaris[101] 1990 Orsten Fauna
Wujicaris[102] 2010 Maotianshan Shales
Cambropachycope[101] 1990 Orsten Fauna
Ercaia[103] 2001 Maotianshan Shales
Mosineia[104] 2010 Blackberry Hill
Cambrocaris[105] 1991 Orsten Fauna
Bredocaris[100] 1983 Orsten Fauna
Dala[100] 1983 Orsten Fauna
Sandtorpia[106] 2009 Orsten Fauna
Walossekia[100] 1983 Orsten Fauna
Rehbachiella[100] 1983 Orsten Fauna
Martinssonia[107] 1986 Orsten Fauna
Musacaris[108] 2010 Orsten Fauna
Henningsmoenicaris[101] 1990 Orsten Fauna
Oelandocaris[100] 1983 Orsten Fauna
Aengapentastomum[109] 2005 Orsten Fauna
Boeckelericambria[99] 1994 Orsten Fauna
Haffnericambria[99] 1994 Orsten Fauna
Klausmuelleria[110] 2003 Comley Limestone
Mictomerus[104] 2010 Potsdam Group
Apankura[111] 2004 Santa Rosita Formation
Arenosicaris[104] 2010 Blackberry Hill
Dietericambria[112] 2022 Henson Gletscher Formation
Ercaicunia[113] 1999 Maotianshan Shales
Yicaris[114] 2007 Maotianshan Shales


Genus Year of Discovery Location of Fossils
Fritzaspis[115] 2007 Campito Formation
Bathyuriscus 1873 Burgess Shale
Urokodia[116] 1989 Maotianshan Shales
Acanthomeridion[117] 1989 Maotianshan Shales
Beckwithia 1931 Weeks Formation
Tremaglaspis[118] 2003 Weeks Formation
Australaglaspis[119] 2010 Stony Point
Aglaspella[120][121] 1939 Sandu Formation

St. Lawrence Formation

Hesselbonia[120] 2013 Trempealeau Formation
Aglaspis 1862 Potsdam Sandstone
Aglaspoides[121] 1939 St. Lawrence Formation
Chraspedops[121] 1939 St. Lawrence Formation
Cyclopites[121] 1939 St. Lawrence Formation
Flobertia[121] 1992 St. Lawrence Formation
Setaspis[121] 1939 St. Lawrence Formation
Tuboculops[121] 1992 St. Lawrence Formation
Olenoides 1877 Burgess Shale

Marjum Formation

Hanburia 1916 Burgess Shale

Stephen Formation

Xiuqiella 1974
Acontheus 1851
Hemirhodon 1937
Thoracocare 1974 Spence Shale Formation
Abakania 1973 Sanashtykgol Formation
Zacanthoides 1888 Burgess Shale
Changaspis 1961
Ogygopsis 1889 Burgess Shale
Oryctocephalus 1886 Burgess Shale
Chancia 1924 Burgess Shale
Ehmaniella 1951 Burgess Shale
Elrathia 1924 Wheeler Formation

Burgess Shale

Ellipsocephalus 1883 Słowiec Sandstone Formation
Phantaspis[122] 2020 Mantou Formation
Lisania 1911
Bailiaspis 1936 Whitesands Bay Formation

Jbel Lmgaysmat Formation

Manuels River Formation

Maentwrog Formations

Solenoparia 1935
Westergaardites 1937
Yunnanocephalus 1936 Maotianshan Shales
Asaphiscus 1873
Olenus 1827
Quitacetra 1967
Ejinaspis 1986
Asthenopsis 1939
Agraulos 1847 Menevia Formation

Jince Formation

Manuels River Formation

Coosella 1936
Norwoodia 1916
Tricrepicephalus 1935 Chefu Formation

Deadwood Formation

Little Elk Creek

Meniscopsia[123] 2011
Acrocephalella 1963
Cambroproteus[124] 2015
Cedaria 1924 Weeks Formation
Conocoryphe 1847
Galeaspis 1983
Genevievella 1936 Mungerbar Formation
Meteoraspis 1890 Limestone Formation
Validaspis 1977
Palaeolenus 1912
Ptychoparia 1847
Kootenia 1939 Burgess Shale

Emu Bay Shale

Misszhouia[125] 1985 Maotianshan Shales
Naraoia[126][127] 1912 Burgess Shale

Stephen Formation

Maotianshan Shales

Bertie Group

Balang Formation

Emu Bay Shale

Buenaspis[128] 1999 Sirius Passet
Emucaris[129] 2010 Emu Bay Shale
Kangacaris[129][130] 2010 Emu Bay Shale

Maotianshan Shales

Liwia 1988 Zawiszyn Formation
Estaingia 1964
Bathynotus 1860
Bajanaspis 1978 Edreisk Formation
Kleptothule[131] 1995 Sirius Passet
Acadoparadoxides 1957
Paradoxides 1822 Duda Formation
Eoredlichia 1950 Maotianshan Shales
Profallotaspis 1965
Lochmanolenellus 1952 Poleta Formation
Olenellus 1861 Pioche Shale

Carrara Formation

Kinzers Formation

Archaeaspis 1965 Campito Formation
Parapaokannia 1980
Plutonides 1985 Purley Shale Formation
Redlichia 1902
Repinaella 1965 Pestrotsvet Formation
Ushbaspis 1965 Shabakty Formation
Balcoracania 1970 Balcoracana Creek
Emuella 1970 Emu Bay Shale
Megapharanaspis[132] 2006
Wutingaspis 1944
Mesolenellus 1993 Sofiekammen Formation
Olenelloides 1894
Biceratops 1971 Pioche Shale
Bradyfallotaspis 1972
Eopeachella[133] 2009 Delamar Mountains
Laudonia 1956
Mummaspis 1992 Mural Formation
Nephrolenellus 1993 Pyramid Shale Member

Carrara Formation

Nevadella 1936 Poleta Formation

Sekwi Formation

Paranephrolenellus[134] 2007
Wanneria 1910 Kinzers Formation
Bristolia 1956 Carrara Formation
Emigrantia 1910
Bolbolenellus 1993
Buenellus 1988 Sirius Passet
Callavia 1890 Brigus Formation
Chengjiangaspis 1980 Maotianshan Shales
Cirquella 1993
Daguinaspis 1950 Amouslek Formation
Fallotaspis 1953
Holmia 1890 Sparagmite Formation
Hydrocephalus 1846
Fritzolenellus 1998
Mesonacis 1859 Forteau Formation
Xystridura 1936
Peachella 1910
Richterops 1954
Archaeagnostus 1939
Agnostus 1822 Marjum Biota

Linyi Lagerstätte

Orsten Fauna

Itagnostus 1979
Peronopsis 1847
Acmarhachis 1938
Acadagnostus 1939
Agnostotes 1963
Lejopyge 1828
Glyptagnostus 1936
Egyngolia 1980
Jinghediscus 1985
Pagetia 1916 Burgess Shale
Acimetopus 1966
Pentagnostus 1940
Calodiscus 1935 Schodack Formation
Mallagnostus 1935
Runcinodiscus 1976
Serrodiscus 1941 Brigus Formation
Weymouthia 1913
Analox 1966
Litometopus 1966
Meniscuchus 1975
Ninadiscus 1980 Egyngolskaya Suite
Oodiscus 1966
Semadiscus 1978
Stigmadiscus 1966
Parapagetia 1964
Tannudiscus 1959 Purley Shale Formation
Chelediscus 1966
Bathydiscus 1966
Cobboldites 1943
Acidiscus 1966
Condylopyge 1847
Delgadella 1908 Tyuser Formation
Dicerodiscus 1954
Pleuroctenium 1847
Ptychagnostus 1909
Bolboparia 1966
Tchernyshevioides 1975
Toragnostus 1988
Tsunyidiscus 1966
Birmanites 1934
Kendallina 1953 Bison Creek Formation
Dikelocephalus 1852
Orygmaspis 1938 Morgan Creek Member
Helmetia 1912 Burgess Shale

Jince Formation

Tegopelte[135] 1975 Burgess Shale

Kicking Horse Shale

Retifacies[136] 1989 Maotianshan Shales
Schmalenseeia[137] 1903
Burlingia[138] 1908
Entomaspis 1931
Neodrepanura[139] 2006 Maotianshan Shales
Emeraldella[140] 1912 Burgess Shale

Wheeler Shale

Molaria[141] 1912 Burgess Shale

Sirius Passet

Sidneyia[142][143] 1911 Burgess Shale

Maotianshan Shales

Bailongia[144] 2021 Maotianshan Shales
Luohuilinella[145][146] 2012 Maotianshan Shales
Panlongia[147] 2006 Guanshan fauna
Pygmaclypeatus[148] 2000 Maotianshan Shales
Sinoburius[149] 1991 Maotianshan Shales
Kodymirus[150] 1965 Paseky Shale
Glypharthrus[151][152] 1939 McKay Group

Clay Creek

Guole Formation

Aaveqaspis[153] 2009 Sirius Passet
Thulaspis[154] 2023 Sirius Passet
Saperion[155] 1991 Maotianshan Shales
Squamacula[66][156] 1997 Maotianshan Shales

Emu Bay Shale

Falcatamacaris[157] 2014 Weeks Formation
Zhiwenia[158] 2019 Xiaoshiba Lagerstatte
Australimicola[156] 2012 Emu Bay Shale
Tonglaiia[143] 2023 Maotianshan Shales
Arthroaspis[159] 2013 Sirius Passet
Kwanyinaspis[160] 2005 Maotianshan Shales
Campanamuta[161] 2011 Sirius Passet
Rhombicalvaria 1987 Maotianshan Shales
Haifengella[162] 2014 Maotianshan Shales
Skioldia 1997 Maotianshan Shales
Kuamaia 1987 Maotianshan Shales
Cindarella 1996 Maotianshan Shales
Xandarella 1991 Maotianshan Shales
Zhugeia[143] 2023 Maotianshan Shales
Eozetetes 2016 Emu Bay Shale
Genus Year of Discovery Location of Fossils
Marrella[163] 1912 Burgess Shale

Kaili Formation

Balang Formation

Skania[164] 1931 Maotianshan Shales

Burgess Shale

Primicaris[165] 2003 Maotianshan Shales
Kiisortoqia[166] 2010 Sirius Passet
Carnarvonia 1912 Burgess Shale
Vladicaris[150] 1995 Paseky Shale
Burgessia 1912 Burgess Shale
Khankaspis 1969 Snegurovka Formation
Strabops 1901 Potosi Dolomite
Paleomerus 1956 Ociesęki Sandstone Formation

Mickwitzia Sandstone

Parapaleomerus 1999 Maotianshan Shales
Worthenella 1911 Burgess Shale
Chuandianella[167] 1975 Maotianshan Shales
Bushizheia[168] 2020 Maotianshan Shales
Combinivalvula 1987 Maotianshan Shales
Diplopyge 1999 Maotianshan Shales
Dongshanocaris 1999 Maotianshan Shales
Jianshania 1999 Maotianshan Shales
Pseudoiulia 1998 Maotianshan Shales
Priscansermarinus 1981 Burgess Shale
Kootenichela[169] 2013 Burgess Shale
Forfexicaris[80] 1999 Maotianshan Shales
Yunnanocaris[80] 1999 Maotianshan Shales
Kockurus[150] 1995 Paseky Shale
Occacaris[80] 1999 Maotianshan Shales
Sarotrocercus[170] 1981 Burgess Shale
Messorocaris[171] 2017 Wheeler Shale
Dicranocaris[172] 2008 Wheeler Shale
Nettapezoura[172] 2008 Wheeler Shale
Siriocaris[173] 2009 Sirius Passet

Notes

[edit]
1.^ (here taken to mean all members of Deuteropoda, Dinocaridida and everything closer to those than any other extant phylum)

References

[edit]
  1. ^ Whiteaves JF (1892). "Description of a new genus and species of phyllocarid Crustacea from the Middle Cambrian of Mount Stephen, B.C." Canadian Record of Science. 5 (4): 205–208.
  2. ^ a b Paterson, John R.; García-Bellido, Diego C.; Edgecombe, Gregory D. (January 2023). "The early Cambrian Emu Bay Shale radiodonts revisited: morphology and systematics". Journal of Systematic Palaeontology. 21 (1). Bibcode:2023JSPal..2125066P. doi:10.1080/14772019.2023.2225066.
  3. ^ a b c Xian-Guang, Hou; Bergström, Jan; Ahlberg, Per (September 1995). "Anomalocaris and other large animals in the lower Cambrian Chengjiang fauna of southwest China". GFF. 117 (3): 163–183. Bibcode:1995GFF...117..163X. doi:10.1080/11035899509546213.
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