List of things named after J. R. R. Tolkien and his works
The British author J. R. R. Tolkien (1892–1973) and the names of fictional characters and places he invented for his legendarium have had a substantial impact on culture, and have become the namesakes of various things around and outside the world, including street names, mountains, companies, species of animals and plants, asteroids, and other notable objects.
Astronomy
[edit]- The asteroids 2991 Bilbo and 2675 Tolkien were both discovered and named in 1982.[1][2]
- The Kuiper Belt object 385446 Manwë and its moon Thorondor were discovered in 2003.[3][4]
- Earendel, the most-distant known star.[5]
- The nickname "Eye of Sauron" has been given to multiple eye-like objects, namely the planetary nebulae M 1-42[6][7] and Helix Nebula,[8][9] the star system HR 4796A,[10][11] and the intermediate spiral Seyfert galaxy NGC 4151.[12]
- The Mars-crossing asteroid 378214 Sauron was discovered in 2007.[13]
- The trans-Neptunian object 174567 Varda and its moon Ilmarë were discovered in 2006 and 2011 (respectively) and named in 2014.[14]
Geography of Titan
[edit]By convention, certain classes of features on Saturn's moon Titan are named after elements from Middle-earth.[15] Colles (small hills or knobs) are named for characters,[16] while montes (mountains) are named for mountains of Middle-earth.[17]
Colles
[edit]Collis[16] | Coordinates | Diameter (km) | Named after |
---|---|---|---|
Arwen Colles | 7°30′S 250°00′W / 7.5°S 250.0°W | 64 | Arwen, character from The Lord of the Rings |
Bilbo Colles | 4°12′S 38°36′W / 4.2°S 38.6°W | 164 | Bilbo Baggins, titular character of The Hobbit |
Faramir Colles | 4°00′N 153°48′W / 4.0°N 153.8°W | 82 | Faramir, character from The Lord of the Rings |
Gandalf Colles | 14°36′N 209°30′W / 14.6°N 209.5°W | 102 | Gandalf, character from The Lord of the Rings |
Handir Colles | 10°00′N 356°42′W / 10.0°N 356.7°W | 100 | Handir, character from The Silmarillion |
Nimloth Colles | 11°54′N 151°18′W / 11.9°N 151.3°W | 90 | Nimloth, name of a character and a tree from Middle-earth |
Montes
[edit]Other Planetary bodies
[edit]Mercury
[edit]A crater adjacent the planet's north pole, Tolkien, is named after the eponymous author.
Pluto System
[edit]Various maculae on Pluto and Charon are unofficially named after subjects in Middle Earth.
Maculae | Object | Coordinates | Named after |
---|---|---|---|
Balrog Macula | Pluto | 10°S 80°W / 10°S 80°W | Balrog, a race of demons from the mythos.[18][19] |
Mordor Macula | Charon | 80°N 00°W / 80°N -0°E | Mordor[20][21] It is named after the shadow lands in Tolkien's The Lord of the Rings, which it resembles in shape.[22] |
Morgoth Macula | Pluto | 20°S 172°E / 20°S 172°E | Morgoth, the dark lord of Arda and the main antagonist up to the First Age. |
Companies and other entities
[edit]- Iron Crown Enterprises produces role playing, board, miniature, and collectible card games since 1980. Many of ICE's better-known products were related to Tolkien's world of Middle-earth. It was named after the crown worn by Morgoth.[23]
- Middle-earth Enterprises, formerly known as Tolkien Enterprises, is a trading name for a division of The Saul Zaentz Company, located in Berkeley, California. The company owns the worldwide exclusive rights to certain elements of J. R. R. Tolkien's two most famous literary works: The Lord of the Rings and The Hobbit. These elements include the titles of the works, the names of characters contained within as well as the names of places, objects and events within them, and certain short phrases and sayings from the works.[24]
- Palantir Technologies is a private American software and services company, specializing in data analysis. Named after the crystal balls from Tolkien's legendarium, Palantir's original clients were federal agencies of the United States Intelligence Community like CIA and NSA.[25]
- Lembas Capital is a San Francisco-based investment firm named after the Elven waybread that appears in The Lord of the Rings and The Silmarillion. The company invests in both public equity and private equity.[26]
- The Tolkien Estate is the legal body which manages the property of J. R. R. Tolkien, including the copyright in his works. The individual copyrights have for the most part been assigned by the Estate to subsidiary entities such as the J. R. R. Tolkien Discretionary Settlement and The Tolkien Trust.[27][28][failed verification]
- Anduril Industries[29]
- Mithril, a decentralized social media platform[30]
- The Rivendell Winery operated from 1987 to December 2008 in New York's Hudson River Valley; in 2003 Rivendell's 2003 Dry Riesling captured the Governor's Cup at the 19th annual New York Wine and Food Classic.[31]
- Rivendell Bicycle Works, a steel frame bicycle producer in Walnut Creek, California, U.S.
- An astrophysical method for locating gamma ray bursts has been named BALROG, for "BAyesian Location Reconstruction Of GRBs".[32]
- A mining company called Durin Drilling is named about the Dwarven ling of kings named Durin.[33]
Genes and proteins
[edit]- Smaug, a protein that inhibits translation of mRNA nanos (Greek for dwarf) in Drosophila embryos. Named after the dragon Smaug from The Hobbit.[34]
- Glorund, a protein that inhibits translation of mRNA nanos in Drosophila ovaries. Named after Glórund, an early name for Glaurung, the first dragon in Tolkien's legendarium.[34]
- Bard, a gene in Drosophila that encodes the protein Bard, which is essential in degrading the protein Smaug. Named after Bard the Bowman, who killed Smaug in The Hobbit.[35]
Individual plants and animals
[edit]- Iluvatar is a redwood tree in Prairie Creek Redwoods State Park in Northern California that has been confirmed to be at least 20.5 feet (6.2 m) in diameter at breast height, and 320 feet (98 m) in height. Measured by botanist Stephen C. Sillett, it is the world's third-largest coast redwood, the largest being Lost Monarch.[36]
- Silmaril is a retired American thoroughbred mare racehorse named after the Silmarils featured in The Silmarillion.[37]
Geographical features
[edit]Mountains
[edit]- Three mountains in the Cadwallader Range of British Columbia, Canada: Mount Shadowfax, Mount Gandalf and Mount Aragorn.[38][39]
- On 1 December 2012, a bid was launched for the New Zealand Geographic Board to name a mountain peak near Milford Sound after Tolkien to mark Tolkien's 121st birthday.[40]
Seamounts
[edit]Several undersea features in the North Atlantic Ocean, west of Ireland and south of Iceland,[41] including:
- Eriador Seamount[42]
- Rohan Seamount[43]
- Gondor Seamount[44]
- Fangorn Bank[45]
- Edoras Bank[46]
- Isengard Ridge[47]
At least three seamounts in the Indian Ocean, including:[48]
Music
[edit]- Amon Amarth, a Swedish melodic death metal band, that takes its name from the Sindarin name of Mount Doom.[49]
- Burzum, a Norwegian music project founded by Varg Vikernes in 1991. The word "burzum" means "darkness" in the Black Speech, a fictional language crafted by Tolkien.[50]
- Ephel Duath, an Italian avant garde metal/hardcore punk band, formed in 1998 took their name after the mountain range in The Lord of the Rings.[51]
- Gorgoroth, a Norwegian black metal band, named after the dead plateau of darkness in the land of Mordor.[52]
- Iluvatar (band) , a neo-prog band from Baltimore, US, took their name from Eru Iluvatar.[53]
- Marillion, a British rock band, formed in 1979, was named "The Silmarillion", but was shortened to Marillion in 1981 to avoid potential copyright conflicts.[54][55]
- Shadowfax, a new-age group, took its name from Gandalf the White's horse Shadowfax.[56]
Ships
[edit]- J.R. Tolkien, a gaff-topsail schooner of Netherlands registry used for passenger cruises on the Baltic Sea and elsewhere in European waters, was built in 1964, and renamed in honour of Tolkien in 1998.[57][58][59]
Places
[edit]Streets
[edit]The "Tolkien Road" in Eastbourne, East Sussex, was named after Tolkien whereas the "Tolkien Way" in Stoke-on-Trent is named after Tolkien's eldest son, Fr. John Francis Tolkien, who was the priest in charge at the nearby Roman Catholic Church of Our Lady of the Angels and St. Peter in Chains.[60] In the Hall Green and Moseley areas of Birmingham, parks and walkways are dedicated to J. R. R. Tolkien, especially at the Millstream Way and Moseley Bog.[61] Collectively the parks are known as the Shire Country Parks.[61] In Weston-super-Mare, Somerset, England there is a collection of roads in the 'Weston Village' named after locales of Middle Earth, namely Hobbiton Road, Bree Close, Arnor Close, Rivendell, Westmarch Way and Buckland Green.[62]
In the Dutch town of Geldrop, near Eindhoven, the streets of an entire new neighbourhood are named after Tolkien himself ("Laan van Tolkien") and some of the best-known characters from his books.[63]
Two streets at Warsaw, Poland were named in 2022 after J.R.R. Tolkien and Gandalf. Both streets are located at neighbourhood commonly called Mordor.[64]
Housing
[edit]The Bend, Oregon housing development Forest Creek (formerly "The Shire") features the Tolkien-inspired names Ring Bearer Court, Shire Lane, and Wizard Lane.[65] One of the student housing complexes at the University of California, Irvine is named Middle Earth; its halls and other facilities were selected from Tolkien's legendarium.[66]
In the Silicon Valley towns of Saratoga and San Jose in California, there are two housing developments with street names drawn from Tolkien's works. About a dozen Tolkien-derived street names also appear scattered throughout the town of Lake Forest, California. The Columbia, Maryland, neighbourhood of Hobbit's Glen and its street names (including Rivendell Lane, Tooks Way, and Oakenshield Circle) come from Tolkien's works.[67]
Taxonomy
[edit]It has been noted that "Tolkien has been accorded formal taxonomic commemoration like no other author."[68] In the field of taxonomy, over 200 taxa (genera and species) have been given scientific names honouring, or deriving from, characters or other fictional elements from The Lord of the Rings, The Hobbit, and other works set in Middle-earth.[25]
Several taxa have been named after the character Gollum (also known as Sméagol), as well as for various hobbits, the small humanlike creatures such as Bilbo and Frodo Baggins. Various elves, dwarves, and other creatures that appear in his writings. Tolkien himself has been honoured in the names of several species. In 1978, paleontologist Leigh Van Valen named over 20 taxa of extinct mammals after Tolkien lore in a single paper.[69][70] The entomologist Karl-Johan Hedqvist, also a fan of Tolkien, named several wasp genera after Tolkien's characters.[71] In 1999, entomologist Lauri Kaila described 46 new species of Elachista moths and named 38 of them after Tolkien mythology.[25][72]
In 2004, the extinct hominid Homo floresiensis was described, and quickly earned the nickname "hobbit" due to its small size.[73]
- Pink background shading indicates that this species name has been synonymised and is no longer in use
J. R. R. Tolkien
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Martesia tolkieni Kennedy, 1974 | Clam | J. R. R. Tolkien | "The name tolkieni honors the late J.R.R. Tolkien, creator of The Hobbit, The Lord of the Rings, and many delightful creatures of long ago in the time of Middle-earth." | [74] |
Leucothoe tolkieni Vinogradov, 1990 | Amphipod | J. R. R. Tolkien | [75][76] | |
Gabrius tolkieni Schillhammer 1997 | Beetle | J. R. R. Tolkien | [77] | |
Tolkienia † Lieberman & Kloc, 1997 | Trilobite | J. R. R. Tolkien | A genus of Devonian trilobites that has been found in Spain, France and the United States. | [78] |
Khamul tolkeini Gates, 2008 | Wasp | J. R. R. Tolkien | "named in honor of J. R. R. Tolkein [sic] for his profound impact on the fantasy literature genre." | [79] |
Shireplitis tolkieni Fernández-Triana & Ward, 2013 | Wasp | J. R. R. Tolkien | [80] | |
Drassodella tolkieni Mbo et al, 2019 | Spider | J. R. R. Tolkien | "Named after John Ronald Reuel Tolkien, who was born in Bloemfontein, Free State, South Africa on 3rd January 1892 and died on 2nd September 1973. He is internationally recognised for his authorship of The Hobbit, The Lord of the Rings and The Silmarillion, amongst other works. His fictional 'Middle Earth' is believed to have been inspired in part by the exceptional natural scenery of Hogsback, the type locality of this species." | [81] |
Hyloscirtus tolkieni Sánchez-Nivicela et al, 2023 | Frog | J. R. R. Tolkien | "The specific epithet tolkieni is in honour of the writer, poet, philologist, and academic John Ronald Reuel Tolkien (J.R.R. Tolkien, 1892–1973), creator of Middle-earth and author of fantasy works like “The Hobbit” and “The Lord of the Rings”. The amazing colours of the new species evoke the magnificent creatures that seem to only exist in fantasy worlds."; "In a stream in the forest there lived a Hyloscirtus. Not a nasty, dirty stream, with spoor of contamination and a muddy smell, nor yet a dry, bare, sandy stream with nothing in it to perch on or to eat: it was a Hyloscirtus-stream, and that means environmental quality." | [82] |
Ainur
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Macropsidius sauroni (Hamilton, 1972) | Leafhopper | Sauron | [83][84] | |
Balrogia Hedqvist, 1977 | Wasp | Balrogs | "The new genus is named after a monster called Balrog in Tolkin's book 'The Lord of the Rings'." | [85] |
Mimatuta morgoth † Van Valen, 1978 | Fossil mammal | Morgoth | "Fëanor's name for Melkor, the power-lustful Vala of The Silmarillion. Reference is to the Hell Creek Formation." | [70] |
Mithrandir † Van Valen, 1978 | Fossil mammal | Gandalf (Mithrandir) | "Mithrandir (Sindarin, gray wanderer), Elvish name for Olorin, wisest of the Istari in The Lord of the Rings. Reference is to the subtleness of the differences between the subgenera." | |
Niphredil radagasti † Van Valen, 1978 | Fossil mammal | Radagast | "Radagast, naturalist of the Istari in The Lord of the Rings." | |
Helferella gothmogoides Williams & Weir, 1988 | Beetle | Gothmog, Lord of the Balrogs | "The species name is derived from Gothmog, a commander within the evil host of Mordor, and oides, latin suffix signifying 'resembling'." | [86] |
Macrostyphlus gandalf Morrone, 1994 | Beetle | Gandalf | [87] | |
Semicytherura balrogi Brouwers, 1994 | Ostracod | Balrogs | "After the Balrog, an evil character in J.R.R. Tolkien's adventures of Middle Earth" | [88] |
Semicytherura tauronae Brouwers, 1994 | Ostracod | Oromë (Tauron) | "After Tauron, a character in J.R.R. Tolkien's adventures of Middle Earth." | |
Sauron Eskov, 1995 | Spider | Sauron | The zoologist Kirill Eskov, author of The Last Ringbearer, a retelling of The Lord of the Rings from Mordor's point of view, has named a genus of linyphiid sheet weaver spiders Sauron after the ruler of Mordor.[89] | [90] |
Elachista olorinella Kaila, 1999 | Moth | Gandalf (Olórin) | From Gandalf's alternate name Olórin | [72] |
Elachista tauronella Kaila, 1999 | Moth | Oromë (Tauron) | ||
Paraortygoides radagasti † Dyke & Gulas, 2002 | Bird | Radagast | "For the wizard of Middle Earth, Radagast the Brown, rabid communicator with birds" | [91] |
Gandalfia Willems et al, 2005 | Flatworm | Gandalf | [92] | |
Litoria sauroni Richards & Oliver, 2006 | Frog | Sauron | "From the character Sauron, referred to as 'the eye' in Tolkien’s 'The Lord of the Rings', in reference to the striking red and black mottled eye of this taxon." | [93] |
Gandalfus McLay, 2007 | Crab | Gandalf | "Gandalfus is derived from the name of the omnipotent 'Gandalf', a character in J. R. Tolkien's 'Lord of the Rings', which was made into a film by Peter Jackson, in New Zealand, the home of the type species." | [94] |
Khamul gothmogi Gates, 2008 | Wasp | Gothmog, Lord of the Balrogs | "named in honor of Gothmog, Lord of the Balrogs, High Captain of Angband" | [95] |
Liolaemus tulkas Quinteros et al, 2008 | Lizard | Tulkas | "In the mythology of J. R.R. Tolkien, 'Tulkas' is one of the ainur or powers that helped shape arda or middle earth. One of the characteristics of Tulkas is that of running faster than any other creature. Liolaemus tulkas is very fast in short sprints." | [96] |
Sauroniops † Cau et al, 2012 | Dinosaur | Sauron | "The genus name is formed by Sauron, fictional character created by J.R.R. Tolkien (1892—1973), and όψη, Greek, 'eye'." | [97] |
Yavanna † Vera, 2013 | Tree fern | Yavanna | "The generic name is proposed after Yavanna, fictional character of The Silmarillion written by J.R.R. Tolkien. In Tolkien’s writings, Yavanna is a godlike entity (an Ainur), creator of the plants and animals that inhabit the Middle Earth." | [98] |
Anthracosuchus balrogus † Hastings et al, 2014 | Crocodyliform | Balrogs | "The specific term, balrogus, is named from a literary beast discovered within a deep mine, originally written by Tolkien" | [99] |
Terataner balrog Hita García, 2017 | Ant | Balrogs | For the animal's "dark" lifestyle as a predator, and for its robustly armoured and horned body | [100] |
Lepanus sauroni Gunter & Weir, 2019 | Dung beetle | Sauron | "This species is named sauroni for the distinctive pygidial depression with elongate turbercle that resembles the Eye of Sauron from Peter Jackson’s film adaptation of The Lord of the Rings." | [101] |
Baalrog Monjaraz-Ruedas, Prendini & Francke, 2019 | Schizomid | Balrogs | "The genus name is a compound word derived from two different words. Baal is a Mayan [sic, actually Hebrew] word for "devil." "Balrogs" are fictitious demons from the "Legendarium" and "The Lord of the Rings" by J.R.R. Tolkien." | [102] |
Orome † Bauzá, Gelfo & López, 2019 | Fossil mammal | Oromë | A genus of fossil Notoungulates from the Eocene of Argentina. | [103] |
Chespiritos balrogiformis Kuwahara & Marshall, 2020 | Fly | Balrogs | "This name refers to the fifth sternite, which resembles the head of the Balrog from J.R.R. Tolkien’s The Lord of The Rings series, specifically the one represented in Peter Jackson's movie Lord of the Rings: The Fellowship of the Ring." | [104] |
Afrodrassex balrog Haddad & Booysen, 2022 | Ground spider | Balrogs | "In Peter Jackson’s movies based on the books, the Balrog is depicted as wielding a long whip of fire, reminiscent of the very long embolus of this species." | [105] |
Austrosphecodes balrog Gonçalves & Pereira, 2022 | Bee | Balrogs | "A tall and menacing being who can shroud itself in fire, darkness, and shadow, created by J. R. R. Tolkien." | [106] |
Saurona Huertas & Willmott, 2023 | Butterfly | Sauron | "The generic name is based on that of the main antagonist, Sauron, in J. R. R. Tolkien’s novel ‘The Lord of the Rings’. The name alludes to the distinctive fused orange rings that encircle the VHW ocelli." | [107] |
Myloplus sauron Pereira, Ota, Machado, Collins, Ândrade, Garcia-Ayala, Jégu, Farias & Hrbek, 2024 | Fish | Sauron | "The specific name sauron alludes to the Eye of Sauron, from J. R. R. Tolkien’s “The Lord of the Rings”. The elliptical body of Myloplus sauron, marked with a vertical, black bar tapering toward both ends, resembles the famous vertical-pupilled eye from the novel." | [108] |
Elves
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Gildoria Hedqvist, 1974 | Wasp | Gildor Inglorion | "Named after an eIf-leader in Tolkien: 'The Lord of the Rings'." | [109] |
Legolasia Hedqvist, 1974 | Wasp | Legolas | Now revised to Chlorocytus. | [110] |
Oxyprimus galadrielae † Van Valen, 1978 | Fossil mammal | Galadriel | "Galadriel- (Sindarin [Elvish], radiantly garlanded woman), wise elf-queen of The Lord of the Rings and The Silmarillion." | [70] |
Tinuviel † Van Valen, 1978 | Fossil mammal | Lúthien (Tinúviel) | "Sindarin [Elvish] tinúviel, daughter of twilight, or nightingale, Beren's name for Lúthien in The Silmarillion'" | |
Metapheretima elrondi Easton, 1979 | Earthworm | Elrond | [111] | |
Feanora De Clerck & Schockaert, 1995 | Flatworm | Fëanor | "the genus name is a derivation of a mythological personage from Tolkien's 'Silmarillion'" | [112] |
Elachista lomionella Kaila, 1997 | Moth | Maeglin (Lómion) | [113] | |
Elachista amrodella Kaila, 1999 | Moth | Amrod | [72] | |
Elachista aredhella Kaila, 1999 | Moth | Aredhel | ||
Elachista caranthirella Kaila, 1999 | Moth | Caranthir | ||
Elachista celegormella Kaila, 1999 | Moth | Celegorm | ||
Elachista curufinella Kaila, 1999 | Moth | Curufin | ||
Elachista daeronella Kaila, 1999 | Moth | Daeron | ||
Elachista finarfinella Kaila, 1999 | Moth | Finarfin | ||
Elachista galadella Kaila, 1999 | Moth | Galadhrim, the Silvan Elves | ||
Elachista gildorella Kaila, 1999 | Moth | Gildor Inglorion | ||
Elachista guilinella Kaila, 1999 | Moth | Guilin | ||
Elachista indisella Kaila, 1999 | Moth | Indis | ||
Elachista maglorella Kaila, 1999 | Moth | Maglor | ||
Elachista miriella Kaila, 1999 | Moth | Míriel Serindë | ||
Elachista serindella Kaila, 1999 | Moth | Míriel Serindë | ||
Elachista telerella Kaila, 1999 | Moth | Teleri | ||
Elachista turgonella Kaila, 1999 | Moth | Turgon | ||
Gallogramma galadrieli † Garrouste et al, 2017 | Prehistoric insect | Galadriel | [114] | |
Pseudophallus galadrielae Dallevo-Gomes, Mattox, & Toledo-Piza, 2020 | Pipefish | Galadriel | "The epithet galadrielae refers to the character Galadriel in the trilogy 'The Lord of the Rings' by J. R. R. Tolkien. The elf ruler of Lothlórien is bearer of the ring Nenya, also known as the ring of water. It is used herein in reference to the additional bony rings diagnostic of the new species and its association with freshwater habitats." | [115] |
Dwarves
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Balinia Hedqvist, 1978 | Wasp | Balin | "named after Balin, a dwarf in Tolkien's 'The Lord of the Rings'." | [117] |
Gimlia Hedqvist, 1978 | Wasp | Gimli | "named after Gimli, a dwarf in Tolkien's book 'The Lord of the Rings'."
Now revised to Allocerastichus.[118] | |
Oinia Hedqvist, 1978 | Wasp | Óin | "named after Oin, a dwarf in Tolkien's book 'The Lord of the Rings'." | |
Bomburodon † (Van Valen, 1978) | Fossil mammal | Bombur | "Bombur, a fat dwarf in The Hobbit and The Lord of the Rings. Reference is to size and morphology." | [70] |
Deltatherium durini † Van Valen, 1978 | Fossil mammal | Durin | "Name of many dwarf-kings in The Lord of the Rings; Durin I beganKhazad-dûm. Allusion is to size." | |
Metapheretima kilii Easton, 1979 | Earthworm | Kíli | [111] | |
Metapheretima dorii Easton, 1979 | Earthworm | Dori | ||
Geocharidius gimlii Erwin, 1982 | Ground beetle | Gimli | "Gimlii, after the dwarf Gimli, son of Gloin (one of the 12 companions of Thorin Oakenshield), who accompanied the Hobbit, Frodo, on his trip south, in Book I of The Lord of the Rings by Tolkien, in reference to the small size of these beetles." | [121] |
Elachista ibunella Kaila, 1999 | Moth | Ibûn | [72] | |
Elachista telcharella Kaila, 1999 | Moth | Telchar | ||
Cacosternum thorini Conradie, 2014 | Frog | Thorin Oakenshield | "The species name is derived from Thorin II Oakenshield, a fictional character in the J.R.R. Tolkien novel ‘The Hobbit’. [...] It is widely known and accepted that Hogsback was the inspiration behind J.R.R. Tolkien's novels ‘The Hobbit’ and the ‘Lord of the Rings’ series. The name references its small size, its complicated call and the fact that it is found at the base of a large mountain in Hogsback." | [122] |
Geocharidius balini Sokolov & Kavanaugh, 2014 | Ground beetle | Balin | The specific epithet is "based on the given name of the dwarf Balin, a refounder of the underground kingdom of Moria, one of Thorin Oakenshield's Company of Dwarves who had accompanied Bilbo Baggins on the Quest of Erebor" | [123] |
Gervasiella oakenshieldi Paladini & Cavichioli, 2015 | True bug | Thorin Oakenshield | Named for the "fictional character surname of the novel The Hobbit, Thorin Oakenshield" | [124] |
Nebela gimlii Singer et al, 2015 | Testate amoeba | Gimli | "The name of this species refers to the name of Gimli, one of the dwarfs in J.R.R. Tolkien’s masterpiece 'The Lord of the Rings', because of its small size (the smallest known member of the Nebela collaris complex) and stout shape. In addition, it has been found abundantly in a forest, and Gimli was unique among his kind to have been travelling in the woods." | [125] |
Aspidoras azaghal Tencatt et al, 2020 | Catfish | Azaghâl | "Azaghâl was the king of the Broadbeam Dwarves, one of the seven dwarf clans, and Lord of the dwarven realm of Belegost in the Blue Mountains during Middle Earth's First Age. The name comes from a double allusion, first about the region where the species was found, Terra do Meio, freely translated as 'Middle Earth' in English, [the] name of the fictional world of Tolkien's legendarium, and second by the fact that the new species occurs in a mountainous region and presents a relatively small size, which are both typical features of the fictional dwarves." | [126] |
Spaeleoleptes gimli Pereira et al., 2024 | Harvestman | Gimli | "Refers to the dwarf Gimli, one of the main characters from J.R.R. Tolkien’s famous novel 'The Lord of the Rings'. This name was chosen because in Tolkien’s novel the dwarves are a race associated with mining and have a strong admiration for caves. Particularly, Gimli explicitly expresses his fascination for the Glittering Caves of Aglarond with their extensive series of spectacular speleothems, and after the defeat of Sauron he was given the lordship over this marvelous cave. The intention of the specific name is to make a metaphorical association with the close relationship of this species with its subterranean habitat." | [127] |
Race of Men
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Anisonchus eowynae † Van Valen, 1978 | Fossil mammal | Éowyn | "Éowyn, woman of Rohan in The Lord of the Rings, who killed the chief of the Nazgûl and was cured of his poison by athelas.
Now revised to Anisonchus athelas.[128] |
[70] |
Paradzickia morwen Blagoderov, 1998 | Fungus gnat | Morwen | "After the character in the books of J.R.R. Tolkien." | [129] |
Paradzickia hador Blagoderov, 1998 | Fungus gnat | Hador | "After the character in the books of J.R.R. Tolkien." | |
Elachista aerinella Kaila, 1999 | Moth | Aerin | [72] | |
Elachista arthadella Kaila, 1999 | Moth | Arthad | ||
Elachista beorella Kaila, 1999 | Moth | Bëor | ||
Elachista bregorella Kaila, 1999 | Moth | Bregor | ||
Elachista dagnirella Kaila, 1999 | Moth | Dagnir | ||
Elachista eilinella Kaila, 1999 | Moth | Eilinel | ||
Elachista gorlimella Kaila, 1999 | Moth | Gorlim | ||
Elachista haldarella Kaila, 1999 | Moth | Haldar | ||
Elachista marachella Kaila, 1999 | Moth | Marach | ||
Elachista morwenella Kaila, 1999 | Moth | Morwen | ||
Elachista neithanella Kaila, 1999 | Moth | Túrin Turambar (Neithan) | Neithan is a pseudonym of Túrin.[25] | |
Elachista nienorella Kaila, 1999 | Moth | Niënor | ||
Elachista ragnorella Kaila, 1999 | Moth | Ragnor | ||
Elachista rianella Kaila, 1999 | Moth | Rían | ||
Elachista tuorella Kaila, 1999 | Moth | Tuor | ||
Elachista turinella Kaila, 1999 | Moth | Túrin Turambar | ||
Idiopyrgus eowynae Salvador & Bichuette, 2024 | Snail | Éowyn | "Éowyn exemplifies courage, resilience, and resistance against darkness, both internal and external, standing against Gríma Wormtongue and the Witch-king of Angmar." | [130] |
Hobbits
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Gollum Compagno, 1973 | Shark | Gollum | [131] | |
Smeagolia (Hedqvist, 1973) | Wasp | Gollum (Sméagol) | Now revised to Muscidifurax. | [132] |
Pericompsus bilbo Erwin, 1974 | Beetle | Bilbo Baggins | "These beetles are short and robust much like Bilbo." | [133] |
Gollumiella Hedqvist, 1978 | Wasp | Gollum | [134] | |
Smeagol Climo, 1980 | Gastropod | Gollum (Sméagol) | [25][135] | |
Syconycteris hobbit Ziegler, 1982 | Bat | Hobbits | [25][136] | |
Paragiopagurus hobbiti (Macpherson, 1983) | Hermit crab | Hobbits | "The name hobbiti is given in allusion to the famous fictional race of small creatures created by J. R. Tolkien (hobbits) which, like the new species, dwell in burrows." | [137] |
Gollumia Riedel, 1988 | Snail | Gollum | "Nach Gollum (oder Smeagol) - ein Fabelwesen, Ausgeburt von J. R. R. TOLKIEN, geheimnisvolle und rätselhafte Kreatur, die in Dunkelheit, am liebsten unterirdisch lebte." | [138] |
Macrostyphlus bilbo Morrone, 1994 | Beetle | Bilbo Baggins | [87] | |
Macrostyphlus frodo Morrone, 1994 | Beetle | Frodo Baggins | ||
Marjumia bagginsi † Melzak & Westrop, 1994 | Trilobite | Bilbo Baggins | [139] | |
Iandumoema smeagol Pinto-da-Rocha, 1996 | Harvestman | Gollum (Sméagol) | [140] | |
Lotharingius frodoi † Mattioli, 1996 | Coccolithophore | Frodo Baggins | [141] | |
Frodospira † Wagner, 1999 | Gastropod | Frodo Baggins | [142] | |
Galaxias gollumoides McDowall & Chadderton, 1999 | Ray-finned fish | Gollum | Gollumoides means "Gollum-like".[25] | [143] |
Breviceps bagginsi Minter, 2003 | Frog | Bilbo Baggins | [144] | |
Peperomia hobbitoides Wendt | Piperaceae | Hobbits | "Peperomia hobbitoides is a small and humble plant that lives in an almost fairyland-like environment of wet karst outcrops in rain forest, and it is strongly and faithfully tied to this home substrate. Indeed, it spends perhaps the greater part of the year in holes and depressions in the rock as a resting tuber. It is edible, an attribute of high esteem among hobbits." | [145] |
Gollumjapyx smeagol Sendra & Ortuño, 2006 | Dipluran | Gollum (Sméagol) | [146] | |
Ingerophrynus gollum Grismer, 2007 | Toad | Gollum | [147] | |
Laparocerus hobbit Machado, 2008 | Beetle | Hobbits | "The specific epithet ... refers to the Hobbits ... a literary fictitious race of people who have big and hairy feet; a metaphor of the swollen and hairy tarsi characteristic of this species." | [148] |
Saurodocus hobbit Yerman & Krapp-Schickel, 2008 | Amphipod | Hobbits | "Named after the fictional small people of 'halflings' in the fantasy novels written by J. R. R. Tolkien in the traditions of a fairy tale" | [149] |
Abacophrastus hobbit Will, 2011 | Beetle | Hobbits | "an allusion to the setose dorsal surface of the tarsi, analogous to the hairy feet of Tolkien’s Hobbits" | [150] |
Shireplitis bilboi Fernández-Triana & Ward, 2013 | Wasp | Bilbo Baggins | [80] | |
Shireplitis frodoi Fernández-Triana & Ward, 2013 | Wasp | Frodo Baggins | ||
Shireplitis meriadoci Fernández-Triana & Ward, 2013 | Wasp | Meriadoc Brandybuck | ||
Shireplitis peregrini Fernández-Triana & Ward, 2013 | Wasp | Peregrin Took | ||
Shireplitis samwisei Fernández-Triana & Ward, 2013 | Wasp | Samwise Gamgee | ||
Tennesseellum gollum Dupérré, 2013 | Spider | Gollum | Named "on reference to the evil persona Gollum, given the 'evil look' of the male of this species due to the large cheliceral tubercles." | [151] |
Tetramorium gollum Hita Garcia & Fisher, 2014 | Ant | Gollum | "The new species is named after the fictional character "Gollum" from J.R.R. Tolkien's novels 'The Hobbit' and 'The Lord of the Rings'." | [152] |
Tetramorium hobbit Hita Garcia & Fisher, 2014 | Ant | Hobbits | "This very hairy new species is named after the fictional people from J.R.R. Tolkien’s novels 'The Hobbit' and 'The Lord of the Rings'." | |
Ansonia smeagol Davies et al, 2016 | Toad | Gollum (Sméagol) | [153] | |
Meoneura hobbitoides (Stucke, 2016) | Fly | Hobbits | [154] | |
Aglaophenia baggins Soto Ángel & Peña Cantero, 2017 | Hydrozoan | Baggins family | [155] | |
Meoneura bilboi Stuke & Freidberg, 2017 | Fly | Bilbo Baggins | A tiny fly (length<2 mm) "dedicated to Bilbo Baggins [...] who, being an ordinary hobbit, found the ring of power in J.R.R. Tolkienʼs fantasy novel The Hobbit." | [156] |
Odontonia bagginsi de Gier & Fransen, 2018 | Shrimp | Baggins family | [157] | |
Aenigmachanna gollum Britz et al, 2019 | Labyrinth fish | Gollum | "Named after Gollum, a character from J. R. R. Tolkien’s books ‘The Hobbit’ and ‘The Lord of the Rings’, a creature that went underground and during its subterranean life changed its morphological features." | [158] |
Psylla frodobagginsi Martoni, 2019 | Psyllid | Frodo Baggins | [159] | |
Goniurosaurus gollum Qi et al, 2020 | Gecko | Gollum | "This new species and Gollum have similar cave-dwelling habit and emaciated body. We suggest the common name as 'Gollum Leopard Gecko'" | [160] |
Sinopesa gollum Lin & Li, 2021 | Spider | Gollum | "named after Gollum,[...] who lived in a cave, as does this new species" | [161] |
Chiloglanis frodobagginsi Schmidt, Friel, Bart & Pezold, 2023 | Catfish | Frodo Baggins | "Chiloglanis frodobagginsi is named after another diminutive traveller, Frodo Baggins" | [162] |
Idiopyrgus meriadoci Salvador & Bichuette, 2024 | Snail | Merry Brandybuck | "Besides standing with Éowyn against the Witch-king in the Battle of the Pelennor Fields, Merry is also an example of the fight for nature conservation in Middle-earth, pushing the Ents into action and ultimately ending Saruman's threat to Fangorn Forest." | [130] |
Orcs
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Borophagus orc † Webb, 1969 | Fossil mammal | Orcs | "Orc = a ruthless carnivorous creature serving the forces of evil in the Third Age of Middle Earth" | [163] |
Protungulatum gorgun † Van Valen, 1978 | Fossil mammal | Orcs (Gorgûn) | "Etymology: Gorgûn, the Woses' name for orcs in The Lord of the Rings, with reference to the Cretaceous Hell Creek Formation" | [70] |
The Nazgûl
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Nazgulia Hedqvist, 1973 | Wasp | Nazgûl | [132] | |
Khamul Gates, 2008 | Wasp | Khamûl, a Nazgûl | [79] | |
Tetramorium nazgul Hita Garcia & Fisher, 2012 | Ant | Nazgûl | [164] | |
Acledra nazgul Faúndez, Rider & Carvajal, 2016 | True bug | Nazgûl | "After the fictional Nazgûl characters created by J.R.R. Tolkien, who, mounted on winged creatures, could fly long distances, even on steep peaks; on the other hand, this new species has a wide distribution and dispersal capacity throughout the highlands near the Argentinean Andes, which resembles the behaviour of the Nazgûl." | [165][166] |
Abavorana nazgul Quah et al, 2017 | Frog | Nazgûl | Named "in reference to the 'Nazgûl', characters created by J.R.R. Tolkien in The Lord of the Rings (1954). Also known as Ring-wraiths, they were nine men who succumbed to Sauron’s power and were transformed into white ghostly figures wearing black cloaks—the colouration that the new population being described herein shares." | [167] |
Potamalpheops nazgul Christodoulou, Iliffe & De Grave, 2019 | True shrimp | Nazgûl | "Named after J.R.R. Tolkienʼs fictional characters, the Nazgul, who dwell in the realm of shadows, akin to the habitat of this new shrimp species" | [168] |
Mischocyttarus nazgul Borges & Silveira, 2019 | Wasp | Nazgûl | "The specific epithet is a reference to the Nazgul Kings" | [169] |
Ents
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Entia Hedqvist (1974) | Wasp | Ents | Now revised to Boucekastichus. | [170][171] |
Fimbrethil † Van Valen, 1978 | Fossil mammal | Fimbrethil, the lost wife of Treebeard | "Fimbrethil, entwife loved by Fangorn in The Lord of the Rings. Reference is to partly primate-like morphology and the disappearance of both Finbrethils." | [70] |
Unicauda fimbrethilae Rosser, 2016 | Myxozoan | Fimbrethil, the lost wife of Treebeard | [173] |
Other characters
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Beorn † Cooper, 1964 | Tardigrade | Beorn | Tardigrades or water bears are microscopic animals that look something like bears. | [174] |
Beornia Hedqvist, 1975 | Wasp | Beorn | [175] | |
Earendil † Van Valen, 1978 | Fossil mammal | Eärendil, a Half-elf | "Eärendil, who (in The Silmarillion) sailed with a silmaril to get the aid that defeated Morgoth." | [70] |
Mimotricentes mirielae † Van Valen, 1978 | Fossil mammal | Tar-Míriel, a queen of Númenor | "Míriel (Quenya, Jewel-woman), Númenorian queen in The Silmarillion, forced into marriage and the loss of her throne."
Now revised to Loxolophus hyattianus.[176] | |
Protoselene bombadili † (Van Valen, 1978) | Fossil mammal | Tom Bombadil | "Tom Bombadil, the Hobbit name for a simple, powerful, and very old being. Reference is to these three traits." | |
Elachista diorella Kaila, 1999 | Moth | Dior Eluchíl, a Half-elf | [72] |
Animals
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Ancalagon Conway Morris, 1977 | Priapulid | Ancalagon the Black | [177] | |
Ankalagon † (Van Valen, 1978) | Fossil mammal | Ancalagon the Black | "The mightiest dragon of Morgoth, in The Silmarillion." | [70] |
Claenodon mumak † Van Valen, 1978 | Fossil mammal | Mûmakil, the elephants of Middle-earth | "Mûmakil name used in Ithilien for the animal hobbits called an oliphaunt, resembling a large elephant. Reference is to size." | |
Desmatoclaenus mearae † Van Valen, 1978 | Fossil mammal | Mearas, the horses of Rohan | "Meara, any one of the great horses of Rohan in The Lord of the Rings." | |
Gwaihiria Naumann, 1982 | Wasp | Gwaihir, Lord of the Eagles | [178] | |
Amphiledorus ungoliantae Pekár & Cardoso, 2005 | Spider | Ungoliant | Ungoliant is a giant spider. | [179] |
Nemesia ungoliant Decae, Cardoso & Selden, 2007 | Spider | Ungoliant | [180] | |
Liolaemus smaug Abdala et al, 2010 | Lizard | Smaug | "In Tolkien’s mythology Smaug, the Golden, is the last of the Middle Earth dragons. The name Liolaemus smaug is because this new species exhibit a golden coloration on body." | [181] |
Smaug Stanley et al, 2011 | Lizard | Smaug | [182] | |
Tetramorium smaug Hita Garcia & Fisher, 2012 | Ant | Smaug | [164] | |
Pycnophyes smaug Sánchez et al, 2013 | Kinorhynch | Smaug | "The species name smaug, refers to the dragon Smaug, the greatest and most powerful in the later part of the Third Age in the books of J.R.R. Tolkien." | [183] |
Glaurung † Bulanov & Sennikov, 2015 | Weigeltisaurid | Glaurung | [184] | |
Planois smaug Carvajal, Faúndez & Rider, 2015 | True bug | Smaug | [185][186] | |
Tamolia ancalagon Carvajal et al, 2015 | True bug | Ancalagon the Black | Named "for Ancalagon the Black, the largest dragon in J. R. R. Tokien’s universe; because of the dark coloration and aspect of this new species, as well as its size compared to many other heteropterans." | [187] |
Cristaphyes glaurung Sørensen & Grzelak, 2018 | Kinorhynch | Glaurung | [188] | |
Cristaphyes scatha Sørensen & Grzelak, 2018 | Kinorhynch | Scatha the Worm | ||
Ochyrocera laracna Brescovit, Cizauskas & Mota, 2018 | Spider | Shelob | "Laracna" is Shelob's name in Portuguese | [189] |
Ochyrocera ungoliant Brescovit, Cizauskas & Mota, 2018 | Spider | Ungoliant | ||
Pycnophyes ancalagon Sørensen & Grzelak, 2018 | Kinorhynch | Ancalagon the Black | [188] | |
Rhabdias glaurungi Willkens et al., 2020 | Roundworm | Glaurung | This species is named after the fictional character ‘Glaurung’, the first of the Dragons from ‘The Silmarillion’ | [190] |
Cnemaspis smaug Pal et al, 2021 | Gecko | Smaug | "The species is named after "Smaug", the dragon from J. R. R. Tolkien's 1937 novel, The Hobbit. The name is derived from the old German verb 'smeuganan' meaning "to creep" or "to squeeze through a hole". The type specimens of this species were found within crevices of rocks and boulders inside the forest. Additionally, like dragons, the dorsum is armoured with large conical tubercles." | [191] |
Objects and locations
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Anisonchus athelas † Van Valen, 1978 | Fossil mammal | Athelas, a plant | "Sindarin (Elvish) athelas, kingsfoil, a healing plant in The Lord of the Rings. Reference is to the joining of phylogenies." | [70] |
Earendil undomiel † Van Valen, 1978 | Fossil mammal | Undómiel | "Quenya (Elvish) undómiel, evening star, which Eärendil with his silmaril became." | |
Fimbrethil ambaronae † Van Valen, 1978 | Fossil mammal | Fangorn (also named Ambaróna) | "Quenya (Elvish) Ambaróna, one of Fangorn's shorter names for his forest. Reference is to the dimness of the forest and of the affinities of this species."
Now revised to Oxyacodon agapetillus.[192] | |
Litaletes ondolinde † Van Valen, 1978 | Fossil mammal | Gondolin (also named Ondolindë) | "Quenya (elvish) ondo, rock, and lindë, song. Reference is to Rock Bench and to the hidden city Ondolindë or Gondolin of The Silmarillion. The Rock Bench specimens and others were formerly as hidden (and unsorted)." | |
Niphredil † Van Valen, 1978 | Fossil mammal | Niphredil, a flower | "Sindarin (Elvish) niphredil, white-flowering forb of open woods in Neldoreth and Lothlorien." | |
Platymastus palantir † Van Valen, 1978 | Fossil mammal | Palantir | "Quenya (Elvish) palantir, distant watcher, one of 7 globes made by Fëanor that gave visions through spacetime. Reference is to the long duration of the genus." | |
Thangorodrim † Van Valen, 1978 | Fossil mammal | Thangorodrim | "Thangorodrim, the mountainous triple fortress of Morgoth in The Silmarillion. Reference is to Purgatory Hill."
Now revised to Oxyclaenus.[193] | |
Metapheretima andurili Easton, 1979 | Earthworm | Andúril, sword reforged by Aragorn | [111] | |
Metapheretima stingi Easton, 1979 | Earthworm | Sting, Bilbo and Frodo's dagger | ||
Metapheretima orcrista Easton, 1979 | Earthworm | Orcrist, Thorin Oakenshield's sword | ||
Metapheretima glamdringi Easton, 1979 | Earthworm | Glamdring, Gandalf's sword | ||
Pseudopallenis palantir Kolibáč, 1997 | Beetle | Palantir | [194] | |
Asthenodipsas lasgalenensis Loredo et al, 2013 | Snake | Mirkwood (also named Eryn Lasgalen) | "The specific epithet lasgalenensis is derived from the name Eryn Lasgalen which means in the "Wood of Greenleaves" in the fictional Sindarian language from J.R.R Tolkien's The Lord of the Rings (1955). It was the name used by the Wood Elves for the Mirkwood Forest after its cleansing following the War of the Ring. This name was chosen because Tolkien's (1955) description of this forest showed great similarity to the cloudy, upland forests within which this species is found." | [195] |
Shireplitis Fernández-Triana & Ward, 2013 | Wasp | The Shire | "[T]he genus is endemic to New Zealand, where a human replicate of The Shire is built". | [80] |
Lopholatilus ereborensis † Carnevale & Godfrey, 2014 | Tilefish | Erebor, the Lonely Mountain | [151] | |
Desmia mordor Landry & Solis, 2016 | Moth | Mordor | The specific name "means ‘Black Land’ in Sindarin, a fictional language used in The Lord of the Rings, the epic high-fantasy novel written by English author J. R. R. Tolkien [...] Mordor is volcanic and partly arid, like the Galápagos." | [196] |
Arctoceras erebori † Piazza, 2017 | Ammonoid | Erebor, the Lonely Mountain | [197] | |
Astyanax lorien Zanata, Burger & Camelier, 2018 | Ray-finned fish | Lothlórien | lorien, from the Quenya language meaning "Dream Land", used in allusion to the "beautiful areas" inhabited by the Brazilian species | [198] |
Acantopsis bruinen Boyd et al., 2018 | Ray-finned fish | River Bruinen, "Loudwater" | Specific epithet bruinen for the Loudwater of Rivendell and the flood that took the form of great horses, alluding to the common name “horseface loach” for the genus | [199] |
Cristaphyes dordaidelosensis Sørensen & Grzelak, 2018 | Kinorhynch | Dor Daidelos | "The species name dordaidelosensis, meaning 'living in Dor Daidelos,' is inspired by the book Silmarillion by JRR Tolkien. According to the book, Dor Daidelos is 'The Region of Everlasting Cold' and the northernmost region of Middle Earth in the First Age." | [188] |
Uroplatus fangorn Ratsoavina et al, Scherz, 2020 | Gecko | Fangorn | "The species epithet fangorn is [...] derived from the name of a deep, dark woodland in J.R.R. Tolkien’s Middle Earth. [...] We use this name in reference not only to the similarities between Fangorn forest and the forests in which the new species occurs, but also to the tree-like appearance of Uroplatus geckos, which make them seem like the 'tree spirits' that are Tolkien’s Ents (Flieger 2013)." | [200] |
Hyperlais orodruinella Korb, Gorbunov & Melyakh, 2023 | Moth | Orodruin (Mount Doom) | "We name the new species after Mount Orodruin. In J.R.R. Tolkien’s The Lord of the Rings, the One Ring was forged on Mount Orodruin by the Dark Lord Sauron. The name shows the characteristic feature of the moth: black triangular wings with a yellow suffusion in the basal part, resembling a dark volcano with an erupting top" | [201] |
Elvish words
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Aletodon mellon † Van Valen, 1978 | Fossil mammal | mellon | "Sindarin (Elvish) mellon, friend, the password of the west gate of Khazad-dûm in The Lord of the Rings. Reference is to similarly to P. palantir, presumptive diet of plants" | [70] |
Chriacus calenancus † Van Valen, 1978 | Fossil mammal | calen, anca | "Sindarin (Elvish) calen, green; anca, Jaws. Reference is to inferred herbivory" | |
Litomylus alphamon † Van Valen 1978 | Fossil mammal | alph, amon | "Sindarin (Elvish) alph, swan, and amon, hill. Reference is to the locality"
Now revised to Litocherus lacunatus.[202] | |
Mimatuta minuial † Van Valen, 1978 | Fossil mammal | minuial | "Sindarin (Elvish) minuial, the time at dawn when the stars fade. Reference is to the dawn of the Cenozoic and the fading of the Mesozoic stars." | |
Thangorodrim thalion † Van Valen, 1978 | Fossil mammal | thalion | "Sindarin (Elvish) thalion, strong. Reference is to the massive morphology of the jaws and the generic name"
Now revised to Oxyclaenus pugnax.[203] | |
Elachista aranella Kaila, 1999 | Moth | aran- | "E. aranella seems to owe its name to aran-, the 'royal prefix used by the Kings of Arthedain after Malvegil and by the Chieftains of the Dúnedain of the North to indicate their claim to all of Arnor'".[204] | [72] |
Helicops nentur Costa et al, 2016 | Snake | nen, tur | "The name nentur is formed by the (Quenya) words nen (water) and tur (ruler, master)," referencing the aquatic habits of the species | [205] |
Hylaeus mellon Dathe and Proshchalykin, 2016 | Bee | mellon | "mellon (High Elvish [Sindarin]): friend" | [206] |
Epimeria anguloce d’Acoz & Verheye, 2017 | Amphipod | angulócë | "Angulócë, dragon — Tolkien’s Quenya language ( Faulskanger 2008). The name, which is a noun in apposition, alludes to the dragon-like facies of the species." | [207] |
Thiodina perian Bustamante & Ruiz, 2017 | Jumping spider | perian | "The specific name is a noun in apposition taken from the fictional language Sindarin created by J.R.R. Tolkien. The Sindarin is one of the languages spoken by the Elves. The word 'perian' is translated as 'small' and was used to refer to the Hobbits, the smaller kind of Middle Earth; in reference to its small size, to date the smallest thiodinine ever discovered" | [208] |
Vanima Zacca, Casagrande & Mielke, 2020 | Butterfly | vanima | In Quenya (elvish) vanima means beautiful | [209] |
Other works
[edit]Taxon | Type | Named for | Notes | Ref |
---|---|---|---|---|
Aspidoras psammatides Britto, Lima & Santos, 2005 | Catfish | Psamathos Psamathides, Roverandom | "Psammatides, after 'Psammatos psammatides' [sic], 'the sand sorcerer', a character of J.R.R. Tolkien’s book 'Roverandom', from the Greek psammos, sand, and ides, son of. In allusion to the sand-dwelling behavior of the species." | [210] |
See also
[edit]- Reception of J. R. R. Tolkien
- Works inspired by J. R. R. Tolkien
- List of organisms named after works of fiction
Notes
[edit]References
[edit]- ^ "(2991) Bilbo=1982 HV=1982 KB2=1975 JC=1979 SY3". IAU Minor Planet Center. Retrieved 10 March 2016.
- ^ "JPL Small-Body Database Browser: 2675 Tolkien (1982 GB)". Jet Propulsion Laboratory. Retrieved 10 March 2016.
- ^ "JPL Small-Body Database Browser: 385446 Manwe (2003 QW111)" (2013-11-20 last obs.). Jet Propulsion Laboratory. Retrieved 16 March 2017.
- ^ Johnston, Wm. Robert (20 September 2014). "(385446) Manwe and Thorondor". Asteroids with Satellites Database—Johnston's Archive. Retrieved 1 August 2018.
- ^ Gohd, Chelsea (2022-03-31). "Meet Earendel: Hubble telescope's distant star discovery gets a Tolkien-inspired name". Space.com. Retrieved 2022-03-31.
- ^ Anderson, Thomas (2 March 2013). "The Eye of Sauron is nebula ESO 456-67". Tech Guru Daily.
- ^ Hoffman, Mark (25 February 2013). "Another Eye Of Sauron Nebula". Science World Report.
- ^ "The Eye of Sauron (aka NGC7293)". Sky and Telescope.
- ^ Atkinson, Nancy (4 October 2012). "Eye-Like Helix Nebula Turns Blue in New Image". Universe Today. Archived from the original on 14 July 2014.
- ^ Graham, Flora (4 June 2014). "Eye of Sauron star spotted by planet-hunting camera". New Scientist.
- ^ NewsCorp Australia (6 June 2014). "Sauron's Eye has been found. And it's watching us all, from above, in the form of ringed star HR 4796A". Courier Mail.
- ^ Chandra X-ray Observatory (10 March 2011). "NGC 4151: An Active Black Hole in the "Eye of Sauron"". Chandra X-ray Center.
- ^ "378214 Sauron (2007 AP11)". JPL. Retrieved 9 September 2021.
- ^ "174567 Varda (2003 MW12)". Minor Planet Center. Retrieved 24 November 2019.
- ^ "Categories for Naming Features on Planets and Satellites". Gazetteer of Planetary Nomenclature. International Astronomical Union (IAU) Working Group for Planetary System Nomenclature (WGPSN). Retrieved 28 February 2016.
- ^ a b "Gazetteer of Planetary Nomenclature: Titan Colles". Planetarynames.wr.usgs.gov. Retrieved June 26, 2015.
- ^ "Gazetteer of Planetary Nomenclature: Titan Mons, Montes". Gazetteer of Planetary Nomenclature. International Astronomical Union (IAU) Working Group for Planetary System Nomenclature (WGPSN). Retrieved 28 February 2016.
- ^ Chang, Kenneth (14 July 2015). "NASA's New Horizons Spacecraft Completes Flyby of Pluto". The New York Times. Retrieved 14 July 2015.
- ^ Rogers, Adam (14 July 2015). "The New, Nerdy Mythology of Pluto's Place Names". Wired. Retrieved 14 July 2015.
- ^ Nemiroff, Robert; Bonnell, Jerry (17 July 2015). "APOD: 2015 July 17 – Charon". Astronomy Picture of the Day. Retrieved 17 July 2015.
- ^ Lendino, Jamie (15 July 2015). "NASA unveils historic, high-resolution Pluto, Charon, and Hydra photos from New Horizons mission". Extremetech.com. Retrieved 12 March 2016.
- ^ Stockton, Nick (2015). "Charon Comes into Focus in New Horizons' Latest Photos". Wired. Retrieved 12 March 2016.
- ^ Appelcline, Shannon (2011). Designers & Dragons. Mongoose Publishing. p. 133. ISBN 978-1-907702-58-7.
- ^ "About". Middle-earth Enterprises. Archived from the original on 2011-07-14.
- ^ a b c d e f g Larsen 2007
- ^ "Strategy". Lembas Capital. 2019. Archived from the original on 30 April 2019. Retrieved 29 June 2020.
- ^ "The Tolkien Trust - Extract from the Central Register of Charities maintained by the Charity Commission for England and Wales". Charity Commission. Archived from the original on 2009-02-19. Retrieved 2016-03-20.
- ^ "WWF-UK Annual Review 2007" (PDF). WWF-UK. Retrieved 21 June 2024.
- ^ "Home". Anduril Industries. Retrieved 2018-08-27.
- ^ "MITHRIL - A Social Network that rewards all content creators". mith.io. Retrieved 2018-08-27.
- ^ Thompson, Lenn (26 October 2007). "Rivendell Winery Location for Sale in the Hudson Valley". New York Cork Report. Retrieved 29 June 2020.
- ^ Burgess, J. Michael; Yu, Hoi-Fung; Greiner, Jochen; Mortlock, Daniel J. (11 May 2018). "Awakening the BALROG: BAyesian Location Reconstruction Of GRBs". Monthly Notices of the Royal Astronomical Society. 476 (2): 1427–1444. arXiv:1610.07385. Bibcode:2018MNRAS.476.1427B. doi:10.1093/mnras/stx2853.
- ^ https://www.durindrilling.com/
- ^ a b "What's in a name? The stories behind some of science's most curious names". www.phdnet.mpg.de. Retrieved 2022-09-28.
- ^ Cao, Wen Xi; Karaiskakis, Angelo; Lin, Sichun; Angers, Stephane; Lipshitz, Howard D (2022-01-04). Duronio, R (ed.). "The F-box protein Bard (CG14317) targets the Smaug RNA-binding protein for destruction during the Drosophila maternal-to-zygotic transition". Genetics. 220 (1): iyab177. doi:10.1093/genetics/iyab177. ISSN 1943-2631. PMC 8733446. PMID 34757425.
- ^ Gymnosperm Database
- ^ Staff, Jeff Brammer News-Post (26 August 2009). "CHRIS GROVE: Frederick trainer is a good story among an industry in need of one". The Frederick News-Post. Retrieved 2022-09-26.
- ^ "Mount Gandalf". BC Geographical Names. The Province of British Columbia. Retrieved 12 December 2014.
- ^ "Mount Shadowfax". BC Geographical Names. The Province of British Columbia. Retrieved 12 December 2014.
- ^ "Unnamed peak could be named Mt Tolkien". Stuff.co.nz. Fairfax New Zealand Limited. Retrieved 12 December 2012.
- ^ "Rockall: The lost continent of Middle Earth". Volcano Cafe. 2 June 2017. Retrieved 11 December 2021.
- ^ "Eriador Seamount". MarineRegions.org. Retrieved 11 December 2021.
- ^ "Rohan Seamount". Earthref.org. Retrieved 11 December 2021.
- ^ "Gondor Seamount". Earthref.org. Retrieved 11 December 2021.
- ^ "Fangorn Bank". MarineRegions.org. Retrieved 11 December 2021.
- ^ Barton, A. J.; White, R. S. (1995). "The Edoras Bank margin; continental break-up in the presence of a mantle plume". Journal of the Geological Society. 152 (6): 971–974. Bibcode:1995JGSoc.152..971B. doi:10.1144/gsl.jgs.1995.152.01.15. ISSN 0016-7649. S2CID 129853250.
- ^ "Isengard Ridge". MarineRegions.org. Retrieved 11 December 2021.
- ^ a b c d "An undersea volcano discovered near Christmas Island looks like the Eye of Sauron". Phys.org. Retrieved 11 December 2021.
- ^ "Biography | Amon Amarth". 2011-04-24. Archived from the original on 2011-04-24. Retrieved 2021-11-25.
- ^ Vikernes, Varg (December 2004). "A Burzum Story: Part I". burzum.org. Retrieved 25 May 2013.
- ^ Lee, Cosmo (13 March 2006). "Ephel Duath - Pain Necessary to Know - Review" Archived 2008-10-07 at the Wayback Machine. Stylus Magazine. Retrieved 15 December 2011.
- ^ "Gorgoroth". allmusic.com. Retrieved 9 September 2021.
- ^ "Iluvatar". allmusic.com. Retrieved 9 September 2021.
- ^ "Marillion". allmusic.com. Retrieved 9 September 2021.
- ^ Silveyra, Marcelo (2002). "Chapter 1 – Writing Down The Script". Progfreaks.com. Archived from the original on 2008-06-04.
- ^ "Shadowfax". allmusic.com. Retrieved 9 September 2021.
- ^ "The Yacht J R Tolkien". Charter World. 2012. Retrieved 20 January 2012.
- ^ Cordner, Chris (23 February 2018). "Tolkien about a wonderful ship with a fascinating past". Sunderland Echo.
She was also renamed in honour of the British author JRR Tolkien.
- ^ Butcher, Amanda (2014). Tall Ships Handbook. Amberley Publishing Limited. p. 72. ISBN 9781445618685.
- ^ "People of Stoke-on-Trent". Retrieved 13 March 2005.
- ^ a b "Nature reserve remembers Baggins". BBC News. Retrieved 10 March 2016.
- ^ "Hobbiton Road, Weston-Super-Mare". Google Maps. Retrieved 12 May 2023.
- ^ Strutner, Sizy. "Every Street In This Dutch Neighborhood Has A Lord Of The Rings Name, Which Is Awesome". The Huffington Post. Retrieved 26 January 2014.
- ^ "Tolkien i Gandalf będą mieć swoje ulice w Mordorze". Retrieved 23 December 2012.
- ^ Burslem, Joel (18 September 2006). "Hobbit Living in Bend". Inman. Retrieved 30 April 2021.
- ^ "Middle Earth". UCI Student Housing. Archived from the original on 29 April 2021. Retrieved 30 April 2021.
- ^ Phillips, Bethan (13 March 2012). "The Hobbit in battle with Hollywood". Southern Daily Echo. Retrieved 10 March 2016.
- ^ Beck, Julie (3 May 2015). "Science's Love Affair with The Lord of the Rings". The Atlantic. Retrieved 10 March 2016.
- ^ Gee 2004, p. 55
- ^ a b c d e f g h i j k Van Valen, L. M. (1978). "The beginning of the Age of Mammals" (PDF). Evolutionary Theory. 4: 45–80. Taxonomic summary
- ^ Hansson, Christer (2010). "Till minne av Karl-Johan Hedqvist" [In memory of Karl-Johan Hedqvist] (PDF). Entomologisk Tidskrift (in Swedish). 131: 137–144.
- ^ a b c d e f g Kaila, Lauri (1999). "A revision of the Nearctic species of the genus Elachista s.l. III. The bifasciella, praelineata, saccharella and freyerella groups (Lepidoptera, Elachistidae)". Acta Zoologica Fennica. 211: 1–235.
- ^ Aiello, Leslie C. (2010). "Five years of Homo floresiensis". American Journal of Physical Anthropology. 142 (2): 167–179. doi:10.1002/ajpa.21255. PMID 20229502.
- ^ Kennedy, George L. (1974). "West American Cenozoic Pholadidae (Mollusca: Bivalvia)". Memoirs of the San Diego Society of Natural History. 8 (59): 59–60.
- ^ Vinogradov, G.M. (1990). "Amphipoda (Crustacea) in the pelagic zone of the southeastern Pacific Ocean (in Russian)". Trudy Instituta Okeanologii Akademii Nauk SSSR. 124: 27–104.
- ^ Thomas, James Darwin (22 July 2009). "Leucothoidae species (version 7.09) [includes Anamixidae (sensu lato)]" (PDF). Nova Southeastern University. Archived from the original (PDF) on 19 March 2012. Retrieved 20 March 2016.
- ^ Schillhammer, H. (1997). "Taxonomic revision of the oriental species of Gabrius Stephens (Coleoptera: Staphylinidae)". Monographs on Coleoptera: 1–139.
- ^ Lieberman, B.S.; Kloc, G.J. (1997). "Evolutionary and biogeographical patterns in the Asteropyginae (Trilobita, Devonian) Delo, 1935". Bulletin of the American Museum of Natural History. 232. hdl:2246/1623.
- ^ a b Gates, M. W. (2008). "Description of Khamul, gen. n.(Hymenoptera: Chalcidoidea: Eurytomidae), with a hypothesis of its phylogenetic placement". Zootaxa (1898): 1–33.
- ^ a b c Fernandez-Triana, J; Ward, DF; Cardinal, S; Van Achterberg, C (2013). "A review of Paroplitis (Braconidae, Microgastrinae), and description of a new genus from New Zealand, Shireplitis, with convergent morphological traits". Zootaxa. 3722 (4): 549–568. doi:10.11646/zootaxa.3722.4.6. hdl:2292/21460. PMID 26171541.
- ^ Mbo, Zingisile; Haddad, Charles R. (2019-04-10). "A revision of the endemic South African long-jawed ground spider genus Drassodella Hewitt, 1916 (Araneae: Gallieniellidae)". Zootaxa. 4582 (1): zootaxa.4582.1.1. doi:10.11646/zootaxa.4582.1.1. ISSN 1175-5334. PMID 31716194. S2CID 145898564.
- ^ Sánchez-Nivicela, Juan C.; Falcón-Reibán, José M.; Cisneros-Heredia, Diego F. (19 January 2023). "A new stream treefrog of the genus Hyloscirtus (Amphibia, Hylidae) from the Río Negro-Sopladora National Park, Ecuador". ZooKeys (1141): 75–92. Bibcode:2023ZooK.1141...75S. doi:10.3897/zookeys.1141.90290. ISSN 1313-2970. PMC 10208809. PMID 37234964. Retrieved 3 March 2023.
- ^ Hamilton, K. G. Andrew (1972-12-01). "The Manitoban fauna of leafhoppers I. Descriptions of new species and colour forms". The Canadian Entomologist. 104: 825–831. doi:10.4039/Ent104825-6. S2CID 86516993.
- ^ Registry-Migration.Gbif.Org (2021), Macropsidius sauroni (Hamilton, 1972) in GBIF Secretariat (2021), GBIF Secretariat, doi:10.15468/39omei, retrieved 2022-09-25
- ^ Hedqvist, Karl-Johan (1977). "Two new genera and species of Pteromalidae from Brazil (Hymenoptera)". Insect Systematics & Evolution. 8 (1): 78–80. doi:10.1163/187631277X00134. ISSN 1399-560X.
- ^ Williams, G. A.; Weir, T. A. (1988). "Further New Species of Australian Mastogeniinae (Coleoptera: Buprestidae)". Australian Journal of Entomology. 27 (3): 179–181. doi:10.1111/j.1440-6055.1988.tb01518.x. ISSN 1326-6756.
- ^ a b Morrone, Juan J. (2011-12-05). "NAnnotated checklist of the tribe Listroderini (Coleoptera: Curculionidae: Cyclominae)". Zootaxa. 3119 (1): 1. doi:10.11646/zootaxa.3119.1.1. ISSN 1175-5334.
- ^ Brouwers, E.M. (1994). "Systematic paleontology of Quaternary ostracode assemblages from the Gulf of Alaska; Part 3, Family Cytheruridae". U.S.Geological Survey Professional Paper. Professional Paper. 1544: 11. doi:10.3133/pp1544.
- ^ Eskov, Kirill Y.; Marusik, Y. M. (1995). "On the spiders from Saur Mt. range, eastern Kazakhstan (Arachnida: Araneae)". Beiträge zur Araneologie. 4 (1994): 55–94.
- ^ Eskov, K. Y.; Marusik, Y. M. (1995). "On the spiders from Saur Mt. range, eastern Kazakhstan (Arachnida: Araneae)". Beiträge zur Araneologie. 4: 55–94.
- ^ Dyke, Gareth J.; Gulas, Bonnie E. (2002). "The fossil galliform bird Paraortygoides from the Lower Eocene of the United Kingdom" (PDF). American Museum Novitates (3360): 1–14. doi:10.1206/0003-0082(2002)360<0001:TFGBPF>2.0.CO;2. hdl:2246/2913. S2CID 59131455.
- ^ Willems, W.; Artois, T.; Vermin, W.; Backeljau, T.; Schockaert, E. (2005). ""Typhloplanoida" (Platyhelminthes: Rhabdocoela) from the Indian Ocean, with the description of six new taxa". Journal of Natural History. 39 (19): 1561–1582. Bibcode:2005JNatH..39.1561W. CiteSeerX 10.1.1.688.5041. doi:10.1080/00222930400014239. S2CID 85606704.
- ^ Richards, Steven J.; Oliver, P. M. (2006). "Two new species of large green canopy-dwelling frogs (Anura: Hylidae: Litoria) from Papua New Guinea". Zootaxa. 1295 (1295): 41–60. doi:10.11646/zootaxa.1295.1.3. S2CID 88309117.
- ^ McLay, Colin (2007). "New crabs from hydrothermal vents of the Kermadec Ridge submarine volcanoes, New Zealand: Gandalfus gen. nov.(Bythograeidae) and Xenograpsus (Varunidae)(Decapoda: Brachyura)" (PDF). Zootaxa. 1524: 1–22. doi:10.11646/zootaxa.1524.1.1.
- ^ Gates, M. W. (2008-10-13). "Description of Khamul, gen. n. (Hymenoptera: Chalcidoidea: Eurytomidae), with a hypothesis of its phylogenetic placement". Zootaxa. 1898 (1): 1–33. doi:10.11646/zootaxa.1898.1.1. ISSN 1175-5334. S2CID 85968352.
- ^ Abdala, Cristian Simón; Quinteros, Andrés Sebastián; Espinoza, Robert E. (2008). "Two New Species of Liolaemus (Iguania: Liolaemidae) from the Puna of Northwestern Argentina". Herpetologica. 64 (4): 458–471. doi:10.1655/08-022R1.1. ISSN 0018-0831. S2CID 85932508.
- ^ Cau, Andrea; Dalla Vecchia, Fabio M.; Fabbri, Matteo (2013). "A thick-skulled theropod (Dinosauria, Saurischia) from the Upper Cretaceous of Morocco with implications for carcharodontosaurid cranial evolution". Cretaceous Research. 40: 251–260. Bibcode:2013CrRes..40..251C. doi:10.1016/j.cretres.2012.09.002.
- ^ Vera, Ezequiel Ignacio (2013). "New cyathealean tree fern, Yavanna chimaerica gen. et sp. nov., from the Early Cretaceous of Livingston Island, Antarctica". Cretaceous Research. 44: 214–222. Bibcode:2013CrRes..44..214V. doi:10.1016/j.cretres.2013.04.011. hdl:11336/8854.
- ^ Hastings, Alexander K.; Bloch, Jonathan I.; Jaramillo, Carlos A. (2015-11-17). "A new blunt-snouted dyrosaurid, Anthracosuchus balrogus gen. et sp. nov. (Crocodylomorpha, Mesoeucrocodylia), from the Palaeocene of Colombia". Historical Biology. 27 (8): 998–1020. Bibcode:2015HBio...27..998H. doi:10.1080/08912963.2014.918968. ISSN 0891-2963. S2CID 86315462.
- ^ Hita Garcia, Francisco; Fischer, Georg; Liu, Cong; et al. (22 March 2017). Wong, William Oki (ed.). "X-Ray microtomography for ant taxonomy: An exploration and case study with two new Terataner (Hymenoptera, Formicidae, Myrmicinae) species from Madagascar". PLOS ONE. 12 (3): e0172641. Bibcode:2017PLoSO..1272641H. doi:10.1371/journal.pone.0172641. ISSN 1932-6203. PMC 5362212. PMID 28328931.
- ^ Gunter, Nicole L.; Weir, Thomas A. (2019-03-04). "Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group". Zootaxa. 4564 (1): 41. doi:10.11646/zootaxa.4564.1.2. ISSN 1175-5334. PMID 31716515. S2CID 91878235.
- ^ Monjaraz-Ruedas, R.; Prendini, L.; Francke, O. F. (2019). "Systematics of the short-tailed whipscorpion genus Stenochrus Chamberlin, 1922 (Schizomida: Hubbardiidae), with descriptions of six new genera and five new species". Bulletin of the American Museum of Natural History. 2019 (435): 1–91. doi:10.1206/0003-0090.435.1.1. S2CID 196646084.
- ^ Bauzá, N.; Gelfo, J. N.; López, G. M. (2019). "Early steps in the radiation of notoungulate mammals in southern South America: A new henricosborniid from the Eocene of Patagonia". Acta Palaeontologica Polonica. 64 (3): 597–603. doi:10.4202/app.00565.2018. hdl:11336/127842.
- ^ Kuwahara, G.K.; Marshall, S.A. (20 April 2020). "A revision of the Neotropical genus Chespiritos (Diptera: Sphaeroceridae: Limosininae)". Eur. J. Entomol. 117: 164–189. doi:10.14411/eje.2020.019.
- ^ Haddad, Charles R.; Booysen, Ruan (2022-10-04). "The ground spider genera Leptodrassex Murphy, 2007 and Leptopilos Levy, 2009 (Araneae: Gnaphosidae) in southern Africa, including the description of a new genus and seven new species". Zootaxa. 5194 (1): 1–32. doi:10.11646/zootaxa.5194.1.1. ISSN 1175-5334. PMID 37045347. S2CID 252709842.
- ^ Gonçalves, Rodrigo Barbosa; Pereira, Felipe Walter (2022-05-12). "New species of the cuckoo bee genus Austrosphecodes Michener, 1978 (Hymenoptera: Apoidea: Sphecodini) and a key for Brazilian species". European Journal of Taxonomy (819): 55–89. doi:10.5852/ejt.2022.819.1777. ISSN 2118-9773. S2CID 248760289.
- ^ Espeland, Marianne; Nakahara, Shinichi; Zacca, Thamara; Barbosa, Eduardo P.; Huertas, Blanca; Marín, Mario A.; Lamas, Gerardo; Benmesbah, Mohamed; Brévignon, Christian; Casagrande, Mirna M.; Fåhraeus, Christer; Grishin, Nick; Kawahara, Akito Y.; Mielke, Olaf H. H.; Miller, Jacqueline Y.; Nakamura, Ichiro; Navas, Vanessa; Patrusky, Brooke; Pyrcz, Tomasz W.; Richards, Lindsay; Tan, Denise; Tyler, Stephanie; Viloria, Ángel; Warren, Andrew D.; Xiao, Lei; Freitas, André V. L.; Willmott, Keith R. (10 April 2023). "Combining target enrichment and Sanger sequencing data to clarify the systematics of the diverse Neotropical butterfly subtribe Euptychiina (Nymphalidae, Satyrinae)". Systematic Entomology. 48 (4): 498–570. Bibcode:2023SysEn..48..498E. doi:10.1111/syen.12590. S2CID 258075865.
- ^ Pereira, VD; Ota, Rafaela Priscila; Machado, Valéria Nogueira; Collins, Rupert A; Ândrade, Marcelo C; Garcia-Ayala; Jégu, Miguel; Farias, Izeni; Hrbek, Tomas (2024). "Integrative taxonomy of the black-barred disk pacus (Characiformes: Serrasalmidae), including the redescription of Myloplus schomburgkii and the description of two new species". Neotropical Ichthyology. 22 (2). doi:10.1590/1982-0224-2023-0095.
- ^ "Contribution to the knowledge of the family Braconidae from Canary Islands. (Hymenoptera, Ichneumonoidea). I. A new genus and species of subfamily Doryctinae" (PDF). Vieraea. 3 (1–2): 29–32. 1973.
- ^ Hedqvist, Karl-Johan (1974). "Legolasia dinotiscoides gen. n., sp. n. and Trychnosoma ernobii sp. n., Two New Pteromalids from Sweden (Pteromalidae)". Entomologisk Tidskrift. 95: 117–121.
- ^ a b c Easton, Edward Glynn (1979). "A revision of the "acaecate" earthworms of the Pheretima group (Megascolecidae: Oligochaeta): Archipheretima, Metapheretima, Planapheretima, Pleionogaster, and Polypheretima". Bulletin of the British Museum (Natural History) Zoology. 35: 78–114. doi:10.5962/bhl.part.20451.
- ^ De Clerck, Gert G.; Schockaert, Ernest R. (1995). "Two peculiar new genera of Typhloplanoida from the Western Indian Ocean". Hydrobiologia. 305 (1–3): 3–9. doi:10.1007/BF00036355. ISSN 0018-8158. S2CID 38699976.
- ^ Kaila, L. (1997). A revision of the Nearctic species of Elachista s. l. II. The argentella group (Lepidoptera, Elachistidae). Vol. 206. Acta Zoologica Fennica. pp. 1–93.
- ^ Garrouste, Romain; Lapeyrie, Jean; Steyer, Jean-Sebastien; Giner, Stephen; Nel, Andre (2017). "Insects in the Red Middle Permian of Southern France: first Protanisoptera (Odonatoptera) and new Caloneurodea (Panorthoptera), with biostratigraphical implications". Historical Biology. 30 (4): 546–553. doi:10.1080/08912963.2017.1301448. S2CID 133336679.
- ^ Dallevo-Gomes, Caio I. A.; Mattox, George M. T.; Toledo-Piza, Mônica (2020-10-05). "Taxonomic review of the pipefish genus Pseudophallus Herald, with the description of a new species (Syngnathiformes: Syngnathidae)". Zootaxa. 4859 (1): 81–112. doi:10.11646/zootaxa.4859.1.3. ISSN 1175-5334. PMID 33056206. S2CID 222823500.
- ^ a b "Acrias". www.mindat.org. Retrieved 2022-09-25.
- ^ Hedqvist, Karl-Johan (1978). "Four new genera and species of the subfamily Euderinae from Brazil (Hymenoptera: Chalcidoidea)". Insect Systematics & Evolution. 9 (1): 63–67. doi:10.1163/187631278X00223. ISSN 1399-560X.
- ^ "Allocerastichus". www.mindat.org. Retrieved 2022-09-26.
- ^ Hedqvist, Karl-Johan (1978). "A new subfamily and two new genera and species from the New World (Hymenoptera, Chalcidoidea: Pteromalidae)" (PDF). Entomologica Scandinavica. 9 (2): 135–139. doi:10.1163/187631278x00070.
- ^ Williamson, Thomas E.; Carr, Thomas D. (2012). "Bomburodon, a new name for the Paleocene mammal Bomburia Van Valen, 1978". Journal of Paleontology. 86 (3): 567. Bibcode:2012JPal...86..567W. doi:10.1666/12-013.1. S2CID 129558800.
- ^ Erwin, T. L. (1982). "Small terrestrial ground-beetles of Central America (Carabidae: Bembidiina and Anillina)". Proceedings of the California Academy of Sciences. 4. 42: 455–496 – via BHL.
- ^ Conradie, Werner (2014-04-04). "The King of the Dwarves: a new cryptic species of Dainty Frog (Anura: Pyxicephalidae: Cacosternum) from the eastern Great Escarpment of South Africa". Zootaxa. 3785 (3): 438–452. doi:10.11646/zootaxa.3785.3.6. ISSN 1175-5334. PMID 24872236.
- ^ Sokolov, Igor; Kavanaugh, David (2014-09-30). "The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation". ZooKeys (443): 61–118. Bibcode:2014ZooK..443...61S. doi:10.3897/zookeys.443.7880. ISSN 1313-2970. PMC 4205504. PMID 25349497.
- ^ Paladini, Andressa; Cavichioli, Rodney Ramiro (2015). "A new genus and new species of spittlebug (Hemiptera: Cercopidae: Ischnorhininae) from Southern Brazil". Zoologia (Curitiba). 32 (1): 47–52. doi:10.1590/S1984-46702015000100007. ISSN 1984-4670.
- ^ Singer, David; Kosakyan, Anush; Pillonel, Amandine; Mitchell, Edward A.D.; Lara, Enrique (2015). "Eight species in the Nebela collaris complex: Nebela gimlii (Arcellinida, Hyalospheniidae), a new species described from a Swiss raised bog". European Journal of Protistology. 51 (1): 79–85. doi:10.1016/j.ejop.2014.11.004. PMID 25555254.
- ^ Tencatt, Luiz F. C.; Muriel-Cunha, Janice; Zuanon, Jansen; Ferreira, Marlon F. C.; Britto, Marcelo R. (2020). "A journey through the Amazon Middle Earth reveals Aspidoras azaghal (Siluriformes: Callichthyidae), a new species of armoured catfish from the rio Xingu basin, Brazil". Journal of Fish Biology. 97 (4): 1072–1086. Bibcode:2020JFBio..97.1072T. doi:10.1111/jfb.14467. ISSN 0022-1112. PMID 32672364. S2CID 220584976.
- ^ Pereira, Maria Paula; Gallão, Jonas E.; Bichuette, Maria E.; Pérez-González, Abel (2024-01-31). "Hidden in the caves: a new troglobitic species of Spaeleoleptes and the type species redescription (Opiliones, Laniatores)". European Journal of Taxonomy (921): 36–63. doi:10.5852/ejt.2024.921.2409.
- ^ "Fossilworks: Anisonchus athelas". Paleobiology Database. Retrieved 2022-09-25.
- ^ Blagoderov, V. A. (1998). "Fungus Gnats (Diptera, Mycetophilidae) from the Lower Cretaceous of Mongolia". Paleontological Journal. 32 (6): 598–604. CiteSeerX 10.1.1.616.9726.
- ^ a b Salvador, Rodrigo Brincalepe; Bichuette, Maria Elina (2024-11-08). "Idiopyrgus Pilsbry, 1911 (Gastropoda, Tomichiidae): a relict genus radiating into subterranean environments". Zoosystematics and Evolution. 100 (4): 1543–1556. doi:10.3897/zse.100.136428. ISSN 1860-0743.
- ^ Compagno, L. J. V. (1972). "Ctenacis and Gollum, two new genera of sharks (Selachii; Carcharhinidae)". Proceedings of the California Academy of Sciences. 39 (14): 257–272.
- ^ a b Hedqvist, Karl-Johan (1974). "Two New Genera and Species of the Family Pteromalidae from Sweden" (PDF). Entomologica Scandinavica. 4 (3): 237–240. doi:10.1163/1876312X74X00100.
- ^ Erwin, T. L. (1974). "Studies of the subtribe Tachyina (Coleoptera: Garabidae: Bembidiini), part II: a revision of the New World-Australian genus Pericompsus LeConte". Smithsonian Contributions to Zoology. 162: 1–96.
- ^ Hedqvist, Karl-Johan (1978). "Some Chalcidoidea collected in the Philippine, Bismarck and Solomon Islands, 2. Eucharitidae, with keys and check-lists to Indo-Australian genera (Insecta, Hymenoptera)". Steenstrupia. 4: 227–248.
- ^ Climo, F. M. (1980). "Smeagolida, a new order of gymnomorph mollusc from New Zealand based on a new genus and species". New Zealand Journal of Zoology. 7 (4): 513–522. doi:10.1080/03014223.1980.11760683.
- ^ Helgen, K.; Bonaccorso, F. (2008). "Syconycteris hobbit". IUCN Red List of Threatened Species. 2008: e.T21183A9246881. doi:10.2305/IUCN.UK.2008.RLTS.T21183A9246881.en.
- ^ "Parapagurus hobbiti, New Species (Decapoda, Anomura, Parapaguridae), a Hermit Crab From the Valdivia Bank, Southeast Atlantic". Journal of Crustacean Biology. 3 (3): 472–476. 1983-01-01. doi:10.1163/193724083X00120. ISSN 0278-0372.
- ^ Riedel, Adolf (1988). "Eine neue Zonitiden(?)-Art und -Gattung aus Süd-Anatolien (Gastropoda, Stylommatophora)". Annalen des Naturhistorischen Museums in Wien (in German). 90: 193.
- ^ Melzak, Adam; Westrop, Stephen R. (June 1994). "Mid-Cambrian (Marjuman) trilobites from the Pika Formation, southern Canadian Rocky Mountains, Alberta". Canadian Journal of Earth Sciences. 31 (6): 969–985. Bibcode:1994CaJES..31..969M. doi:10.1139/e94-086.
- ^ Pinto-da-Rocha, Ricardo; Fonseca-Ferreira, Rafael; Bichuette, Maria (2015). "A new highly specialized cave harvestman from Brazil and the first blind species of the genus: Iandumoema smeagol sp. n. (Arachnida, Opiliones, Gonyleptidae)". ZooKeys (537): 79–95. Bibcode:2015ZooK..537...79P. doi:10.3897/zookeys.537.6073. PMC 4714048. PMID 26798238.
- ^ Mattioli, Emanuela (1996). "New calcareous nannofossil species from the Early Jurassic of Tethys". Rivista Italiana di Paleontologia e Stratigrafia. 102 (3): 397–412.
- ^ Wagner, P. J. (1999). "The utility of fossil data in phylogenetic analyses: a likelihood example using Ordovician-Silurian species of the Lophospiridae (Gastropoda: Murchisoniina)". American Malacological Bulletin. 15 (1): 1–31.
- ^ Froese, Rainer; Pauly, Daniel (eds.). "Galaxias gollumoides". FishBase. March 2006 version.
- ^ Minter, Leslie R. (June 2003). "Two new cryptic species of Breviceps (Anura: Microhylidae) from Southern Africa". African Journal of Herpetology. 52 (1): 9–21. Bibcode:2003AfJH...52....9M. doi:10.1080/21564574.2003.9635473. S2CID 84275858.
- ^ Wendt, Tom (2003). "Peperomia hobbitoides (Piperaceae), a New Species of Karstophile from the Rain Forests of the Isthmus of Tehuantepec, Mexico". Lundellia. 6 (1): 37–43. doi:10.25224/1097-993X-6.1.9. ISSN 1097-993X. S2CID 31333362.
- ^ A. Sendra; V. Ortuno; A. Moreno; S. Montagud; S. Teruel (2006). "Gollumjapyx smeagol gen. n., an enigmatic hypogean japygid (Diplura: Japygidae) from the eastern Iberian Peninsula" (PDF). Zootaxa. 1372: 35–52. doi:10.11646/zootaxa.1372.1.4.
- ^ Grismer, L. Lee (2007). "A new species of Ingerophrynus (Anura: Bufonidae) from a lowland rain forest in southern peninsular Malaysia". Journal of Herpetology. 41 (2): 225–230. doi:10.1670/0022-1511(2007)41[225:ANSOIA]2.0.CO;2. S2CID 198157730.
- ^ Machado, A. (2008). "New taxa of Laparocerus Schönherr, 1834 from Madeira and Porto Santo, with a key to the species of the Madeiran archipelago (Coleoptera, Curculionidae, Entiminae)". Graellsia. 64 (2): 307–328. doi:10.3989/graellsia.2008.v64.i2.40.
- ^ Yerman, Michelle N.; Krapp-Schickel, Traudl (2008). "A new genus and two new species of Saurodocus (Crustacea: Amphipoda: Melitidae) from Lizard Island, Queensland, Australia". Zootaxa. 1820 (1820): 60–66. doi:10.11646/zootaxa.1820.1.5.[permanent dead link ]
- ^ Will, Kipling (16 November 2011). "Taxonomic review of the Pterostichini and Loxandrini fauna of New Caledonia (Coleoptera, Carabidae)". ZooKeys (147): 337–397. Bibcode:2011ZooK..147..337W. doi:10.3897/zookeys.147.1943. PMC 3286263. PMID 22371668.
- ^ a b Carnevale, Giorgio; Godfrey, Stephen J. (2014). "Tilefish (Teleostei, Malacanthidae) remains from the Miocene Calvert Formation, Maryland and Virginia: taxonomical and paleoecological remarks". Journal of Vertebrate Paleontology. 34 (5): 1018–1032. Bibcode:2014JVPal..34.1018C. doi:10.1080/02724634.2014.845202. S2CID 129018789.
- ^ Hita Garcia, Francisco; Fisher, Brian (2014-06-04). "The hyper-diverse ant genus Tetramorium Mayr (Hymenoptera, Formicidae) in the Malagasy region ‑ taxonomic revision of the T. naganum, T. plesiarum, T. schaufussii, and T. severini species groups". ZooKeys (413): 1–170. Bibcode:2014ZooK..413....1H. doi:10.3897/zookeys.413.7172. ISSN 1313-2970. PMC 4086027. PMID 25009414.
- ^ Davis, H. R.; Grismer, L. L.; Klabacka, R. L.; Muin, M. A.; Quah, E. S. H.; Anuar, S.; Wood, P. J.; Sites, J. W. (2016). "The phylogenetic relationships of a new Stream Toad of the genus Ansonia Stoliczka, 1870 (Anura: Bufonidae) from a montane region in Peninsular Malaysia". Zootaxa. 4103 (2): 137–53. doi:10.11646/zootaxa.4103.2.4. PMID 27394624. S2CID 11042543.
- ^ Stuke, Jens-Hermann (2016-02-29). "Carnidae (Diptera) in the Canadian National Collection of Insects (Ottawa), with the description of five new species". Zootaxa. 4084 (4): 540–556. doi:10.11646/zootaxa.4084.4.5. ISSN 1175-5334. PMID 27394280.
- ^ Soto Àngel, Joan J.; Peña Cantero, Álvaro L. (2017). "Inhabitant or visitor? Unexpected finding of Aglaophenia (Cnidaria, Hydrozoa) in Antarctic waters". Antarctic Science. 29 (4): 331–339. Bibcode:2017AntSc..29..331S. doi:10.1017/S0954102017000049. S2CID 231658260.
- ^ Stuke JH, Freidberg A (December 2017). "The genera Meoneura Nitzsch and Carnus Rondani (Diptera: Carnidae) in Israel, with the description of ten new species, new records and identification keys" (PDF). Israel Journal of Entomology. 47: 173–214. Archived from the original (PDF) on 4 October 2021. Retrieved 8 November 2022.
- ^ de Gier, Werner; Fransen, Charles H.J.M. (2018). "Odontonia plurellicola sp. n. and Odontonia bagginsi sp. n., two new ascidian-associated shrimp from Ternate and Tidore, Indonesia, with a phylogenetic reconstruction of the genus (Crustacea, Decapoda, Palaemonidae)". ZooKeys (765): 123–160. Bibcode:2018ZooK..765..123D. doi:10.3897/zookeys.765.25277. PMC 6002420. PMID 29910665.
- ^ Britz, Ralf; Anoop, V.K.; Dahanukar, Neelesh; Raghavan, Rajeev (2019-05-09). "The subterranean Aenigmachanna gollum, a new genus and species of snakehead (Teleostei: Channidae) from Kerala, South India". Zootaxa. 4603 (2): 377. doi:10.11646/zootaxa.4603.2.10. ISSN 1175-5334. PMID 31717234. S2CID 164781147.
- ^ Martoni, Francesco (2019). "Resolving an 87-year-old taxonomical curiosity with the description of Psylla frodobagginsi sp. nov. (Hemiptera: Sternorrhyncha: Psyllidae), a second distinct Psylla species on the New Zealand endemic plant kōwhai". PLOS ONE. 14 (9): e0221316. Bibcode:2019PLoSO..1421316M. doi:10.1371/journal.pone.0221316. PMC 6750570. PMID 31532775.
- ^ Qi, Shuo; Wang, Jian; Grismer, L. Lee; Chen, Hong-Hui; Lyu, Zhi-Tong; Wang, Ying-Yong (2020-11-11). "The Stoor Hobbit of Guangdong: Goniurosaurus gollum sp. nov., a cave-dwelling Leopard Gecko (Squamata, Eublepharidae) from South China". ZooKeys (991): 137–153. Bibcode:2020ZooK..991..137Q. doi:10.3897/zookeys.991.54935. ISSN 1313-2970. PMC 7674404. PMID 33223904.
- ^ Lin, Yejie; Marusik, Yuri M.; Gao, Caixia (2021). "Twenty-three new spider species (Arachnida: Araneae) from Asia". Zoological Systematics. 46 (2). Xu, Hao; Zhang, Xiaoqing; Wang, Ziyi; Zhu, Wenhui; Li, Shuqiang: 91–152. doi:10.11865/zs.2021201.
- ^ Schmidt, Ray C.; Bragança, Pedro H. N.; Friel, John P.; Pezold, Frank; Tweddle, Denis; Bart Jr., Henry L. (2023-07-31). "Two New Species of Suckermouth Catfishes (Mochokidae: Chiloglanis) from Upper Guinean Forest Streams in West Africa". Ichthyology & Herpetology. 111 (3): 376–389. doi:10.1643/i2022067. S2CID 260178838.
- ^ Webb, S. David (1969). "The Pliocene Canidae of Florida" (PDF). Bulletin of the Florida State Museum Biological Sciences. 14 (4). University of Florida, Gainesville: 289.
- ^ a b Hita Garcia, Francisco; Fisher, Brian L. (19 December 2012). "The ant genus Tetramorium Mayr (Hymenoptera: Formicidae) in the Malagasy region—taxonomic revision of the T. kelleri and T. tortuosum species groups" (PDF). Zootaxa (3592): 1–85. ISBN 978-1-77557-073-8. ISSN 1175-5334.
- ^ Faúndez, Eduardo; Rider, David; Carvajal, Mariom (2016). "Sobre la identidad de Acledra (Acledra) gregalis Berg, 1878, y descripción de una especie nueva de Acledra Signoret, 1864 (Hemiptera: Heteroptera: Pentatomidae: Pentatominae)". Arquivos Entomolóxicos (in Spanish). 16: 67–72.
- ^ "South American Stink Bug Named after J. R. R. Tolkien's Nazgûl". Entomology Today. September 7, 2016. Retrieved December 13, 2018.
- ^ Quah, Evan S.H.; Anuar, Shahrulm.S.; Grismer, L.L.; Wood, Perry L. Jr.; Azizah, Siti M.N.; Muin, Mohd Abdul (2017-09-15). "A new species of frog of the genus Abavorana Oliver, Prendini, Kraus & Raxworthy 2015 (Anura: Ranidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia". Zootaxa. 4320 (2): 272. doi:10.11646/zootaxa.4320.2.4. ISSN 1175-5334.
- ^ Christodoulou, Magdalini; Iliffe, Thomas M.; Grave, Sammy De (2019-03-16). "A new anchialine cave dwelling species of Potamalpheops Powell, 1979 from the Solomon Islands (Crustacea, Decapoda, Alpheidae)". Crustacean Research. 48: 11–21. doi:10.18353/crustacea.48.0_11. ISSN 0287-3478. S2CID 134623520.
- ^ Borges, Rafael Cabral; Silveira, Orlando Tobias (2019-08-20). "Revision of the species-group of Mischocyttarus (Omega) filiformis (de Saussure 1854), with description of three new species (Hymenoptera, Vespidae)". Zootaxa. 4657 (3): 545–564. doi:10.11646/zootaxa.4657.3.7. ISSN 1175-5334. PMID 31716774. S2CID 202019600.
- ^ "A new genus and species from Romania [sic!], representing a new family (Hymenoptera, Chalcidoidea)" (PDF). Polskie Pismo Entomologiczne. 44 (2): 253–254. 1974.
- ^ "Taxon details". Fauna Europaea. Archived from the original on August 16, 2016. Retrieved 20 March 2016.
- ^ "Fossilworks: Ankalagon". Paleobiology Database. Retrieved 2022-09-25.
- ^ Rosser, Thomas G.; Alberson, Neely R.; Baumgartner, Wes A.; Mauel, Michael J.; Pote, Linda M.; Griffin, Matt J. (2016). "Morphological, Histological, and Molecular Description of Unicauda fimbrethilae n. sp. (Cnidaria: Myxosporea: Myxobolidae) from the Intestinal Tract of Channel Catfish Ictalurus punctatus". Journal of Parasitology. 102 (1): 105–113. doi:10.1645/15-810. PMID 26377372. S2CID 10700892.
- ^ Cooper, Kenneth W. (1964). "The first fossil tardigrade: Beorn leggi Cooper, from Cretaceous amber". Psyche: A Journal of Entomology. 71 (2): 41–48. doi:10.1155/1964/48418.
- ^ Hedqvist, Karl-Johan (1975). "Notes on Chalcidoidea. VIII. The Swedish Chrysolampini with description of a new genus and species (Hym., Chalcidoidea, Pteromalidae)" (PDF). Entomologisk Tidskrift. 96 (3–4): 133–136.
- ^ "Fossilworks: Loxolophus hyattianus". Paleobiology Database. Retrieved 2022-09-25.
- ^ Conway Morris, S (1977). "Fossil priapulid worms" (PDF). Special Papers in Palaeontology. 20: 55.
- ^ Gee 2004, p. 54
- ^ Pekár, S.; Cardoso, P. (2005). "Ant-eating spiders (Araneae: Zodariidae) of Portugal: additions to the current knowledge". Zootaxa. 1009 (1009): 51–60. doi:10.11646/zootaxa.1009.1.5.
- ^ Decae, A.; Cardoso, P.; Selden, P. (2007). "Taxonomic review of the Portuguese Nemesiidae (Araneae, Mygalomorphae)" (PDF). Revista Ibérica de Aracnología. 14: 1–18.
- ^ Abdala, Cristian Simón; Quinteros, Andrés Sebastián; Scrocchi, Gustavo José; Stazzonelli, Juan Carlos (2010). "Three new species of the Liolaemus elongatus group (Iguania: Liolaemidae) from Argentina". Cuadernos de Herpetología. 24 (2): 96.
- ^ Stanley, Edward L.; Bauer, Aaron M.; Jackman, Todd R.; Branch, William R.; Mouton, P. Le Fras N. (2011). "Between a rock and a hard polytomy: Rapid radiation in the rupicolous girdled lizards (Squamata: Cordylidae)" (PDF). Molecular Phylogenetics and Evolution. 58 (1): 53–70. Bibcode:2011MolPE..58...53S. doi:10.1016/j.ympev.2010.08.024. PMID 20816817.
- ^ Sánchez, Nuria; Soo Rho, Hyun; Min, Won Gi; Kim, Dongsung; Vinther Sørensen, Martin (2013-06-30). "Four new species of Pycnophyes (Kinorhyncha: Homalorhagida) from Korea and the East China Sea". Scientia Marina. 77 (2): 353–380. doi:10.3989/scimar.03769.15A. ISSN 1886-8134. S2CID 56005860.
- ^ Bulanov, V. V.; Sennikov, A. G. (2015). "Glaurung schneideri gen. et sp. nov., a new weigeltisaurid (Reptilia) from the Kupfershiefer (Upper Permian) of Germany". Paleontological Journal. 49 (12): 1353–1364. Bibcode:2015PalJ...49.1353B. doi:10.1134/S0031030115120035. S2CID 87461613.
- ^ Faúndez, Eduardo (19 June 2015). "Patagonian Shield Bug Named After Middle's Earth's Smaug the Dragon". Entomology Today. Entomological Society of America. Retrieved 20 March 2016.
- ^ Carvajal, Mariom A.; Faúndez, Eduardo I.; Rider, David A. (2015). "Contribución al conocimiento de los Acanthosomatidae (Hemiptera: Heteroptera) de la Región de Magallanes, con descripción de una nueva especie". Anales del Instituto de la Patagonia (Chile). 43 (1): 145–151. doi:10.4067/s0718-686x2015000100013.
- ^ Carvajal, Mariom A.; Faundez, Eduardo I.; Rider, David A. (2015-12-31). "Tamolia ancalagon Carvajal, Faundez & Rider, 2015, n. sp". Zootaxa. doi:10.5281/ZENODO.6094575.
- ^ a b c Sørensen, Martin Vinther; Grzelak, Katarzyna (2018). "New mud dragons from Svalbard: three new species of Cristaphyes and the first Arctic species of Pycnophyes (Kinorhyncha: Allomalorhagida: Pycnophyidae)". PeerJ. 6: e5653. doi:10.7717/peerj.5653. PMC 6166639. PMID 30280030.
- ^ Brescovit, Antonio D.; Cizauskas, Igor; Mota, Leandro P. (2018). "Seven new species of the spider genus Ochyrocera from caves in Floresta Nacional de Carajás, PA, Brazil (Araneae, Ochyroceratidae)". ZooKeys (726): 87–130. Bibcode:2018ZooK..726...87B. doi:10.3897/zookeys.726.19778. PMC 5799772. PMID 29416386.
- ^ Willkens Y, Rebêlo GL, Santos JN, Furtado AP, Vilela RV, Tkach VV, Kuzmin Y, Melo, F (2020). "Rhabdias glaurungi sp. nov. (Nematoda: Rhabdiasidae), parasite of Scinax gr. ruber (Laurenti, 1768) (Anura: Hylidae), from the Brazilian Amazon". Journal of Helminthology. 94: E54. doi:10.1017/S0022149X19000476. PMID 31630693. S2CID 196678433.
- ^ Pal, S.; Mirza, Z. A.; Dsouza, P. & Shanker, K. (2021). "Diversifying on the Ark: multiple new endemic lineages of dwarf geckos from the Western Ghats provide insights into the systematics and biogeography of South Asian Cnemaspis (Reptilia: Squamata)". Zoological Research. 42 (6): 675–691. doi:10.24272/j.issn.2095-8137.2021.074. PMC 8645882. PMID 34581029.
- ^ "Fossilworks: Oxyacodon agapetillus". Paleobiology Database. Retrieved 2022-09-25.
- ^ "Fossilworks: Oxyclaenus". Paleobiology Database. Retrieved 2022-09-25.
- ^ Kolibac, J. (1997). "Classification of the subfamilies of Cleridae (Coleoptera: Cleroidea)". Acta Musei Moraviae, Scientiae Naturales. 81: 307–361.
- ^ Loredo, Ariel I.; Wood, Perry Jr. L.; Quah, Evan S. H.; Anuar, Shahrul; Greer, Lee F.; Ahmad, Norhayati; Grismer, Lee (2013-05-28). "Cryptic speciation within Asthenodipsas vertebralis (Boulenger, 1900) (Squamata: Pareatidae), the description of a new species from Peninsular Malaysia, and the resurrection of A. tropidonotus (Lidth de Jude, 1923) from Sumatra: an integrative taxonomic analysis". Zootaxa. 3664 (4): 505–524. doi:10.11646/zootaxa.3664.4.5. ISSN 1175-5334. PMID 26266316.
- ^ Landry, Bernard (2016-12-31). "Taxonomic revision of the Spilomelinae (Lepidoptera, Pyralidae s. l.) of the Galápagos Islands, Ecuador". Revue suisse de Zoologie. 123 (2): 315–399. doi:10.5281/ZENODO.155309. S2CID 229199262.
- ^ Piazza, Veronica; Hammer, Øyvind; Jattiot, Romain (2017). "New late Smithian (Early Triassic) ammonoids from the Lusitandiadalan Member, Vikinghøgda Formation, Svalbard" (PDF). Norwegian Journal of Geology. 97 (2): 105–117. doi:10.17850/njg97-2-03.
- ^ ZANATA, ANGELA M.; BURGER, RAFAEL; CAMELIER, PRISCILA (2018). "Two new species of Astyanax Baird & Girard (Characiformes: Characidae) from the upper rio Paraguaçu basin, Chapada Diamantina, Bahia, Brazil". Zootaxa. 4438 (3): 471–490. doi:10.11646/zootaxa.4438.3.3. PMID 30313131. S2CID 52977293.
- ^ Boyd, David A.; So, Nam; Thach, Phanara; Page, Lawrence M. (2018). "Acantopsis bruinen, a new species of horseface loach from Southeast Asia (Teleostei: Cobitidae)". Ichthyological Exploration of Freshwaters. 29 (1): 1–8. doi:10.23788/IEF-1096.
- ^ Ratsoavina, Fanomezana M.; Glaw, Frank; Raselimanana, Achille P.; Rakotoarison, Andolalao; Vieites, David R.; Hawlitschek, Oliver; Vences, Miguel; Scherz, Mark D. (2020-12-15). "Towards completion of the species inventory of small-sized leaf-tailed geckos: two new species of Uroplatus from northern Madagascar". Zootaxa. 4895 (2): 251–271. doi:10.11646/zootaxa.4895.2.5. ISSN 1175-5334. PMID 33756904. S2CID 230563236.
- ^ Korb, S.; Gorbunov, P.; Melyakh, S. (2023). "A new remarkable species of the genus Hyperlais Marion, 1959 (Lepidoptera: Crambidae) from Kyrgyzstan". Zootaxa. 5318 (3): 432–436. doi:10.11646/zootaxa.5318.3.8. PMID 37518371 – via ResearchGate.
- ^ "Fossilworks: Litocherus lacunatus". Paleobiology Database. Retrieved 2022-09-26.
- ^ "Fossilworks: Oxyclaenus pugnax". Paleobiology Database. Retrieved 2022-09-25.
- ^ Larsen 2007, citing: Foster, Robert (2001). The Complete Guide to Middle-earth. New York: Ballantine. p. 23.
- ^ Costa, Henrique C.; Santana, Diego J.; Leal, Fernando; Koroiva, Ricardo; Garcia, Paulo C.A. (2016). "A New Species of Helicops (Serpentes: Dipsadidae: Hydropsini) from Southeastern Brazil". Herpetologica. 72 (2): 157–166. doi:10.1655/HERPETOLOGICA-D-15-00059. S2CID 88849028.
- ^ Dathe, Holger H.; Proshchalykin, Maxim Yu. (2016-06-10). "The genus Hylaeus Fabricius in Mongolia, an updated species inventory (Hymenoptera: Apoidea, Colletidae)". Zootaxa. 4121 (4): 351–382. doi:10.11646/zootaxa.4121.4.1. ISSN 1175-5334. PMID 27395230.
- ^ D'Udekem d'Acoz, Cédric; Verheye, Marie L. (2017-10-17). "Epimeria of the Southern Ocean with notes on their relatives (Crustacea, Amphipoda, Eusiroidea)". European Journal of Taxonomy (359). doi:10.5852/ejt.2017.359. ISSN 2118-9773.
- ^ Bustamante, Abel A.; Ruiz, Gustavo R.S. (2017-12-05). "Systematics of Thiodinini (Araneae: Salticidae: Salticinae), with description of a new genus and twelve new species". Zootaxa. 4362 (3): 301–347. doi:10.11646/zootaxa.4362.3.1. ISSN 1175-5334. PMID 29245433.
- ^ Zacca, T.; Casagrande, M. M.; Mielke, O. H.; Huertas, B.; Espeland, M.; Freitas, A. V. L.; Willmott, K. R.; Nakahara, N.; Lamas, G. (2020). "Revalidation of Vareuptychia Forster, 1964, description of Vanima gen. nov., and notes on Euptychia cleophes Godman & Salvin, 1889 (Lepidoptera: Nymphalidae: Satyrinae)". Zootaxa. 4858 (1): zootaxa.4858.1.1. doi:10.11646/zootaxa.4858.1.1. PMID 33056239. S2CID 222839782.
- ^ Britto, Marcelo R.; Lima, Flávio C. T.; Santos, Alexandre C. A. (2005). "A new Aspidoras (Siluriformes: Callichthyidae) from rio Paraguaçu basin, Chapada Diamantina, Bahia, Brazil". Neotropical Ichthyology. 3 (4): 473–479. doi:10.1590/S1679-62252005000400004. ISSN 1679-6225.
- Works cited
- Gee, Henry (2004). The Science of Middle-earth (1st ed.). Cold Spring Harbor, New York: Cold Spring Press. ISBN 978-1593600235.
- Larsen, Kristine (2007). "Sauron, Mount Doom, and Elvish Moths: The Influence of Tolkien on Modern Science". Tolkien Studies. 4 (1): 223–234. doi:10.1353/tks.2007.0024. S2CID 170563966.