Jump to content

Stegodon

From Wikipedia, the free encyclopedia
(Redirected from Stegodon sompoensis)

Stegodon
Temporal range: Late Miocene–Late Pleistocene, 11.6–0.01 Ma
Stegodon skeleton at the Gansu Provincial Museum
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Proboscidea
Family: †Stegodontidae
Genus: †Stegodon
Falconer, 1847
Species
  • †S. aurorae (Matsumoto, 1918)
  • †S. elephantoides (Clift, 1828)
  • †S. florensis Hooijer, 1957
  • †S. ganesha
    (Faloner and Cautley, 1846)
  • †S. kaisensis Hopwood, 1939
  • †S. luzonensis
    von Koenigswald, 1956
  • †S. miensis (Matsumoto, 1941)
  • †S. mindanensis (Naumann, 1890)
  • †S. orientalis Owen, 1870
  • †S. protoaurorae (Aiba et al., 2010)
  • †S. sompoensis Hooijer, 1964
  • †S. sondaari van den Bergh, 1999
  • †S. trigonocephalus (Martin, 1887)
  • †S. zdanskyi Hopwood, 1935

Stegodon ("roofed tooth" from the Ancient Greek words στέγω, stégō, 'to cover', + ὀδούς, odoús, 'tooth' because of the distinctive ridges on the animal's molars) is an extinct genus of proboscidean, related to elephants. It was originally assigned to the family Elephantidae along with modern elephants but is now placed in the extinct family Stegodontidae. Like elephants, Stegodon had teeth with plate-like lophs that are different from those of more primitive proboscideans like gomphotheres and mammutids.[1] Fossils of the genus are known from Africa and across much of Asia, as far southeast as Timor (with a single record in southeast Europe). The oldest fossils of the genus are found in Late Miocene strata in Asia, likely originating from the more archaic Stegolophodon, subsequently migrating into Africa.[2] While the genus became extinct in Africa during the Pliocene, Stegodon persisted in South, Southeast and Eastern Asia into the Late Pleistocene.[3]

Morphology

[edit]

The skull of Stegodon is relatively tall but short.[1] The lower jaw in comparison to early elephantimorphs is shortened (brevirostrine), and lacks lower tusks/incisors. The molar teeth are superficially like those of elephants, consisting of parallel lamellae that form ridges but are generally relatively low crowned (brachydont),[1][4] the numbers of ridges are greater in later species.[5] Members of the genus lack permanent premolars.[6] The tusks are proportionally large, with those of the biggest species being among the largest known tusks in proboscideans, with a particularly large tusk of S. ganesa from the Early Pleistocene of India measured to be 3.89 metres (12.8 ft) long, with an estimated mass of approximately 140 kilograms (310 lb), substantially larger than the largest recorded modern elephant tusk.[7]

Stegodon molars
  • Molar of Stegodon orientalis
    Molar of Stegodon orientalis
  • Molar of Stegodon trigonocephalus
    Molar of Stegodon trigonocephalus

Size

[edit]
Skeletal restoration of a large male S. zdanskyi with a shoulder height of around 3.87 metres

The Chinese S. zdanskyi is suggested to be the largest species, and is known from an old male (50-plus years old) from the Yellow River that is 3.87 m (12.7 ft) tall and would have weighed approximately 12.7 tonnes (12.5 long tons; 14.0 short tons) in life. It had a humerus 1.21 m (4.0 ft) long, a femur 1.46 m (4.8 ft) long, and a pelvis 2 m (6.6 ft) wide. The Indian S. ganesa is suggested to have a shoulder height of about 3.10 m (10.2 ft), and a body mass of around 6.5 tonnes (6.4 long tons; 7.2 short tons). The Javanese species S. trigonocephalus is suggested to have been around 2.75–2.8 m (9.0–9.2 ft) tall, with a body mass of around 5 tonnes (4.9 long tons; 5.5 short tons).[8] S. orientalis was around the size of an Asian elephant (Elephas maximus).[9]

Dwarfism

[edit]
Main article: Dwarf_elephant § Indonesia_and_the_Philippines
Estimated sizes of dwarf Stegodon species from Flores compared to a human

Similar to modern-day elephants, stegodonts were likely good swimmers,[10][11] allowing them to disperse to remote islands in Indonesia, the Philippines and Japan. Once present on the islands, due to the process of insular dwarfism, as a result of decreased land area and the reduction of predation and competition pressure, they reduced in body size, with the degree of dwarfism varying between islands as the result of local conditions. One of the smallest species, Stegodon sumbaensis from Sumba in Indonesia, is estimated at around 8% of the size of mainland Stegodon species, with a body mass of 250 kilograms (550 lb).[12] Sometimes the same island was colonised multiple times by Stegodon, as in Flores, where the Early Pleistocene strongly dwarfed species Stegodon sondaari, which was 120 cm (3.9 ft) tall at the shoulder and weighed about 350–400 kilograms (770–880 lb),[8] was replaced by the species Stegodon florensis during the Middle Pleistocene which was initially substantially larger, but progressively reduced in size over time, with the earlier subspecies Stegodon florensis florensis from the Middle Pleistocene estimated to be around 50% the size of mainland Stegodon species with a shoulder height of around 190 cm (6.2 ft) and a body mass of around 1.7 tons, while the later Stegodon florensis insularis from the Late Pleistocene is estimated to be around 17% the size of mainland Stegodon species, with a shoulder height of around 130 cm (4.3 ft), and a body mass of about 570 kilograms (1,260 lb).[12][13]

During Pliocene-Early Pleistocene (from around 4-1 million years ago), a succession of endemic dwarf species of Stegodon, probably representing a single lineage lived in the Japanese archipelago, probably derived from the mainland Chinese S. zdanskyi. In chronological succession these species are Stegodon miensis (4-3 million years ago) Stegodon protoaurorae (3-2 million years ago) and Stegodon aurorae, (2-1 million years ago) which show a progressive size reduction through time, possibly as a result of reducing land area of the Japanese archipelago.[14] The latest and smallest species S. aurorae is estimated to be 25% the size of its mainland ancestor with a body mass of around 2,122 kilograms (4,678 lb).[15] S. aurorae also shows morphological straits associated with dwarfism, like shortened limbs.[16]

Ecology

[edit]
Life restoration of the Indian species Stegodon ganesa

Like modern elephants, but unlike more primitive proboscideans, Stegodon is thought to have chewed using a proal movement (a forward stroke from the back to the front) of the lower jaws. This jaw movement is thought to have evolved independently in elephants and stegodontids.[1] Stegodon populations from the Late Pliocene of the India (including Stegodon insignis) are suggested to have been variable mixed feeders, while those from the earliest Pleistocene (including Stegodon ganesa) of the same region are suggested to have been nearly pure grazers based on isotopic analysis.[17] Based on dental microwear analysis, populations of Stegodon from the Pleistocene of China (Stegodon orientalis and Stegodon huananensis) and mainland southeast Asia (S. orientalis) were found to be browsers, with clear niche differentiation from sympatric Elephas populations, which tended towards mixed feeding (both browsing and grazing),[18][19] though isotopic analysis of Stegodon cf. orientalis specimens from the late Middle Pleistocene of Thailand suggests that these individuals were mixed feeders that consumed a significant amount of C4 grass.[20] Specimens of Stegodon trigonocephalus from the Early-Middle Pleistocene of Java were found to be mixed feeders to grazers, with a diet similar to that of sympatric Elephas hysudrindicus.[21] The dwarf species from Flores, Stegodon sondaari and Stegodon florensis, are suggested to have been mixed feeders and grazers, respectively, based on stable carbon isotopes.[13] Specimens of Stegodon kaiesensis from the Pliocene of East Africa were found to be browsers to mixed feeders, based on mesowear analysis.[22]

Tracks of a group of Stegodon from the Late Pliocene of Japan suggest that like modern elephants, Stegodon lived in social herds.[23]

On Flores, where dwarf Stegodon species were the only large herbivores, they were likely the main prey of the Komodo dragon.[24]

Taxonomy

[edit]
Fossils of S. aurorae (left) and S. orientalis (right) at the National Museum of Nature and Science, Tokyo
Jaw fossil of S. sondaari at the Bandung Geological Museum
Skull of S. ganesha

In the past, stegodonts were believed to be the ancestors of the true elephants and mammoths, but currently they are believed to have no modern descendants. Stegodon is likely derived from Stegolophodon, an extinct genus known from the Miocene of Asia,[2] with transitional fossils between the two genera known from the Late Miocene of Southeast Asia and Yunnan in South China.[1] Stegodon is more closely related to elephants and mammoths than to mastodons.[25] Like elephants, stegodontids are believed to have derived from gomphotheres.[26]

Phylogeny

[edit]

The following cladogram shows the placement of the genus Stegodon among other proboscideans, based on hyoid characteristics:[25]

Mammut americanum (American mastodon)

Gomphotherium sp.

Stegodon zdanskyi

Loxodonta africana (African bush elephant)

Elephas maximus (Asian elephant)

Mammuthus columbi (Columbian mammoth)

List of species

[edit]
  • Stegodon kaisensis Late Miocene – Pliocene, Africa
  • Stegodon zdanskyi Late Miocene – Pliocene, China
  • Stegodon huananensis Early Pleistocene, China
  • Stegodon orientalis Middle – Late Pleistocene, China, Southeast Asia, Japan, Taiwan
  • Stegodon namadicus/S. insignis/S. ganesa Pliocene – Late Pleistocene, India
  • Stegodon miensis Pliocene, Japan
  • Stegodon protoaurorae Late Pliocene – Early Pleistocene, Japan
  • Stegodon aurorae Early Pleistocene – early Middle Pleistocene, Japan
  • Stegodon sondaari Early Pleistocene, Flores, Indonesia
  • Stegodon florensis Middle – Late Pleistocene, Flores, Indonesia
  • Stegodon luzonensis Middle Pleistocene, Luzon, Philippines
  • Stegodon trigonocephalus late Early Pleistocene – early Late Pleistocene, Java, Indonesia
  • Stegodon sompoensis Late Pliocene – Early Pleistocene, Sulawesi, Indonesia
  • Stegodon sumbaensis Middle – Late Pleistocene, Sumba, Indonesia
  • Stegodon timorensis Middle Pleistocene, Timor, Indonesia
  • Stegodon mindanensis Pleistocene Mindanao, Philippines

An indeterminate Stegodon molar of an uncertain locality and age is known from Greece, representing the only record of the genus in Europe.[27] Indeterminate remains are also known from the Early Pleistocene and early Middle Pleistocene of Israel.[28]

Relationship with humans

[edit]

Remains at a number of sites suggest that humans (in a broad sense, including archaic humans) interacted with Stegodon.[29] At a cave deposit on Gele Mountain near Chongqing in southwest China, a mandible of Stegodon orientalis was used to make a handaxe, with dating suggesting the bone is around 170,000 years old.[30] At the late Middle Pleistocene Panxian Dadong cave site in southern Guizhou Province, southwest China, dating to around 300-190,000 years ago,[31] numerous remains of juvenile (0-12 years of age) and a much smaller number of adult remains of adult Stegodon orientalis, representing a minimum of 12 individuals were found at the site in association with stone tools and human remains. It suggested that Stegodon remains were brought to the cave by humans though none of the elements show clear evidence of processing.[9][32] At the Xinlong Cave site in the Three Gorges area of Chongqing, suggested to date to around 200-130,000 years ago, two Stegodon cf. orientalis tusks have been found along with human remains. These tusks appear to have been delibrately engraved with patterns and are suggested to have been brought into the cave by humans.[33] At the Late Pleistocene Ma’anshan site also in Guizhou, remains of Stegodon orientalis including both adults and juveniles among other animals are found in two layers, the older dating to around 53,000 years Before Present (BP), with the younger dates to around 19,295-31,155 years BP with the minimum number of individuals being 7 and 2 for the older and younger layers respectively, with the older layer containing adults and juveniles while in the younger later only juveniles are present. Bones at the site display cut marks indicating butchery, and are thought to have been accumulated at the site by people, likely by hunting or possibly scavenging in the case of the large adults found in the older layer.[34]

At Liang Bua cave on Flores dating to around 80-50,000 years ago, remains of the dwarf Stegodon species Stegodon florensis are associated with stone tools produced by the dwarf archaic human species Homo floresiensis, with a small number of the bones bearing cut marks. The ambiguous circumstantial association between bones and stone tools, and the rarity of cut marks makes it unclear to what if to any degree, hunting of Stegodon was practiced by Homo floresiensis.[35][36]

Evolution and extinction

[edit]

The oldest fossils of Stegodon in Asia date to the Late Miocene, around 8-11 million years ago,[2] with the oldest fossils of the genus in Africa being around 7-6 million years old.[16] Stegodon became extinct in Africa during the late Pliocene, around 3 million years ago suggested to be the result of expansion of grassland habitats.[22]

The Javanese species Stegodon trigonocephalus became extinct around 130-80,000 years ago during the latest Middle Pleistocene-early Late Pleistocene (Marine Isotope Stage 5) following a change to more humid conditions, which may have reduced grazing habitat.[21] The last records of Stegodon florensis date to around 50,000 years ago, around the time of arrival of modern humans to Flores (the earliest evidence of which dates to 46,000 years ago), suggesting that effects of modern human activity were likely the cause of their extinction.[37]

Stegodon became extinct in the Indian subcontinent (Stegodon namadicus/Stegodon sp.), mainland Southeast Asia and China (S. orientalis) at some point during the Late Pleistocene epoch, while Asian elephants, which existed in sympatry with Stegodon in these regions, are still extant. The precise timing of extinction is uncertain for these regions,[38][19][39][40] though in India records of Stegodon may date as recently as 35-30,000 years ago.[38] The survival of the Asian elephant as opposed to Stegodon orientalis in Southeast Asia and South China has been suggested to be due to its more flexible diet in comparison to S. orientalis.[19] Although some authors have claimed a Holocene survival in China for S. orientalis,[41] these claims cannot be substantiated due to loss of specimens and issues regarding dating.[39]

References

[edit]
Wikispecies has information related to Stegodon.
  1. ^ a b c d e Saegusa, Haruo (March 2020). "Stegodontidae and Anancus: Keys to understanding dental evolution in Elephantidae". Quaternary Science Reviews. 231: 106176. Bibcode:2020QSRv..23106176S. doi:10.1016/j.quascirev.2020.106176. S2CID 214094348.
  2. ^ a b c Ao, Hong; Zhang, Peng; Dekkers, Mark J.; Roberts, Andrew P.; An, Zhisheng; Li, Yongxiang; Lu, Fengyan; Lin, Shan; Li, Xingwen (January 2016). "New magnetochronology of Late Miocene mammal fauna, NE Tibetan Plateau, China: Mammal migration and paleoenvironments". Earth and Planetary Science Letters. 434: 220–230. Bibcode:2016E&PSL.434..220A. doi:10.1016/j.epsl.2015.11.019. hdl:1874/330309.
  3. ^ O’Regan, H. J.; Bishop, L. C.; Lamb, A.; Elton, S.; Turner, A. (1 January 2005). "Large mammal turnover in Africa and the Levant between 1.0 and 0.5 Ma". Geological Society, London, Special Publications. 247 (1): 231–249. Bibcode:2005GSLSP.247..231O. doi:10.1144/GSL.SP.2005.247.01.13. ISSN 0305-8719. S2CID 129900545.
  4. ^ Cantalapiedra, Juan L.; Sanisidro, Óscar; Zhang, Hanwen; Alberdi, María T.; Prado, José L.; Blanco, Fernando; Saarinen, Juha (1 July 2021). "The rise and fall of proboscidean ecological diversity". Nature Ecology & Evolution. 5 (9): 1266–1272. Bibcode:2021NatEE...5.1266C. doi:10.1038/s41559-021-01498-w. ISSN 2397-334X. PMID 34211141. S2CID 235712060.
  5. ^ van der Made, J. The Evolution of the Elephants and Their Relatives in the Context of Changing Climate and Geography. In Elefantentreich—Eine Fossilwelt in Europa; Verlag Beier & Beran: Langenweißbach, Germany, 2010; pp. 340–360. ISBN 978-3-939414-48-3.
  6. ^ Sanders, William J. (17 February 2018). "Horizontal tooth displacement and premolar occurrence in elephants and other elephantiform proboscideans". Historical Biology. 30 (1–2): 137–156. Bibcode:2018HBio...30..137S. doi:10.1080/08912963.2017.1297436. ISSN 0891-2963. S2CID 89904463.
  7. ^ Larramendi, Asier (10 December 2023). "Estimating tusk masses in proboscideans: a comprehensive analysis and predictive model". Historical Biology: 1–14. doi:10.1080/08912963.2023.2286272. ISSN 0891-2963. S2CID 266182491.
  8. ^ a b Larramendi, A. (2016). "Shoulder height, body mass and shape of proboscideans" (PDF). Acta Palaeontologica Polonica. 61. doi:10.4202/app.00136.2014. S2CID 2092950.
  9. ^ a b Schepartz, Lynne, and Sari Miller-Antonio. 2010. “Large Mammal Exploitation in Late Middle Pleistocene China: A Comparison of Rhinoceros & Stegodonts at Panxian Dadong.” Before Farming 4: 1–14.
  10. ^ Simpson, G. (1977). "Too Many Lines; The Limits of the Oriental and Australian Zoogeographic Regions". Proceedings of the American Philosophical Society, 121(2), 107–120. Retrieved from http://www.jstor.org/stable/986523
  11. ^ Bird, Michael I.; Condie, Scott A.; O’Connor, Sue; O’Grady, Damien; Reepmeyer, Christian; Ulm, Sean; Zega, Mojca; Saltré, Frédérik; Bradshaw, Corey J. A. (2019). "Early human settlement of Sahul was not an accident". Scientific Reports. 9 (1): 8220. Bibcode:2019NatSR...9.8220B. doi:10.1038/s41598-019-42946-9. PMC 6579762. PMID 31209234.
  12. ^ a b Geer, Alexandra A. E.; Bergh, Gerrit D.; Lyras, George A.; Prasetyo, Unggul W.; Due, Rokus Awe; Setiyabudi, Erick; Drinia, Hara (August 2016). "The effect of area and isolation on insular dwarf proboscideans". Journal of Biogeography. 43 (8): 1656–1666. Bibcode:2016JBiog..43.1656V. doi:10.1111/jbi.12743. ISSN 0305-0270. S2CID 87958022.
  13. ^ a b Puspaningrum, Mika; Van Den Bergh, Gerrit; Chivas, Allan; Setiabudi, Erick; Kurniawan, Iwan; Brumm, Adam; and Sutikna, Thomas, "Preliminary results of dietary and environmental reconstructions of Early to Middle Pleistocene Stegodons from the So'a Basin of Flores, Indonesia, based on enamel stable isotope records" (2014). Faculty of Science, Medicine and Health - Papers: part A. 2035.
  14. ^ Aiba, Hiroaki; Baba, Katsuyoshi; Matsukawa, Masaki (10 March 2010). "A new species of Stegodon (Mammalia, Proboscidea) from the Kazusa Group (lower Pleistocene), Hachioji City, Tokyo, Japan and its evolutionary morphodynamics: STEGODON PROTOAURORAE SP. NOV. AND MORPHODYNAMICS". Palaeontology. 53 (3): 471–490. Bibcode:2010Palgy..53..471A. doi:10.1111/j.1475-4983.2010.00953.x.
  15. ^ Geer, Alexandra A. E.; Bergh, Gerrit D.; Lyras, George A.; Prasetyo, Unggul W.; Due, Rokus Awe; Setiyabudi, Erick; Drinia, Hara (August 2016). "The effect of area and isolation on insular dwarf proboscideans". Journal of Biogeography. 43 (8): 1656–1666. Bibcode:2016JBiog..43.1656V. doi:10.1111/jbi.12743. ISSN 0305-0270. S2CID 87958022.
  16. ^ a b Saegusa, Haruo; Thasod, Yupa; Ratanasthien, Benjavun (2005). "Notes on Asian stegodontids". Quaternary International. 126–128: 31–48. Bibcode:2005QuInt.126...31S. doi:10.1016/j.quaint.2004.04.013.
  17. ^ Patnaik, Rajeev; Singh, Ningthoujam Premjit; Paul, Debajyoti; Sukumar, Raman (November 2019). "Dietary and habitat shifts in relation to climate of Neogene-Quaternary proboscideans and associated mammals of the Indian subcontinent". Quaternary Science Reviews. 224: 105968. Bibcode:2019QSRv..22405968P. doi:10.1016/j.quascirev.2019.105968. S2CID 210307849.
  18. ^ Zhang, Hanwen; Wang, Yuan; Janis, Christine M.; Goodall, Robert H.; Purnell, Mark A. (25 July 2017). "An examination of feeding ecology in Pleistocene proboscideans from southern China (Sinomastodon, Stegodon, Elephas), by means of dental microwear texture analysis". Quaternary International. VIth International Conference on Mammoths and their Relatives, Part 3. 445: 60–70. Bibcode:2017QuInt.445...60Z. doi:10.1016/j.quaint.2016.07.011. hdl:1983/4f6a743a-7b6d-47c8-a56a-fee7e2c515df. ISSN 1040-6182.
  19. ^ a b c Ma, Jiao; Wang, Yuan; Jin, Changzhu; Hu, Yaowu; Bocherens, Hervé (15 May 2019). "Ecological flexibility and differential survival of Pleistocene Stegodon orientalis and Elephas maximus in mainland southeast Asia revealed by stable isotope (C, O) analysis". Quaternary Science Reviews. 212: 33–44. Bibcode:2019QSRv..212...33M. doi:10.1016/j.quascirev.2019.03.021. ISSN 0277-3791. S2CID 135056116.
  20. ^ Suraprasit, Kantapon; Bocherens, Hervé; Chaimanee, Yaowalak; Panha, Somsak; Jaeger, Jean-Jacques (August 2018). "Late Middle Pleistocene ecology and climate in Northeastern Thailand inferred from the stable isotope analysis of Khok Sung herbivore tooth enamel and the land mammal cenogram". Quaternary Science Reviews. 193: 24–42. Bibcode:2018QSRv..193...24S. doi:10.1016/j.quascirev.2018.06.004.
  21. ^ a b Puspaningrum, Mika R.; van den Bergh, Gerrit D.; Chivas, Allan R.; Setiabudi, Erick; Kurniawan, Iwan (January 2020). "Isotopic reconstruction of Proboscidean habitats and diets on Java since the Early Pleistocene: Implications for adaptation and extinction". Quaternary Science Reviews. 228: 106007. Bibcode:2020QSRv..22806007P. doi:10.1016/j.quascirev.2019.106007. S2CID 212876762.
  22. ^ a b Saarinen, Juha; Lister, Adrian M. (14 August 2023). "Fluctuating climate and dietary innovation drove ratcheted evolution of proboscidean dental traits". Nature Ecology & Evolution. 7 (9): 1490–1502. Bibcode:2023NatEE...7.1490S. doi:10.1038/s41559-023-02151-4. ISSN 2397-334X. PMC 10482678. PMID 37580434.
  23. ^ Matsukawa, Masaki; Shibata, Kenichiro (2 October 2015). "Review of Japanese Cenozoic (Miocene–Modern) Vertebrate Tracks". Ichnos. 22 (3–4): 261–290. Bibcode:2015Ichno..22..261M. doi:10.1080/10420940.2015.1064407. ISSN 1042-0940. S2CID 129206332.
  24. ^ Diamond, Jared M. (1987). "Did Komodo dragons evolve to eat pygmy elephants?". Nature. 326 (6116): 832.
  25. ^ a b Shoshani, J.; Tassy, P. (2005). "Advances in proboscidean taxonomy & classification, anatomy & physiology, and ecology & behavior". Quaternary International. 126–128: 5–20. Bibcode:2005QuInt.126....5S. doi:10.1016/j.quaint.2004.04.011.
  26. ^ Wu, Yan; Deng, Tao; Hu, Yaowu; Ma, Jiao; Zhou, Xinying; Mao, Limi; Zhang, Hanwen; Ye, Jie; Wang, Shi-Qi (16 May 2018). "A grazing Gomphotherium in Middle Miocene Central Asia, 10 million years prior to the origin of the Elephantidae". Scientific Reports. 8 (1): 7640. Bibcode:2018NatSR...8.7640W. doi:10.1038/s41598-018-25909-4. ISSN 2045-2322. PMC 5956065. PMID 29769581.
  27. ^ Konidaris, George E.; Tsoukala, Evangelia (2022), Vlachos, Evangelos (ed.), "The Fossil Record of the Neogene Proboscidea (Mammalia) in Greece", Fossil Vertebrates of Greece Vol. 1, Cham: Springer International Publishing, pp. 299–344, doi:10.1007/978-3-030-68398-6_12, ISBN 978-3-030-68397-9, S2CID 245023119, retrieved 28 February 2023
  28. ^ Lister, Adrian M.; Dirks, Wendy; Assaf, Amnon; Chazan, Michael; Goldberg, Paul; Applbaum, Yaakov H.; Greenbaum, Nathalie; Horwitz, Liora Kolska (September 2013). "New fossil remains of Elephas from the southern Levant: Implications for the evolutionary history of the Asian elephant". Palaeogeography, Palaeoclimatology, Palaeoecology. 386: 119–130. Bibcode:2013PPP...386..119L. doi:10.1016/j.palaeo.2013.05.013.
  29. ^ Haynes, Gary (March 2022). "Late Quaternary Proboscidean Sites in Africa and Eurasia with Possible or Probable Evidence for Hominin Involvement". Quaternary. 5 (1): 18. doi:10.3390/quat5010018. ISSN 2571-550X.
  30. ^ Wei, Guangbiao; He, Cunding; Hu, Yue; Yu, Kefu; Chen, Shaokun; Pang, Libo; Wu, Yan; Huang, Wanbo; Yuan, Wenge (April 2017). "First discovery of a bone handaxe in China". Quaternary International. 434: 121–128. Bibcode:2017QuInt.434..121W. doi:10.1016/j.quaint.2014.12.022.
  31. ^ Zhang, Jia-Fu; Huang, Wei-Wen; Hu, Yue; Yang, Shi-Xia; Zhou, Li-Ping (October 2015). "Optical dating of flowstone and silty carbonate-rich sediments from Panxian Dadong Cave, Guizhou, southwestern China". Quaternary Geochronology. 30: 479–486. Bibcode:2015QuGeo..30..479Z. doi:10.1016/j.quageo.2015.01.011.
  32. ^ Schepartz, L.A.; Stoutamire, S.; Bekken, D.A. (January 2005). "Stegodon orientalis from Panxian Dadong, a Middle Pleistocene archaeological site in Guizhou, South China: taphonomy, population structure and evidence for human interactions". Quaternary International. 126–128: 271–282. doi:10.1016/j.quaint.2004.04.026.
  33. ^ Peng, Hongxia; Ma, Zhibang; Huang, Wanpo; Gao, Jing (December 2014). "230Th/U chronology of a paleolithic site at Xinglong Cave in the three-Gorge region of south China". Quaternary Geochronology. 24: 1–9. doi:10.1016/j.quageo.2014.07.001.
  34. ^ Zhang, Yue; Stiner, Mary C.; Dennell, Robin; Wang, Chunxue; Zhang, Shuangquan; Gao, Xing (August 2010). "Zooarchaeological perspectives on the Chinese Early and Late Paleolithic from the Ma'anshan site (Guizhou, South China)". Journal of Archaeological Science. 37 (8): 2066–2077. Bibcode:2010JArSc..37.2066Z. doi:10.1016/j.jas.2010.03.012.
  35. ^ Matthew W. Tocheri, Thomas Sutikna, Jatmiko, E. Wahyu Saptomo (14 February 2022), "Homo floresiensis", The Oxford Handbook of Early Southeast Asia, Oxford University Press, pp. 38–69, doi:10.1093/oxfordhb/9780199355358.013.2, ISBN 978-0-19-935535-8, retrieved 13 March 2023{{citation}}: CS1 maint: multiple names: authors list (link)
  36. ^ Sutikna, Thomas; Tocheri, Matthew W.; Faith, J. Tyler; Jatmiko; Due Awe, Rokus; Meijer, Hanneke J.M.; Wahyu Saptomo, E.; Roberts, Richard G. (November 2018). "The spatio-temporal distribution of archaeological and faunal finds at Liang Bua (Flores, Indonesia) in light of the revised chronology for Homo floresiensis". Journal of Human Evolution. 124: 52–74. Bibcode:2018JHumE.124...52S. doi:10.1016/j.jhevol.2018.07.001. PMID 30173885.
  37. ^ van den Bergh, Gerrit D.; Alloway, Brent V.; Storey, Michael; Setiawan, Ruly; Yurnaldi, Dida; Kurniawan, Iwan; Moore, Mark W.; Jatmiko; Brumm, Adam; Flude, Stephanie; Sutikna, Thomas; Setiyabudi, Erick; Prasetyo, Unggul W.; Puspaningrum, Mika R.; Yoga, Ifan (October 2022). "An integrative geochronological framework for the Pleistocene So'a basin (Flores, Indonesia), and its implications for faunal turnover and hominin arrival". Quaternary Science Reviews. 294: 107721. Bibcode:2022QSRv..29407721V. doi:10.1016/j.quascirev.2022.107721. hdl:10072/418777. S2CID 252290750.
  38. ^ a b Jukar, A. M.; Lyons, S. K.; Wagner, P. J.; Uhen, M. D. (15 January 2021). "Late Quaternary extinctions in the Indian Subcontinent". Palaeogeography, Palaeoclimatology, Palaeoecology. 562: 110137. Bibcode:2021PPP...56210137J. doi:10.1016/j.palaeo.2020.110137. ISSN 0031-0182.
  39. ^ a b Samuel T. Turvey, Haowen Tong, Anthony J. Stuart and Adrian M. Lister (2013). "Holocene survival of Late Pleistocene megafauna in China: a critical review of the evidence". Quaternary Science Reviews. 76: 156–166. Bibcode:2013QSRv...76..156T. doi:10.1016/j.quascirev.2013.06.030.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  40. ^ Turvey, Samuel T.; Sathe, Vijay; Crees, Jennifer J.; Jukar, Advait M.; Chakraborty, Prateek; Lister, Adrian M. (January 2021). "Late Quaternary megafaunal extinctions in India: How much do we know?". Quaternary Science Reviews. 252: 106740. Bibcode:2021QSRv..25206740T. doi:10.1016/j.quascirev.2020.106740. S2CID 234265221.
  41. ^ H. Saegusa, "Comparisons of Stegodon and Elephantid Abundances in the Late Pleistocene of Southern China" Archived 2006-05-08 at the Wayback Machine, The World of Elephants – Second International Congress, (Rome, 2001), 345–349.
Retrieved from "https://en.wikipedia.org/w/index.php?title=Stegodon&oldid=1243693515"