The evolutionary ancestry of arthropods dates back to the Cambrian period. The group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians (or their constituent clades) in a superphylum Ecdysozoa. Overall, however, the basal relationships of animals are not yet well resolved. Likewise, the relationships between various arthropod groups are still actively debated. Today, arthropods contribute to the human food supply both directly as food, and more importantly, indirectly as pollinators of crops. Some species are known to spread severe disease to humans, livestock, and crops. (Full article...)
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Five anatomical images of a Drosophila subobscura (male) on a rotting squash, located near Christchurch Park, Ipswich
Drosophila subobscura is a species of fruit fly in the family Drosophilidae. Originally found around the Mediterranean, it has spread to most of Europe and the Near East. It has been introduced into the west coasts of Canada, the United States, and Chile. Its closest relative is Drosophila madeirensis, found in the Madeira Islands, followed by D. guanche, found in the Canary Islands. These three species form the D. subobscura species subgroup. When they mate, males and females perform an elaborate courtship dance, in which the female can either turn away to end the mating ritual, or stick out her proboscis in response to the male's, allowing copulation to proceed. D. subobscura has been regarded as a model organism for its use in evolutionary-biological studies. (Full article...)
Horse-flies and deer flies are true flies in the family Tabanidae in the insectorderDiptera. The adults are often large and agile in flight. Only female horseflies bite land vertebrates, including humans, to obtain blood. They prefer to fly in sunlight, avoiding dark and shady areas, and are inactive at night. They are found all over the world except for some islands and the polar regions (Hawaii, Greenland, Iceland). Both horse-flies and botflies (Oestridae) are sometimes referred to as gadflies.
Adult horse-flies feed on nectar and plant exudates; males have weak mouthparts, but females have mouthparts strong enough to bite large animals. This is for the purpose of obtaining enough protein from blood to produce eggs. The mouthparts of females are formed into a stout stabbing organ with two pairs of sharp cutting blades, and a spongelike part used to lap up the blood that flows from the wound. The larvae are predaceous and grow in semiaquatic habitats.
The species and others in its family were used in traditional apothecary preparations as "Cantharides". The insect is the source of the terpenoidcantharidin, a toxic blistering agent once used as an exfoliating agent, anti-rheumatic drug and an aphrodisiac. The substance has also found culinary use in some blends of the North African spice mix ras el hanout. Its various supposed benefits have been responsible for accidental poisonings. (Full article...)
Puzzled by the geographical distribution of wildlife and fossils he collected on the voyage, Darwin began detailed investigations and, in 1838, devised his theory of natural selection. Although he discussed his ideas with several naturalists, he needed time for extensive research, and his geological work had priority. He was writing up his theory in 1858 when Alfred Russel Wallace sent him an essay that described the same idea, prompting the immediate joint submission of both their theories to the Linnean Society of London. Darwin's work established evolutionary descent with modification as the dominant scientific explanation of natural diversification. In 1871, he examined human evolution and sexual selection in The Descent of Man, and Selection in Relation to Sex, followed by The Expression of the Emotions in Man and Animals (1872). His research on plants was published in a series of books, and in his final book, The Formation of Vegetable Mould, through the Actions of Worms (1881), he examined earthworms and their effect on soil. (Full article...)
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An individual in its burrow
Neotrypaea californiensis (formerly Callianassa californiensis), the Bay ghost shrimp, is a species of ghost shrimp that lives on the Pacific coast of North America. It is a pale animal which grows to a length of 11.5 cm (4.5 in). One claw is bigger than the other, especially in males, and the enlarged claw is thought to have a function in mating. N. californiensis is a deposit feeder that lives in extensive burrow systems, and is responsible for high rates of bioturbation. It adversely affects oyster farms, and its numbers are controlled in some places by the application of pesticides. It carries out an important role in the ecosystem, and is used by fishermen as bait. (Full article...)
A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genusBombus, part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related genera (e.g., Calyptapis) are known from fossils. They are found primarily in higher altitudes or latitudes in the Northern Hemisphere, although they are also found in South America, where a few lowland tropical species have been identified. European bumblebees have also been introduced to New Zealand and Tasmania. Female bumblebees can sting repeatedly, but generally ignore humans and other animals.
Most bumblebees are social insects that form colonies with a single queen. The colonies are smaller than those of honey bees, growing to as few as 50 individuals in a nest. Cuckoo bumblebees are brood parasitic and do not make nests or form colonies; their queens aggressively invade the nests of other bumblebee species, kill the resident queens and then lay their own eggs, which are cared for by the resident workers. Cuckoo bumblebees were previously classified as a separate genus, but are now usually treated as members of Bombus.
Bumblebees have round bodies covered in soft hair (long branched setae) called 'pile', making them appear and feel fuzzy. They have aposematic (warning) coloration, often consisting of contrasting bands of colour, and different species of bumblebee in a region often resemble each other in mutually protective Müllerian mimicry. Harmless insects such as hoverflies often derive protection from resembling bumblebees, in Batesian mimicry, and may be confused with them. Nest-making bumblebees can be distinguished from similarly large, fuzzy cuckoo bumblebees by the form of the female hind leg. In nesting bumblebees, it is modified to form a pollen basket, a bare shiny area surrounded by a fringe of hairs used to transport pollen, whereas in cuckoo bumblebees, the hind leg is hairy all round, and they never carry pollen. (Full article...)
Mysida is an order of small, shrimp-like crustaceans in the malacostracansuperorderPeracarida. Their common nameopossum shrimps stems from the presence of a brood pouch or "marsupium" in females. The fact that the larvae are reared in this pouch and are not free-swimming characterises the order. The mysid's head bears a pair of stalked eyes and two pairs of antennae. The thorax consists of eight segments each bearing branching limbs, the whole concealed beneath a protective carapace and the abdomen has six segments and usually further small limbs.
His class syllabus on zoology was originally designed for students at Berkeley, but were later published as an invertebrate zoology textbook and field guide, becoming the first compendium of marine invertebrates in the north central California coastal region for specialists working in the area between Hopkins Marine Station and Bodega Marine Laboratory. After Light's death, the book was edited, revised, and expanded by Ralph I. Smith and other contributors, becoming known as Light's Manual. After Smith himself died, the book was renamed the Light and Smith Manual in his honor. (Full article...)
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Portia schultzi is a species of jumping spider which ranges from South Africa in the south to Kenya in the north, and also is found in West Africa and Madagascar. In this species, which is slightly smaller than some other species of the genusPortia, the bodies of females are 5 to 7 mm (0.20 to 0.28 in) long, while those of males are 4 to 6 mm (0.16 to 0.24 in) long. The carapaces of both sexes are orange-brown with dark brown mottling, and covered with dark brown and whitish hairs lying over the surface. Males have white tufts on their thoraces and a broad white band above the bases of the legs, and these features are less conspicuous in females. Both sexes have tufts of orange to dark orange above the eyes, which are fringed with pale orange hairs. Males' abdomens are yellow-orange to orange-brown with blackish mottling, and on the upper sides are black and light orange hairs, and nine white tufts. Those of females are pale yellow and have black markings with scattered white and orange-brown hairs on the upper side. P. schultzi has relatively longer legs than other Portia, and a "lolloping" gait.
While most jumping spiders focus accurately up to about 75 cm (30 in) away, P. schultzi responds to a maximum of about 10 cm (3.9 in) in good light, and ignores everything in very subdued light. For prey, P. schultzi prefers web-based spiders, then jumping spiders, and finally insects. The females of P. schultzi and other Portia species build "capture webs" to catch prey, and often join their own webs on to web-based spiders to catch the other spiders or their prey.
If a P. schultzi female is mature, a male P. schultzi will try to copulate with her, or cohabit with a subadult female and copulate while she is moulting. They usually mate on a web or on a dragline made by the female, and P. schultzi typically copulates for about 100 seconds, while others in the genus can take several minutes or even several hours. Females try to kill and eat their mates during or after copulation, and subadult females mimic adult females to attract males as prey. Contests between Portia females are violent, and embraces in P. schultzi typically take 20 to 60 seconds. Sometimes, one female knocks the other on her back and the other may be killed and eaten if she does not right herself quickly and run away. When hunting, P. schultzi mature females emit olfactory signals that reduce the risk that any other females, males, or juveniles of the same species may contend for the same prey. (Full article...)
Nanahughmilleria is classified in the family Adelophthalmidae, the only clade in the superfamily Adelophthalmoidea. This clade was characterised by their small size, their parabolic (approximately U-shaped) carapaces and the presence of epimera (lateral "extensions" of the segment) on the seventh segment, among others. Nanahughmilleria was different from its relatives by the presence of more spines in its appendages (limbs) and by its genital morphology. The largest species confidently assigned to the genus was N. norvegica at 10 cm (3.9 in), making it a comparatively small eurypterid. (Full article...)
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Polistes exclamans, the Guinea paper wasp, is a socialwasp and is part of the family Vespidae of the order Hymenoptera. It is found throughout the United States, Mexico, the Bahamas, Jamaica and parts of Canada. Due to solitary nest founding by queens, P. exclamans has extended its range in the past few decades and now covers the eastern half of the United States, as well as part of the north. This expansion is typically attributed to changing global climate and temperatures. P. exclamans has three specific castes, including males, workers, and queens, but the dominance hierarchy is further distinguished by age. The older the wasp is, the higher it is in ranking within the colony. In most P. exclamans nests, there is one queen who lays all the eggs in the colony. The physiological similarities between the worker and queen castes have led to experiments attempting to distinguish the characteristics of these two castes and how they are determined, though males have easily identifiable physiological characteristics. Since P. exclamans live in relatively small, open combed nests, they are often subject to predators and parasites, such as Chalcoela iphitalis, Elasmus polistis, and birds. P. exclamans have defense and recognition strategies that help protect against these predators and parasites. (Full article...)
Mayflies have ancestral traits that were probably present in the first flying insects, such as long tails and wings that do not fold flat over the abdomen. Their immature stages are aquatic fresh water forms (called "naiads" or "nymphs"), whose presence indicates a clean, unpolluted and highly oxygenated aquatic environment. They are unique among insect orders in having a fully winged terrestrial preadult stage, the subimago, which moults into a sexually mature adult, the imago.
Mayflies "hatch" (emerge as adults) from spring to autumn, not necessarily in May, in enormous numbers. Some hatches attract tourists. Fly fishermen make use of mayfly hatches by choosing artificial fishing flies that resemble them. One of the most famous English mayflies is Rhithrogena germanica, the fisherman's "March brown mayfly". (Full article...)
Metanephrops challengeri (commonly known as the New Zealand lobster or New Zealand scampi) is a species of slim, pink lobster that lives around the coast of New Zealand. It is typically 13–18 cm (5–7 in) long and weighs around 100 g (3.5 oz). The carapace and abdomen are smooth, and adults are white with pink and brown markings and a conspicuous pair of long, slim claws. M. challengeri lives in burrows at depths of 140–640 m (460–2,100 ft) in a variety of sediments. Although individuals can live for up to 15 years, the species shows low fecundity, where small numbers of larvae hatch at an advanced stage.
M. challengeri is a significant prey item for ling, as well as being an important fishery species for human consumption; trawlers catch around 1,000 t (2,200,000 lb) per year under the limitations of New Zealand's Quota Management System. The species was first collected by the Challenger expedition of 1872–1876, but only described as separate from related species by Heinrich Balss in 1914. Although originally classified in the genus Nephrops, it was moved in 1972 to a new genus, Metanephrops, along with most other species then classified in Nephrops. (Full article...)
Myrmeciites is an extinct form genus of bulldog ants in the subfamily Myrmeciinae of the family Formicidae, which contains three described species and two fossils not placed beyond the genus level. Described in 2006 from Ypresian stage (Early Eocene) deposits, all three of the described species and one unplaced fossil are from British Columbia, Canada, while the second unplaced fossil is from Washington State, USA. These ants were large, with the largest specimens collected reaching 3 centimetres (1.2 in). The behaviour of these ants would have been similar to extant Myrmeciinae ants, such as solitary foraging, nesting either in the soil or trees, and leaving no pheromone trail to food sources. Due to the poor preservation of these ants, their phylogenetic position among Myrmeciinae is unclear, and no type species has been designated. These ants are classified as incertae sedis in Myrmeciinae, but some writers have classified it as incertae sedis within the insect order Hymenoptera. This reclassification, however, has not been accepted; instead, Myrmeciites remains in Myrmeciinae. (Full article...)
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A. decolor
Ameles decolor (also known as the Mediterranean dwarf mantis or the dwarf mantis) is a species of small praying mantis native to the west Mediterranean and North Africa. A. decolor was first described by entomologist Domenico Cyrillo in 1787, and its current classification was established in 1976 by Karl Harz and Alfred Peter Kaltenbach. A. decolor presents as a small, light brown mantis with females tending to appear larger than their male counterparts. The mating patterns of A. decolor are considered some of the most complex amongst praying mantises, with males presenting two different styles of courtship. Their habitat favours shrublands, grasslands, and wooded areas. (Full article...)
... that the Australian jumping spider Portia fimbriata plays a deadly game of hide-and-seek with its favourite prey, Jacksonoides queenlandicus, another jumping spider?
Image 3Body structure of a typical crustacean – krill (from Crustacean)
Image 4Mature queen of a termite colony, showing how the unsclerotised cuticle stretches between the dark sclerites that failed to stretch as the abdomen grew to accommodate her ovaries (from Arthropod exoskeleton)
Image 5 This fully-grown robber crab has tough fabric forming its joints, delicate biomineralized cuticle over its sensory antennae, optic-quality over its eyes, and strong, calcite-reinforced chitin armouring its body and legs; its pincers can break into coconuts (from Arthropod exoskeleton)
Image 7 Honeybee larvae have flexible but delicate unsclerotised cuticles. (from Arthropod exoskeleton)
Image 8In honeypot antrepletes, the abdomens of the workers that hold the sugar solution grow vastly, but only the unsclerotised cuticle can stretch, leaving the unstretched sclerites as dark islands on the clear abdomen (from Arthropod exoskeleton)
Image 10Formation of anterior segments across arthropod taxa based on gene expression and neuroanatomical observations, Note the chelicera(Ch) and chelifore(Chf) arose from somite 1 and thus correspond to the first antenna(An/An1) of other arthropods. (from Chelicerata)
Image 12The house centipedeScutigera coleoptrata has rigid sclerites on each body segment. Supple chitin holds the sclerites together and connects the segments flexibly. Similar chitin connects the joints in the legs. Sclerotised tubular leg segments house the leg muscles, their nerves and attachments, leaving room for the passage of blood to and from the hemocoel (from Arthropod exoskeleton)
Image 16Ghost crab, showing a variety of integument types in its exoskeleton, with transparent biomineralization over the eyes, strong biomineralization over the pincers, and tough chitin fabric in the joints and the bristles on the legs (from Arthropod exoskeleton)
Image 17Some of the various hypotheses of myriapod phylogeny. Morphological studies (trees a and b) support a sister grouping of Diplopoda and Pauropoda, while studies of DNA or amino acid similarities suggest a variety of different relationships, including the relationship of Pauropoda and Symphyla in tree c. (from Myriapoda)
Image 28This Zoea-stage larva is hardly recognisable as a crab, but each time it sheds its cuticle it remodels itself, eventually taking on its final crab form (from Arthropod exoskeleton)
The red rock crab (Grapsus grapsus), also known as "Sally Lightfoot", is one of the commonest crabs along the western seaboard of the Americas. John Steinbeck wrote of them, "Everyone who has seen them has been delighted with them ... These little crabs, with brilliant cloisonné carapaces, walk on their tiptoes, they have remarkable eyes and an extremely fast reaction time." He tried to catch them but to little avail. "If you walk slowly, they move slowly ahead of you in droves. If you hurry, they hurry. When you plunge at them, they seem to disappear in a puff of blue smoke."
Argiope trifasciata, the banded garden or banded orb-weaving spider, is a species of arachnid in the family Araneidae. It is native to North and South America but has spread to other parts of the world. This ventral view of a female A. trifasciata shows her in the centre of her web, which can reach a diameter of 60 cm (24 in). The function of the zig-zag web decorations is unclear, but they may serve to make the spider appear larger or to act as a warning sign.
The light blue soldier crab (Mictyris longicarpus) inhabits beaches in the Indo-Pacific region. Soldier crabs filter sand or mud for microorganisms. They congregate during the low tide, and bury themselves in a corkscrew pattern during high tide, or whenever they are threatened.
Xylotrupes socrates (Siamese rhinoceros beetle, or "fighting beetle"), male, on a banana leaf. This scarab beetle is particularly known for its role in insect fighting in Northern Laos and Thailand.
Gonepteryx rhamni, also known as the common brimstone, is a butterfly in the family Pieridae. It lives throughout the Palearctic zone and is commonly found across Europe, Asia and North Africa. The butterfly relies on two species of buckthorn as hosts for its eggs and larvae, which influences its geographic range and distribution as these plants are commonly found in wetlands. After spending the summer feeding, adults travel to woodland areas to spend seven months hibernating. In spring when their host plants have developed, they return to the wetlands to breed and lay eggs. Both the larval and adult forms of the species have protective coloration and behaviour that decreases their chances of being recognised and preyed upon. The adult common brimstone has sexual dichromism in its wing coloration and iridescence; the male (pictured) has yellow wings and iridescence, while females have greenish-white wings and are not iridescent.
The orb-weaver spiders (family Araneidae) are the familiar builders of spiral wheel-shaped webs often found in gardens, fields and forests. The family is a large one, including over 2800 species in over 160 genera worldwide, making it the third largest known (behind Salticidae and Linyphiidae). The web has always been thought of as an engineering marvel.
The yellow fever mosquito (Aedes aegypti) is a species of mosquito known for its ability to spread yellow fever and dengue fever. The mosquito can be recognized by the white markings on its legs and a marking in the form of a lyre on its thorax. Though originally from Africa, the yellow fever mosquito can now be found in tropical regions around the world.
A Eusthenia species of stonefly. The order contains almost 3,500 known species, including the only known insects that are exclusively aquatic from birth to death. Stoneflies are believed to be one of the most primitive groups of Neoptera and are found worldwide, with the exception of Antarctica.
A female subimago of a March Brown mayfly (Rhithrogena germanica). Mayflies belong to the orderEphemeroptera, and the only insects that have a subimago phase. This stage is a favourite food of many fish, and many fishing flies are modeled to resemble them. They are aquatic insects whose nymph stage usually lasts one year in freshwater. The adults are short-lived, from as little as thirty minutes to a few days depending on the species.
The monarch butterfly (Danaus plexippus) is one of the best-known species of butterfly. Its wings feature an easily recognizable orange and black pattern, with a wingspan of 8.9–10.2 centimetres (3½–4 in).
The longtail tadpole shrimp (Triops longicaudatus) is a freshwater crustacean resembling a miniature horseshoe crab. It is one of the oldest animal species still in existence. Like its relative Triops cancriformis, the longtail tadpole shrimp is considered a living fossil because its basic prehistoric morphology has changed little in the last 70 million years, exactly matching ancient fossils.
Papilio polymnestor, the blue Mormon, is a species of swallowtail butterfly found in southern India and Sri Lanka. It is a woodland species, often seen on forest paths and near streams. The larvae feed on trees in the family Rutaceae, such as citrus. Young larvae are green with white markings and position themselves on the upper surface of leaves, relying on their cryptic colouring, which resembles bird droppings, for protection. Older larvae seek less conspicuous locations, and have a unique habit of securing their balance by weaving silk on the substratum. This adult male P. polymnestor butterfly was photographed in the Indian state of Kerala.
The Ozyptila praticola species of crab spider is found throughout Europe and the Middle East. They do not build webs to trap prey, but are active hunters. Crab spiders are so named because of their first two pairs of legs, which are held out to the side giving them a crab-like appearance. Also, like crabs, these spiders move sideways and backwards more easily than forwards.
A fishing spider with two of its legs missing. Most likely some predator (a bird, or given its habitat a large fish or frog) grasped the spider by the missing limbs which were jettisoned by the spider in response, a process known as autotomy.
A male Roesel's bush-cricket (Metrioptera roeseli), a European bush-cricket named after August Johann Rösel von Rosenhof, a German entomologist. Its song is very similar to that of Savi's Warbler. Its body length as an adult insect is 15 to 18 mm. It is brown with a pale margin to the sides of the pronotum. Its forewings usually reach midway along its abdomen at rest. However there is a macropterous form of this insect (f. diluta), in which the wings reach beyond the tip of the abdomen. This form appears predominantly during hot summers and enables the species to extend its geographical range rapidly while conditions are suitable; such migrations may also be in response to local overpopulation.
Zonocerus variegatus, the variegated grasshopper, is a species of grasshopper in the family Pyrgomorphidae native to tropical western and central Africa. It feeds on a wide variety of plant foods and causes damage to crops, particularly cassava, groundnuts and vegetables, as well as transmitting diseases caused by mosaic viruses between plants. This Z. variegatus grasshopper was photographed in the Bobiri Forest in Ghana.
Sexual dimorphism is the condition where the two sexes of the same species exhibit different characteristics. Differences may include secondary sex characteristics, size, weight, color, or markings, as well as behavioral and cognitive differences. In the butterfly species Colias dimera (also known as the Dimera sulphur), seen here mating in Venezuela, the male on the right is a brighter shade of yellow than the female.
A honey bee extracts nectar from a flower using its proboscis. Tiny hairs covering the bee's body maintain a slight electrostatic charge, causing pollen from the flower's anthers to stick to the bee's hairs, allowing for pollination when the bee moves on to another flower.
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