Achilidae

Achilidae Temporal range:
Achilus flammeus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Hemiptera
Suborder:	Auchenorrhyncha
Infraorder:	Fulgoromorpha
Superfamily:	Fulgoroidea
Family:	Achilidae Stål, 1866

Achilidae is a family of planthoppers, sometimes called "achilids" in the order Hemiptera. There are at least 520 described species in Achilidae.

Subfamilies and Genera

Fulgoromorpha Lists on the Web (FLOW) includes 3 subfamilies:

Achilinae

Cixidia opaca

Authority: Stål, 1866 - 2 tribes

Achilini Stål, 1866

• Achilina Stål, 1866

1. Achilus Kirby, 1819 - type genus
2. Flatachilus Fennah, 1950
3. Olmiana Guglielmino, Bückle & Emeljanov, 2010
4. Ouwea Distant, 1907

• Cixidiina Emeljanov, 1992

1. †Angustachilus Lefebvre, Bourgoin & Nel, 2007
2. Cixidia Fieber, 1866

• Elidipterina Fennah, 1950

1. Booneta Distant, 1907
2. Catonidia Uhler, 1896
3. Elidiptera Spinola, 1839
4. Faventilla Metcalf, 1948
5. Messeis (planthopper) Stål, 1860
6. Metaphradmon Fennah, 1950
7. Paracatonidia – P. webbeda Long, Yang & Chen, 2015
8. Paraphradmon Fennah, 1950
9. Parelidiptera Fennah, 1950
10. Phradmonicus Emeljanov, 1991
11. Prinoessa Fennah, 1950
12. Uniptera Ball, 1933

• Achilini incertae sedis

1. Anabunda Emeljanov, 2005
2. Aneipo Kirkaldy, 1906
3. Bunduica (planthopper) Jacobi, 1909
4. Dipsiathus Emeljanov, 2005
5. Epiona (planthopper) Emeljanov, 2005
6. Mabira (planthopper) Fennah, 1950
7. Nelidia Stål, 1860
8. Parabunda Emeljanov, 2005
9. †Paratesum Emeljanov & Shcherbakov, 2009
10. †Protomenocria Emeljanov & Shcherbakov, 2009
11. †Psycheona Emeljanov & Shcherbakov, 2009
12. Rhinochloris Emeljanov, 2005
13. †Gedanochila Brysz & Szwedo, 2022
14. †Protepiptera Usinger, 1939

Achillini Emeljanov, 1991

1. Achilla (planthopper) Haglund, 1899
2. †Hooleya (planthopper) Cockerell, 1922
3. Maurisca Emeljanov, 2005

Apatesoninae

Authority: Metcalf, 1938 - 4 monotypic tribes

1. Apateson Fowler, 1900: Apatesonini Metcalf, 1938
2. Ilva (planthopper) Stål, 1866: Ilvini Emeljanov, 1991
3. Sevia Stål, 1866: Seviini Emeljanov, 1991
4. Tropiphlepsia Muir, 1924: Tropiphlepsiini Emeljanov, 1991

Myconinae

Agandecca annectans male

Synecdoche impunctata

Authority: Fennah, 1950 - 6 tribes

Amphignomini Emeljanov, 1991

1. Amphignoma Emeljanov, 1991

Mycarini Emeljanov, 1991

1. Acocarinus Emeljanov, 1991
2. Emeljanocarinus Bourgoin & Soulier-Perkins, 2006
3. Katbergella Fennah, 1950
4. Mycarinus Emeljanov, 1991
5. Mycarus Emeljanov, 1991

Myconini Fennah, 1950

1. Ganachilla – G. zhenyuanensis Wang & Huang, 1989
2. Haicixidia – H. jianfengensis Wang, 1989
3. Myconellus Fennah, 1950
4. Myconus (planthopper) Stål, 1860
5. Myrophenges Fennah, 1965

Plectoderini Fennah, 1950. Selected genera

• Abas (planthopper) Fennah, 1950
• Callichlamys (planthopper) Kirkaldy, 1907
• Cernea (planthopper) Williams, 1977
• Cnidus (planthopper) Stål, 1866
• Cythna (planthopper) Kirkaldy, 1906
• Deferunda Distant, 1912
• Francesca (planthopper) Kirkaldy, 1906
• Gordiacea (planthopper) Metcalf, 1948
• Haitiana Dozier, 1936
• Hamba (planthopper) Distant, 1907
• Kosalya (planthopper) Distant, 1906
• Kurandella Fennah, 1950
• Magadha (planthopper) Distant, 1906
• Mahuna Distant, 1907
• Nyonga (planthopper) Synave, 1959
• Opsiplanon Fennah, 1945
• Plectoderes Spinola, 1839
• Quadrana Caldwell, 1951
• Rhinocolura (planthopper) Fennah, 1950
• Spino (planthopper) Fennah, 1950
• Synecdoche (planthopper) O'Brien, 1971
• Taloka (planthopper) Distant, 1907
• Usana (planthopper) Distant, 1906

Rhotalini Fennah, 1950

1. Errada Walker, 1868
2. Errotasa Emeljanov, 2005
3. Hebrotasa Melichar, 1915
4. Rhotala Walker, 1857
5. Taractellus Metcalf, 1948

Waghildini† Szwedo, 2006

1. Waghilde Szwedo, 2006

incertae sedis

1. †Acixiites Hamilton, 1990
2. Leptarciella Fennah, 1958
3. †Niryasaburnia Szwedo, 2004
4. Parasabecoides Synave, 1965
5. Peltatavertexalis – P. horizontalis Xu, Long & Chen, 2019
6. Ridesa Schumacher, 1915
7. Sabecoides Fennah, 1958

To be placed

GBIF also includes the following genera that may have doubtful placement:

1. Chiotasa Melichar, 1915
2. Cionoderus Uhler, 1895
3. Messoides Metcalf, 1930
4. Nablusitypus Kaddumi, 2005
5. Neomenocria Fennah, 1950
6. Okatropis Matsumura, 1910
7. Planusfrons Chun Liang Chen, Chung Tu Yang & Wilson, 1989
8. Plectoderella Fennah, 1950
9. Pyren Fennah, 1950
10. Rhotella Metcalf, 1938
11. Spendon Jacobi, 1928
12. Tabiana Jacobi, 1928
13. Tudea Distant, 1907
14. Winawa Haupt, 1926

Further reading

• List of Achilidae genera

• Bantock, T.; Botting, J. (2013). "British Bugs, an online identification guide to UK Hemiptera". Retrieved 2018-04-06.
• Bartlett, C.R. (2012). "Planthoppers of North America". Retrieved 2018-04-06.
• Borror, Donald J.; Peterson, Roger Tory; White, Richard E. (1998). A Field Guide to Insects. Houghton Mifflin. ISBN 978-0395911709.
• Fennah, R.G. (1969). "Fulgoroidea (Homoptera) from New Caledonia and the Loyalty Islands". Pacific Insects Monographs. 21 (3): 259–260. doi:10.1093/besa/15.3.259b.
• Fennah, R.G. (1956). "Fulgoroidea from southern China". Proceedings of the California Academy of Sciences. Fourth Series. 28 (13): 441–527.
• Gillott, Cedric (1980). Entomology. Plenum Press. ISBN 0-306-40366-8.
• Kellogg, Vernon L. (1905). American insects. H. Holt.
• Majka, C. (2009). "Thomas L. Casey and Rhode Island". ZooKeys (22): 267–283. doi:10.3897/zookeys.22.93.
• Metcalf, Z.P. (1947). General Catalogue of the Hemiptera, Fascicle IV: Fulgoroidea, Part 9: Fulgoridae. Smith College.
• Misof, B.; Liu, S.; Meusemann, K.; Peters, R.S.; et al. (2014). "Phylogenomics resolves the timing and pattern of insect evolution". Science. 346 (6210): 763–767. doi:10.1126/science.1257570. PMID 25378627. S2CID 36008925.
• Urban, Julie M.; Cryan, Jason R. (2007). "Evolution of the planthoppers (Insecta: Hemiptera: Fulgoroidea)". Molecular Phylogenetics and Evolution. 42 (2): 556–572. doi:10.1016/j.ympev.2006.08.009. ISSN 1055-7903. PMID 17011797.
• Zimmerman, Elwood C. (1948). Insects of Hawaii: A Manual of the Insects of the Hawaiian Islands, including an Enumeration of the Species and Notes on their Origin, Distribution, Hosts, Parasites, etc. Vol. 4. University of Hawaii Press.

External links

• Media related to Achilidae at Wikimedia Commons