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Leptosuchus

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Leptosuchus
Temporal range: Late Triassic, 237–208.5 Ma
L. crosbiensis skull
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauromorpha
Clade: Archosauriformes
Order: †Phytosauria
Family: †Parasuchidae
Clade: †Leptosuchomorpha
Genus: †Leptosuchus
Case, 1922
Type species
†Leptosuchus crosbiensis
Case, 1922
Species
  • †L. crosbiensis Case, 1922
  • †L. studeri Case & White, 1934

Leptosuchus is an extinct genus of leptosuchomorph phytosaur with a complex taxonomical history. Fossils have been found from the Dockum Group and lower Chinle Formation outcropping in Texas, New Mexico, and Arizona, United States, and date back to the Carnian stage of the Late Triassic.[1]

Currently there are believed to be four species of Leptosuchus. All species share in common a similar position of the temporal arch below the skull roof and a posterior process of the squamosal that extends farther than the paroccipital process.[2] The type species is L. crosbiensis, which was named in 1922 on the basis of material found from Texas.[3] L. adamanensis was first described in 1930 as a species of Machaeroprosopus from the Blue Mesa Member of Petrified Forest National Park, along with the other two species, L. lithodendrorum and L. gregorii. It was not until the publication of a 1995 paper on tetrapods of southwestern United States that these species were recognized as belonging to the genus Leptosuchus. However, because of the distinctive size of the rostral crest in L. gregorii, it was assigned to its own genus, Smilosuchus, in that same paper.[2] Despite this, L. gregorii has recently been seen as belonging to Leptosuchus, as it is believed that the large, complete crest was independently developed in this particular species.[4]

The close relation of Leptosuchus and Machaeroprosopus with Rutiodon has led some paleontologists to believe that the prior two were synonymous with Rutiodon, with the latter's name having seniority.[5] Differing features seen in the three phytosaurs have been attributed to sexual dimorphism, differing growth stages, or individual variation, while similarities could be seen in the rostral crest and the position of the nares.[6] Previously it was believed that these were different genera due to the geographical isolation that could be seen in North American phytosaurs; Leptosuchus remains were found primarily in southern localities while Rutiodon remains could be found in more eastern localities.[7] Other studies concluded that it was synonymous with Machaeroprosopus or Phytosaurus.[8] A 1998 study again found Leptosuchus to be congeneric with Rutiodon,[9] but since then another study has suggested that its type species R. carolinensis, while still very similar to Leptosuchus, is synonymous with Angistorhinus.[10]

L. imperfecta is known from a partial skull, UMMP 7523, from same locality as the holotype of L. crosbiensis. It was collected as fragments and reassembled. Long and Murry (1995) considered it to be a nomen nudum, as there was never full formal documentation of this specimen. They referred it to L. adamanensis. Camp (1930) and Ballew (1989) referred it to L. crosbiensis. Stocker (2010) stated that the specimen is phylogenetically redundant and might be synonymous with L. crosbiensis, because its traits are identical to those of L. crosbiensis.[11]

References

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  1. ^ Case, E. C. and White, T. E. (1934). "Two new specimens of phytosaurs from the Upper Triassic of Western Texas". Contributions from the Museum of Paleontology, University of Michigan 1(9):133-142.
  2. ^ a b Long, R. A., and Murry, P. A. (1995). "Late Triassic (Carnian and Norian) tetrapods from the southwestern United States". New Mexico Museum of Natural History and Science Bulletin 4:1-254.
  3. ^ Case, E. C. (1922). "New reptiles and stegocephalians from the Upper Triassic of western Texas?!. Carnegie Institution of Washington Publication 321:1-84.
  4. ^ Irmis, R. B. (2005). "The vertebrate fauna of the Upper Triassic Chinle Formation in Northern Arizona". In: Nesbitt, S. J., Parker, W. G. and Irmis, R. B., eds., Guidebook to the Triassic Formations of the Colorado Plateau in northern Arizona: Geology, Paleontology, and History. Mesa Southwest Museum, Bulletin No. 9.
  5. ^ Ballew, K. L. (1989). "A phylogenetic analysis of Phytosauria from the Late Triassic of the western United States". In: Lucas, S. G. and Hunt, A. P., eds., Dawn of the age of dinosaurs in the American Southwest, pp. 309-339. New Mexico Museum of Natural History. Albuquerque.
  6. ^ Gregory, J. T. (1962). "The relationships of the American phytosaur Rutiodon". American Museum Novitates 2095:1-22.
  7. ^ Camp, C. L. (1930). "A study of phytosaurs, with description of new material from western North America". Memoirs of the University of California 10:161 pp. Berkeley.
  8. ^ Colbert, E. H. (1947). Studies of the phytosaurs Machaeroprosopus and Rutiodon. Bulletin of the American Museum of Natural History 88:53-96.
  9. ^ Lucas, S. G. (1998). Global Triassic tetrapod biostratigraphy and biochronology. Palaeogeography, Palaeoclimatology, Palaeoecology 143:347-384.
  10. ^ Hungerbühler, A. and Sues, H.-D. (2001). Status and phylogenetic relationships of the Late Triassic phytosaur Rutiodon carolinensis. Journal of Vertebrate Paleontology 21(3 suppl.):64A.
  11. ^ Michelle R. Stocker (2010). "A new taxon of phytosaur (Archosauria: Pseudosuchia) from the Late Triassic (Norian) Sonsela Member (Chinle Formation) in Arizona, and a critical reevaluation of Leptosuchus Case, 1922". Palaeontology. 53 (5): 997–1022. doi:10.1111/j.1475-4983.2010.00983.x. S2CID 83536253.
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