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List of largest birds

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The common ostrich is the largest living bird
Aepyornis maximus, one of the largest birds ever

The largest extant species of bird measured by mass is the common ostrich (Struthio camelus), closely followed by the Somali ostrich (Struthio molybdophanes). A male ostrich can reach a height of 2.8 metres (9.2 feet) and weigh over 156.8 kg (346 lb),[1] A mass of 200 kg (440 lb) has been cited for the ostrich but no wild ostriches of this weight have been verified.[2] Ostrich eggs are the largest of any bird, averaging 1.4 kg (3.1 lb).[3]

The largest wingspan of any extant bird is that of the wandering albatross (Diomedea exulans) of the Sub-Antarctic oceans. The largest dimensions found in this species are an approximate head-to-tail length of 1.44 m (4.7 ft) and a wingspan of 3.65 m (12.0 ft).

The largest bird of all time was likely the elephant bird Aepyornis maximus, which was estimated to have weighed 275–1,000 kilograms (610–2,200 lb) and stood at 3 metres (9.8 ft) tall.[4][5]

The largest wingspan of all time likely belonged to Pelagornis sandersi at roughly 5.2 m (17 ft).[6] P. sandersi was also likely the largest bird to ever fly.[7]

Largest extinct birds

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Table of heaviest extinct bird species

[edit]
Rank Binomial name Mass estimate

[kg (lb)]

Family Order Geological age
1 Aepyornis maximus 850 kg (1,870 lb)[5][Note 1] Aepyornithidae (greater elephant birds) Aepyornithiformes (elephant birds) Late Pleistocene-Holocene
2 Dromornis stirtoni 800 kg (1,800 lb)[8][Note 2] Dromornithidae (mihirungs) Anseriformes (waterfowl) Late Miocene
3 Pachystruthio dmanisensis 450 kg (990 lb)[9][Note 3] Struthionidae Struthioniformes (ostriches) Early Pleistocene
4 Brontornis burmeisteri 319 kg (703 lb)[11] Brontornithidae (thunderbirds) Anseriformes[Note 4](waterfowl) Early Miocene
5 Pachystruthio indet 300 kg (660 lb)[12] Struthionidae Struthioniformes (ostriches) Early Pleistocene
6 Dromornis planei 300 kg (660 lb)[13] Dromornithidae (mihirungs) Anseriformes (waterfowl) Middle Miocene
7 Genyornis newtoni 275 kg (606 lb)[13] Dromornithidae (mihirungs) Anseriformes (waterfowl) Late Pleistocene
8 Struthio anderssoni 270 kg (600 lb)[14] Struthionidae Struthioniformes (ostriches) Late Pleistocene
9 Dromornis murrayi 250 kg (550 lb)[15] Dromornithidae (mihirungs) Anseriformes (waterfowl) Late Oligocene-Early Miocene
10 Paraphysornis brasilienis 240 kg (530 lb)[16] Phorusrhacidae (terror birds) Cariamiformes Early Miocene
11 Aepyornis hildebrandti 235 kg (518 lb)[5][Note 5] Aepyornithidae (greater elephant birds) Aepyornithiformes (elephant birds) Late Pleistocene-Holocene
12 Dromornis australis 220 kg (490 lb)[13] Dromornithidae (mihirungs) Anseriformes (waterfowl) Pliocene
13 Gastornis gigantea 200 kg (440 lb)[13] Gastornithidae Anseriformes (waterfowl) Eocene
14 Dinornis sp. 192 kg (423 lb)[17][Note 6] Dinornithidae (giant moa) Dinornithiformes (moa) Late Pleistocene-Holocene
15 Ilbandornis woodburnei 175 kg (386 lb)[13] Dromornithidae (mihirungs) Anseriformes (waterfowl) Late Miocene
16 Devincenzia pozzi 161 kg (355 lb)[19] Phorusrhacidae (terror birds) Cariamiformes Neogene
17 Ilbandornis lawsoni 150 kg (330 lb)[13] Dromornithidae (mihirungs) Anseriformes (waterfowl) Late Miocene
18 Titanis walleri 150 kg (330 lb)[20] Phorusrhacidae (terror birds) Cariamiformes Pliocene-Early Pleistocene
19 Gastornis parisiensis 135 kg (298 lb)[21] Gastornithidae Anseriformes (waterfowl) Eocene
20 Gargantuavis philoinos 120 kg (260 lb)[22] Gargantuaviidae Avialae Late Cretaceous
21 Palaeeudyptes klekowskii 116 kg (256 lb)[23] Spheniscidae Sphenisciformes (penguin) Eocene
23 Kelenken guillermoi 100 kg (220 lb)[24] Phorusrhacidae (terror birds) Cariamiformes Middle Miocene
24 Phorusrhacos longissmus 94 kg (207 lb)[11] Phorusrhacidae (terror birds) Cariamiformes Miocene
25 Mullerornis modestus 80 kg (180 lb)[5] Mullerornithidae (lesser elephant birds) Aepyornithiformes (elephant birds) Late Pleistocene-Holocene
26 Pachyornis elephantopus 80 kg (180 lb)[25] Emeidae (lesser moa) Dinornithiformes (moa) Late Pleistocene-Holocene
27 Argentavis magnificens 71 kg (157 lb)[26][Note 7] Teratornithidae (teratorn) Cathartiformes (New World vultures) Miocene
28 Barawertornis tedfordi 70 kg (150 lb)[13] Dromornithidae (mihirungs) Anseriformes (waterfowl) Late Oligocene-Early Miocene
29 Mesembriornis incertus 70 kg (150 lb)[27] Phorusrhacidae (terror birds) Cariamiformes Pliocene
30 Pachyornis australis 67 kg (148 lb)[25] Emeidae (lesser moa) Dinornithiformes (moa) Late Pleistocene-Holocene
31 Emeus crassus 58 kg (128 lb)[25] Emeidae (lesser moa) Dinornithiformes (moa) Late Pleistocene-Holocene
32 Euryapteryx curtus 47.5 kg (105 lb)[25] Emeidae (lesser moa) Dinornithiformes (moa) Late Pleistocene-Holocene
33 Megalapteryx didinus 40 kg (88 lb)[25] Megalapterygidae (upland moa) Dinornithiformes (moa) Late Pleistocene-Holocene
34 Anomalopteryx didiformis 40 kg (88 lb)[25] Emeidae (lesser moa) Dinornithiformes (moa) Late Pleistocene-Holocene
35 Sylviornis neocaledoniae 30.5 kg (67 lb)[27] Sylviornithidae (giant megapodes) Galliformes (landfowl) Late Pleistocene-Holocene
36 Eremopezus eocaenus 30 kg (66 lb)[28] Eremopezidae Eocene
37 Pezophaps solitaria 28 kg (62 lb)[29][Note 8] Columbidae Columbiformes (doves and pigeons) Holocene
38 Pachyornis geranoides 27 kg (60 lb)[25] Emeidae (lesser moa) Dinornithiformes (moa) Late Pleistocene-Holocene
39 Patagornis marshi 23 kg (51 lb)[27] Phorusrhacidae (terror birds) Cariamiformes Miocene
40 Pelagornis sandersi 21.7 kg (48 lb)[30][Note 9] Pelagornithidae Odontopterygiformes (pseudotooth birds) Oligocene

Largest extant birds

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Table of heaviest extant bird species

[edit]

The following table is a list of the heaviest extant bird species based on maximum reported or reliable mass, with the average weight is also given for comparison. These species are almost all flightless, having denser bones and heavier bodies. Flightless birds comprise less than 1% of all extant bird species.

Rank Common name Binomial Name Average mass
[kg (lb)] 		Maximum mass
[kg (lb)] 		Average total length
[cm (ft)] 		Flighted
1 Common ostrich Struthio camelus 104 kg (229 lb)[1] 156.8 kg (346 lb)[1] 210 cm (6.9 ft)[31] No
2 Somali ostrich Struthio molybdophanes 90 kg (200 lb)[1] 130 kg (290 lb)[citation needed] 200 cm (6.6 ft)[1] No
3 Southern cassowary Casuarius casuarius 45 kg (99 lb)[1] 85 kg (187 lb)[32] 155 cm (5.09 ft)[1] No
4 Northern cassowary Casuarius unappendiculatus 44 kg (97 lb)[1] 75 kg (165 lb)[1] 149 cm (4.89 ft)[31] No
5 Emu Dromaius novaehollandiae 33 kg (73 lb)[1][33] 70 kg (150 lb)[citation needed] 153 cm (5.02 ft)[1] No
6 Emperor penguin Aptenodytes forsteri 31.5 kg (69 lb)[31][34] 46 kg (101 lb)[31] 114 cm (3.74 ft)[31] No
7 Greater rhea Rhea americana 23 kg (51 lb)[33] 40 kg (88 lb)[31] 134 cm (4.40 ft)[1] No
8 Dwarf cassowary Casuarius bennetti 19.7 kg (43 lb)[1] 34 kg (75 lb)[1] 105 cm (3.44 ft)[citation needed] No
9 Lesser rhea Rhea pennata 19.6 kg (43 lb)[1] 28.6 kg (63 lb)[1] 96 cm (3.15 ft)[31] No
10 King penguin Aptenodytes patagonicus 13.6 kg (30 lb)[31][34] 20 kg (44 lb)[35] 92 cm (3.02 ft)[citation needed] No
11 Turkey Meleagris gallopavo 13.5 kg (30 lb)[36] 39 kg (86 lb)[37] 100–124.9 cm (3.28–4.10 ft)[citation needed] Yes
12 Trumpeter swan Cygnus buccinator 12.7 kg (28 lb) 17.2 kg (38 lb) 138–165 cm (4.53–5.41 ft) Yes
13 Wandering albatross Diomedea exulans 11.9 kg (26 lb) 16.1 kg (35 lb)[38] 107–135 cm (3.51–4.43 ft) Yes
14 Mute swan Cygnus olor 11.87 kg (26.2 lb) 23 kg (51 lb) 100–130 cm (3.3–4.3 ft)[39] Yes
15 Dalmatian pelican Pelecanus crispus 11.5 kg (25 lb) 15 kg (33 lb)[citation needed] 183 cm (6.00 ft)[citation needed] Yes
16 Kori bustard Ardeotis kori 11.4 kg (25 lb)[31] 20 kg (44 lb)[citation needed] 150 cm (4.9 ft)[31] Yes
17 Whooper swan Cygnus cygnus 11.4 kg (25 lb) 15.5 kg (34 lb) 140–165 cm (4.59–5.41 ft) Yes
18 Andean condor Vultur gryphus 11.3 kg (25 lb)[39] 14.9 kg (33 lb)[39] 100–130 cm (3.3–4.3 ft)[39] Yes
19 California condor Gymnogyps californianus 10.8 kg (24 lb)[citation needed] 14.1 kg (31 lb)[40][41] 109–140 cm (3.58–4.59 ft)[40] Yes
20 Great bustard Otis tarda 10.6 kg (23 lb)[citation needed] 21 kg (46 lb)[42] 115 cm (3.77 ft)[citation needed] Yes

By order

[edit]
The Andean condor is the largest living bird of prey.[43][44]
The Eurasian black vulture is the largest member of Accipitridae.
The secretarybird is the largest bird of prey in terms of height and length.

Birds of prey (Accipitriformes)

[edit]
  • New World vultures are generally considered belonging to this order,[45] although their inclusion is not accepted by all.[46] If included, the largest species of this order, based on body weight and wingspan, is the Andean condor (Vultur gryphus) of western South America. The Andean condor can reach a wingspan of 3.2 m (10 ft)[47] and a weight of 15 kg (33 lb).[48]
  • Excluding New World vultures, the largest extant species is the Eurasian black vulture (Aegypius monachus). The Eurasian black vulture can attain a maximum weight of 14 kg (31 lb), a height of up to 1.2 m (3.9 ft), and a wingspan of 3.1 m (10 ft).[49] Other vultures can be almost as large, with the Himalayan vulture (Gyps himalayensis) reaching lengths of up to 1.5 m (4.9 ft) due to its long neck.[50]
  • The largest living member of this order, in terms of length and height, is the secretarybird (Sagittarius serpentarius) of sub-Saharan Africa. It measures 0.9–1.3 m (3.0–4.3 ft) in height and 1.2–1.5 m (3.9–4.9 ft) in length. Its wingspan can reach 1.2–1.35 m (3.9–4.4 ft) and have a weight of 2.3–4.27 kg (5.1–9.4 lb).[51]
  • The largest predatory bird, specifically the largest eagle, is a source of contention.
    • The harpy eagle (Harpia harpyja) of neotropical forests is often cited as the most massive eagle, with wild females up to 10 kg (22 lb) in weight and captive females occasionally growing to weights of over 12 kg (26 lb).[42]
    • The Steller's sea eagle (Haliaeetus pelagicus) of Asia's North Pacific, with unconfirmed weights of up to 12.7 kg (28 lb), and an average weight of 6.7 kg (15 lb), is sometimes regarded as the heaviest eagle.
    • The up to 1.12 m (3.7 ft) Philippine eagle (Pithecophaga jefferyi) has the greatest length of any eagle. The harpy and Philippine eagles, due to having to navigate in deep forest, are relatively short-winged and do not exceed 2 or 2.2 m (6.6 or 7.2 ft), respectively, in wingspan.[42]
    • The golden eagle (Aquila chrysaetos) is of marginally larger wingspan, with the Himalayan subspecies recorded at 2.77 m (9.1 ft).[42]
    • The white-tailed eagle (Haliaeetus albicilla) measures 66–94 cm (26–37 in) in length with a 1.78–2.45 m (5.8–8.0 ft) wingspan. Its wingspan, with a midpoint of 2.18 m (7.2 ft), is on average the largest of any eagle.
    • The white-tailed eagle is sometimes considered the fourth-largest eagle in the world, and is on average the fourth-heaviest.
    • The martial eagle (Polemaetus bellicosus) is the largest eagle in Africa, and the fifth-heaviest (on average) eagle in the world, with a length of 78–96 cm (31–38 in), a weight of 3–6.2 kg (6.6–13.7 lb) and a wingspan of 188–260 cm (6 ft 2 in – 8 ft 6 in).
    • The longest wingspan of an eagle ever recorded was an Australian wedge-tailed eagle (Aquila audax) at 2.83 m (9.3 ft). However, less substantiated records indicate that the Steller's sea eagle may reach at least 2.74 m (9.0 ft).[52]
    • The now extinct Haast's eagle (Hieraaetus moorei), which existed alongside early aboriginal people in New Zealand, was by far the largest eagle known and perhaps the largest raptor ever. Adult female Haast's are estimated to have averaged up to 1.4 m (4.6 ft) in length, weighing up to 15 kg (33 lb), with a relatively short 3 m (9.8 ft) wingspan.[53]
Migrating trumpeter swans are the largest waterfowl.
  • The largest of the accipitrine hawks is the Eurasian goshawk (Accipiter gentilis) of temperate Eurasia. They range in size variably, but on average measure 53–64 cm (21–25 in) in length, have a wingspan of 103–117 cm (3.38–3.84 ft) and weigh 0.63–1.4 kg (1.4–3.1 lb).[54] The Henst's goshawk (Accipiter henstii) and Meyer's goshawk (Accipiter meyerianus) do rival it in terms of wing size and body mass.
  • Among the buteonine hawks, the largest species are the ferruginous hawk (Buteo regalis) and the upland buzzard (Buteo hemilasius) of North America and Asia respectively. The former can have a wingspan of 133–142 cm (4.36–4.66 ft), weigh 0.98–2.1 kg (2.2–4.6 lb) and measure 56–69 cm (22–27 in) in length.[55] The weight of the upland buzzard, which can be in the range of 0.95–2.05 kg (2.1–4.5 lb), broadly overlaps that of the ferruginous hawk, even though it is slightly larger at 57–72 cm (22–28 in) long and with a wingspan of 143–161 cm (56–63 in).
  • The swamp harrier (Circus approximans) of Australasia is believed to be the largest species of harrier, measuring 50–60 cm (20–24 in) long, having a wingspan of 120–145 cm (47–57 in) and weighing 0.58–1.1 kg (1.3–2.4 lb).[56]
  • The largest species of kite is the red kite (Milvus milvus). With a wingspan of 174–179 cm (69–70 in), it measures 60–70 cm (24–28 in) in length and weighs 0.8–1.3 kg (1.8–2.9 lb).

Waterfowl (Anseriformes)

[edit]
  • The largest waterfowl species by average size is the trumpeter swan (Cygnus buccinator) of Northern North America, which can reach a length of 1.82 m (6.0 ft), a wingspan of 3.1 m (10 ft) and a weight of 17.3 kg (38 lb).[57] The heaviest single waterfowl ever recorded was a cob (Cygnus olor) from Poland which weighed 23 kg (51 lb), and was allegedly too heavy to take flight.[42]
  • The largest species of goose is the Canada goose (Branta canadensis), more specifically the subspecies known as the giant Canada goose (Branta canadensis maxima). Individuals can reach more than 9.1 kg (20 lb) in weight.[58]
  • The largest 'duck' species is the Muscovy duck (Cairina moschata) of the Americas. Males can weigh from 4.5–6.3 kg (9.9–13.9 lb) and can measure up to 86 cm (34 in).[59] However, its genus is now considered to be paraphyletic with the species currently being placed in the subfamily Tadorninae (shelducks and shelgeese). If so, the largest species of the true ducks or dabbling ducks (Anatinae) is the mallard (Anas platyrhynchos).[60] They can measure 50–65 cm (20–26 in) in length, have a wingspan of 82–95 cm (32–37 in) and a weight of 1–1.3 kg (2.2–2.9 lb).[61]

Swifts and allies (Apodiformes)

[edit]
  • The largest species of Apodiformes is the white-naped swift (Streptoprocne semicollaris), endemic to southern Mexico, and the purple needletail (Hirundapus celebensis), of the Philippine islands. Both reach weights of up to 225 g (7.9 oz), lengths of up to 25 cm (9.8 in) and wingspans as long as 0.6 m (2.0 ft).[62]
  • Traditionally included in this order, by far the largest hummingbird species is the giant hummingbird (Patagona gigas) of the Andes Mountains. "Giant" is a relative term among the hummingbirds, the smallest-bodied variety of birds, with the giant hummingbird species weighing up to 24 g (0.85 oz) with a length of 23 cm (9.1 in).[63]
  • The longest hummingbird species, indeed the longest in the order, is the adult male black-tailed trainbearer (Lesbia victoriae), which can measure up to 25.5 cm (10.0 in). The majority of this length is due to the hummingbird's extreme tail streamers. Another size champion among hummingbirds is the sword-billed hummingbird, a fairly large species of which approximately half of its 21 cm (8.3 in) length derives from its bill. This is by far the largest bill-to-body-size ratio of any bird.[64]
The great potoo is, overall, the largest member of the order Caprimulgiformes.

Hornbills, hoopoe, and wood-hoopoes (Bucerotiformes)

[edit]
  • The largest species of Coraciiformes is the southern ground hornbill (Bucorvus leadbeateri), which can reach weights of up to 6.2 kg (14 lb) and grow as long as 1.3 m (4.3 ft).[65] Several arboreal, Asian hornbills can also grow very large, with the great hornbill (Buceros bicornis) weighing up to 4 kg (8.8 lb), and the helmeted hornbill (Rhinoplax vigil) measuring as much as 1.7 m (5.6 ft) in total length.[66][67] The larger hornbills have a wingspan of up to 1.83 m (6.0 ft).[68]

Nightjars and allies (Caprimulgiformes)

[edit]
  • The largest species of this order of nocturnal birds is the neotropical great potoo (Nycitbius grandis), which can grow to a weight of 680 g (1.50 lb) and a height of 60 cm (2.0 ft). Heavier Caprimulgiformes have been recorded in juvenile specimens of the Australian tawny frogmouth (Podargus strigoides), which can weigh up to 1.4 kg (3.1 lb). Other species nearly as large as the potoo are the Papuan frogmouth (Podargus papuensis) of New Guinea and the neotropic, cave-dwelling oilbird (Steatornis caripensis), both growing as large as 48 cm (19 in). The wingspan of the great potoo and the oilbird can be more than 1 m (3.3 ft), the largest of the order.[69][70]
  • The largest species of the nightjar family, the great eared nightjar (Eurostopodus macrotis) of East Asia, is of smaller proportions. Great eared nightjars can reach 150 g (0.33 lb) in weight and 41 cm (16 in) in height.[71]

Shorebirds (Charadriiformes)

[edit]
  • The largest species in this diverse order is the great black-backed gull (Larus marinus) of the North Atlantic, attaining a height as large as 0.79 m (2.6 ft), a wingspan of 1.7 m (5.6 ft) and a weight of up to 2.3 kg (5.1 lb). The glaucous gull (L. hyperboreus) is smaller on average than the black-back but has been weighed as heavy as 2.7 kg (6.0 lb).[33][72]
  • Among the most prominent family of "small waders", the sandpipers reach their maximum size in the Far Eastern curlew (Numenius madagascariensis) at up to 0.60 m (2.0 ft) in length and 1.1 m (3.6 ft) across the wings. The more widespread Eurasian curlew (N. arquata) can weigh up to 1.36 kg (3.0 lb).[73][74]
  • Less variable in size, the largest species of plovers is the Australasian masked lapwing (Vanellus miles) which grows up to 0.4 m (1.3 ft) long with a 0.85 m (2.8 ft) wingspan and a weight of 400 g (14 oz).[75] The widely distributed Caspian tern (Hydroprogne caspia), is relatively large and heavily built. Caspians can range up to 782 g (1.724 lb) in weight, with a 1.4 m (4.6 ft) wingspan and a length of 0.6 m (2.0 ft).[76][77]
  • The largest extant alcid is the sub-Arctic thick-billed murre (Uria lomvia), which can weigh up to 1.48 kg (3.3 lb), with a length of 0.48 m (1.6 ft) and a wingspan of 0.76 m (2.5 ft).[78] However, until its extinction, the flightless great auk (Pinguinus impennis) of the North Atlantic was both the largest alcid and the second-largest member of the order. Great auks could range up to 6.8 kg (15 lb) and 0.9 m (3.0 ft) tall.[79]
  • Miomancalla howardi was the largest charadriiform of all time, weighing approximately 1.5 ft (0.46 m)(?) more than the Great Auk with a height of approximately 1 m (3.3 ft).[80]
The Saddle-billed stork is perhaps the tallest of the storks.

Herons and allies (Ciconiiformes)

[edit]
  • The longest-bodied and tallest species in this order is the saddle-billed stork of Africa (Ephippiorhynchus senegalensis), which often exceeds 1.5 m (4.9 ft) tall and has a wingspan of up to 2.7 m (8.9 ft).[81] Reaching a similar height but more heavily built among the storks are the neotropical jabiru (Jabiru mycteria), the Asian greater adjutant (Leptoptilos dubius) and the African marabou stork (L. crumeniferus), all of which weigh up to 8 to 9 kg (18 to 20 lb).[33][82] The greater adjutant and marabou nearly equal the Andean condor in maximum wingspan, with all three birds believed to reach or exceed a wingspan of 3.16 m (10.4 ft).[42] Standing up to 1.53 m (5.0 ft), with a wingspan of up to 2.3 m (7.5 ft) and a weight up to 5 kg (11 lb), the African goliath heron (Ardea goliath) is the largest of the herons and egrets. Juvenile white-bellied heron (A. insignis) have been reported to weigh up to 8.5 kg (19 lb) with heights of 1.58 m (5.2 ft).[83]
  • Many of the largest flying birds in the fossil record may have been members of the Ciconiiformes. The heaviest flying bird ever, Argentavis magnificens, is part of a group, the teratorns, that is considered an ally of the New World vultures.[84]
  • The largest ibis is the giant ibis (Thaumatibis gigantea). Adults can grow to 102–106 cm (40–42 in) long, with a standing height of up to 100 cm (39 in) and are estimated to weigh approximately 4.2 kg (9.3 lb). Among standard measurements, the wing chord is 52.3–57 cm (20.6–22.4 in), the tail is 30 cm (12 in), the tarsus is 11 cm (4.3 in) and the culmen is 20.8–23.4 cm (8.2–9.2 in). The crested ibis (Nipponia nippon) of Japan is as large as 78.5 cm (30.9 in) in height and 30.9 cm (12.2 in) in length.

Mousebirds (Coliiformes)

[edit]
  • The largest mousebird species, the speckled mousebird (Colius striatus), weighs 2 ounces (57 g) with a height of over 14 inches (36 cm).[85]
The Victoria crowned pigeon is the largest living pigeon.

Pigeons (Columbiformes)

[edit]
  • The largest species of the pigeon/dove complex is the Victoria crowned pigeon (Goura victoria) of Northern New Guinea. Some exceptionally large Victoria crowned pigeons have reached 3.7 kg (8.2 lb) and 85 cm (33 in). The largest arboreal pigeon is the Marquesan imperial pigeon (Ducula galeata), which can grow approximately 0.8 m (2.6 ft) across the wings and can weigh 1 kg (2.2 lb).[86]
  • The largest pigeons and doves known to have existed were the dodo (Raphus cucullatus) and the Rodrigues solitaire (Pezophaps solitaria). Both flightless species may have exceeded 1 m (3.3 ft) in height. The dodo is frequently cited as the largest-ever pigeon, potentially weighing as much as 28 kg (62 lb), although recent estimates have indicated that an average wild dodo weighed much less at approximately 10.2 kg (22 lb).[87][88]

Rollers, Kingfishers, Bee-eaters, motmots, and todies (Coraciiformes)

[edit]
  • The largest kingfisher is the giant kingfisher (Megaceryle maxima), at up to 48 cm (19 in) long and 425 g (15.0 oz) in weight.[89] The common Australian species, the laughing kookaburra (Dacelo novaeguineae), may be heavier still, as individuals exceeding 450 g (0.99 lb) are not uncommon. A kookaburra's wingspan can range up to 0.9 m (3.0 ft).[90]

Cuckoos, coucals and roadrunners (Cuculiformes)

[edit]
  • The largest of the cuckoos is the Australasian channel-billed cuckoo (Scythrops novaehollandiae), which can range up to a weight of 0.93 kg (2.1 lb), a 1 m (3.3 ft) wingspan and a length of 0.66 m (2.2 ft).[91][92]

Falcons (Falconiformes)

[edit]
  • Many authorities now support the split of falcons from the Accipitriformes, despite similar adaptations, due to the genetic evidence showing they are not closely related.[93] The largest species of falcon is the gyrfalcon (Falco rusticolus). Large females of this species can range up to 2.1 kg (4.6 lb), span 1.6 m (5.2 ft) across the wings and measure 0.66 m (2.2 ft) long.[49]
The Indian peafowl is one of the largest living gamebirds.

Gamebirds (Galliformes)

[edit]
  • The heaviest member of this order is the North American wild turkey (Meleagris gallopavo). The largest specimen ever recorded was shot in 2015, and weighed 17.05 kg (37.6 lb).[94] The heaviest domesticated turkey on record weighed 37 kg (82 lb).[47]
  • The longest gamebirds species, if measured from the tip of the bill to the end of the tail coverts, is the male green peafowl (Pavo muticus) of Southeast Asia at a length of up to 3 m (9.8 ft), with two-thirds of the length being made up by the tail coverts. It has a relatively large wingspan for a gamebird, spanning as much as 1.6 m (5.2 ft) across the wings.[95]
  • The largest member of the grouse family is the Eurasian western capercaillie (Tetrao urogallus), weighing up to 6.7 kg (15 lb) with a length of 1 m (3.3 ft).
  • A prehistoric, flightless family, sometimes called (incorrectly) "giant megapodes" (Sylviornis) of New Caledonia were the most massive galliformes ever, with lengths of up to 1.7 m (5.6 ft) weights up to approximately 40 kg (88 lb).[96]

Loons (Gaviiformes)

[edit]
  • The largest species on average is the yellow-billed loon (Gavia adamsii) of the Arctic, at up to 1 m (3.3 ft) and 7 kg (15 lb). One exceptionally large North American Great northern diver (Gavia immer) was weighed at 8 kg (18 lb), heavier than any recorded yellow-billed loon. Wingspans of the largest loons can reach 1.52 m (5.0 ft).[97]
Alongside the great bustard, the kori bustard is the heaviest extant flying bird.

Cranes and allies (Gruiformes)

[edit]
  • The males of the Eurasian great bustard (Otis tarda) and the African kori bustard (Ardeotis kori) are the heaviest birds capable of flight, averaging up to 16 kg (35 lb) and weighing 2 to 3 times as much as their female counterparts. It is not resolved if one of these species is larger than the other, but both can reach a weight of at least 21 kg (46 lb)[47] and measure up to 1.53 m (5.0 ft) long.[42][98] Some kori bustards have been reported from 23 kg (51 lb) to even 40 kg (88 lb), but all such reports are unverified or dubious.[99]
  • The tallest flying bird on earth, also represented in the Gruiformes, is the sarus crane (Grus antigone) of Southern Asia and Australia, which can reach a height of 2 m (6.6 ft).[100] Heavier cranes are reported in other species, the red-crowned crane (Grus japonensis) and the Siberian crane (G. leucogeranus), both from Northeast Asia and both at up to 15 kg (33 lb), as opposed to a top weight of 12.8 kg (28 lb) in the sarus.[42][33][101] Wingspan in both the largest cranes and the largest bustards can range up to 2.5–3 m (8.2–9.8 ft).[102][103]
  • The most species-rich family in this order, the rails, reaches their largest size in the bulky takahē (Porphyrio hochstetteri) of New Zealand, an endangered species that can weigh up to 4.2 kg (9.3 lb) and measure 0.65 m (2.1 ft) long.[104] The aforementioned "terror bird", Brontornis burmeisteri, has traditionally been classified as a member of this order, although this may not be an accurate classification.
The thick-billed raven shares the title of the largest songbird with its common cousin.

Hoatzin (Opisthocomiformes)

[edit]
  • Hoatzin (Opisthocomus hoazin), the only member of its order, is a pheasant-sized South American bird, with a total length of 65 cm (26 in) and a maximum weight of 1 kg (2.2 lb).
The hyacinth macaw is the largest parrot.

Songbirds (Passeriformes)

[edit]
  • The passerine or songbird order comprises more than half of all bird species, and are known for their generally small size, their strong voices and their frequent perching. Corvids are the largest of passerines, particularly the large races of the common raven (Corvus corax) and the Northeast African thick-billed raven (C. crassirostris). Large ravens can weigh 2 kg (4.4 lb), attain a 1.5 m (4.9 ft) wingspan and measure 0.8 m (2.6 ft) long.[105]
  • The closest non-corvid contender to largest size is the Australian superb lyrebird (Menura novaehollandiae), which can reach a length of 1 m (3.3 ft), much of it comprised by their spectacular tail, and a weight of 1 kg (2.2 lb).[106]
  • The largest species in the most species-rich passerine family, Tyrannidae or tyrant-flycatchers, is the great shrike-tyrant of the South Andes (Agriornis lividus), at 99.2 g (3.50 oz) and 31 cm (12 in), although the fork-tailed flycatcher (Tyrannus savana), to 41 cm (16 in), is longer thanks to its extreme tail.[33][107]
  • The namesake of the previous family, the Old World flycatchers, reaches its maximum size in the blue whistling thrush of India Southeast Asia (Myophonus caeruleus), if it is indeed a proper member of the family, at up to 122 g (4.3 oz) and a length of 29 cm (11 in).[108]
  • Closely related to the Old World flycatchers, the thrush family's largest representative is the Great thrush of South America (Turdus fuscater), at up to 175 g (6.2 oz) and 28 to 33 cm (11 to 13 in).[109]
  • The largest bird family in Eurasia is the Old World warblers. As previously classified these warblers could get fairly large, up to 57 g (2.0 oz) and 28 cm (11 in) in the striated grassbird of Southeast Asia (Megalurus palustris). The Old World warblers have been split into several families, however, which leaves the barred warbler of central Eurasia (Sylvia nisoria), up to 36 g (1.3 oz) and 17 cm (6.7 in), as the largest "true warbler".[110]
  • Not to be confused with the previous family, the largest of the well-known New World warblers is the aberrant yellow-breasted chat (Icteria virens), which can exceptionally measure up to 22 cm (8.7 in) and weigh 53 g (1.9 oz).[111][112]
  • Another large family is the bulbuls, the largest of which is the south Asian straw-headed bulbul (Pycnonotus zeylanicus), to 94 g (3.3 oz) and 29 cm (11 in). The diverse, large family of babblers can reach 35 cm (14 in) and 170 g (6.0 oz) in the south Asian greater necklaced laughingthrush (Garrulax pectoralis).[113]
  • The familiar domesticated species, the Java sparrow (Padda oryzivora), is (in the wild) the largest estrildid, at up to 28.3 g (1.00 oz) and 17 cm (6.7 in). The largest honeyeater, perhaps the most diverse Australasian bird family, is the crow honeyeater (Gymnomyza aubryana), at up to 290 g (10 oz)[33] and 30 cm (12 in). The largest of the "true finches" is the collared grosbeak (Mycerobas affinis) of central and south Asia at up to 23 cm (9.1 in) and 80 g (2.8 oz).[114]
  • Among the largest bird families, the emberizids, reaches its largest size in the Abert's towhee (Pipilo aberti) of Southwest United States and north Mexico at up to 23 cm (9.1 in) and 80 g (2.8 oz).[115]
  • Closely related to the previous family is the tanagers, which can range up to 140 g (4.9 oz) in the Andean-forest-dwelling white-capped tanager (Sericossypha albocristata).[33] Another species-rich neotropical family is the ovenbirds, the largest of which, the great rufous woodcreeper (Xiphocolaptes major) of the Amazonian rainforest, can weigh up to 162 g (5.7 oz) and 35 cm (14 in). The specialized antbird family can range up to 156 g (5.5 oz) and 35.5 cm (14.0 in) in the giant antshrike (Batara cinerea).[33] Among the most variably sized passerine families is the icterids.
  • The largest icterid is the olive oropendola (Psarocolius bifasciatus), in which males can range up to 52 cm (1.71 ft) and 550 g (1.21 lb).[116] The latter species competes with the similarly sized Amazonian umbrellabird (Cephalopterus ornatus) as the largest passerine in South America.
The Dalmatian pelican is one of the largest flying birds.

Cormorants and allies (Pelecaniformes)

[edit]
  • Pelicans rank amongst the largest flying birds. The largest species of pelican is the Eurasian Dalmatian pelican (Pelecanus crispus), which can attain a length of 1.83 m (6.0 ft) and a body weight of 15 kg (33 lb). The great white pelican (P. onocrotalus) of Europe and Africa is almost as large. The Australian pelican (P. conspicillatus) is slightly smaller but has the largest bill of any bird, at as much as 49 cm (19 in) long.[47] A large pelican can attain a wingspan of 3.6 m (12 ft), second only to the great albatrosses among all living birds.[117]
  • The largest of the cormorants is the flightless cormorant of the Galapagos Islands (Nannopterum harrisi), at up to 5 kg (11 lb) and 1 m (3.3 ft), although large races in the great cormorant (Phalacrocorax carbo) can weigh up to 5.3 kg (12 lb).[118][119] The spectacled cormorant of the North Pacific (Urile perspicillatus), which became extinct around 1850, was larger still, averaging around 6.4 kg (14 lb) and 1.15 m (3.8 ft).
  • The widely distributed magnificent frigatebird is of note for having an extremely large wingspan, up to 2.5 m (8.2 ft), for its relatively light body, at up to only 1.9 kg (4.2 lb).[120]
  • Pelagornithidae or pseudotooth birds included several species that were behind only Argentavis magnificens in size among all flying birds. Characterized by the tooth-like protrusions along their bills, this unique family has been variously allied with the Pelecaniformes, tubenoses, large waders and even waterfowl. Their true linkage to extant birds remains in question, though pelecaniformes are the group most regularly considered related. Some of the largest pseudotooth birds have included, Osteodontornis of the late Miocene from the North Pacific, Gigantornis eaglesomei, from the Eocene era in what is now Nigeria and Dasornis, from Eocene era Europe. A new, unnamed species has been discovered which may outsize even these giants. Superficially albatross-like, each of these pseudotooth species may have attained lengths of 2.1-metre-long (6.9 ft) and wingspans of at least 6 m (20 ft). Body mass in these slender birds was probably only up to around 29 kg (64 lb).[121][122][123]

Tropicbirds (Phaethontiformes)

[edit]
  • The largest tropicbirds is the red-billed tropicbird (Phaethon aethereus). The adult is a slender, mainly white bird, 48 cm long, excluding the central tail feathers which double the total length, and a one-meter wingspan.

Flamingos (Phoenicopteriformes)

[edit]
  • The largest flamingo is the greater flamingo (Phoenicopterus roseus) of Eurasia and Africa. One of the tallest flying birds in existence when standing upright (exceeded only by the tallest cranes), this species typically weighs 3.5 kg (7.7 lb) and stands up to 1.53 m (5.0 ft) tall. At maximum, a male can weigh up to 4.55 kg (10.0 lb) and stand as high as 1.87 m (6.1 ft).[124] Wingspan is relatively small in flamingos, ranging up to 1.65 m (5.4 ft).[125]
The toco toucan is the largest species in the order Piciformes as well as one of the most colorful.

Woodpeckers and allies (Piciformes)

[edit]
  • The largest species of this order is the toco toucan (Ramphastos toco) of the neotropic forest. Large specimens of this toucan can weigh to 870 g (1.92 lb) and 0.65 m (2.1 ft), at which size the beak alone can measure approximately 20 cm (7.9 in).[126]
  • Until the 20th century, the largest woodpecker was the imperial woodpecker (Campephilus imperialis) of Mexico, with a length of up to 0.6 m (2.0 ft). This species is generally believed to have gone extinct following habitat destruction and hunting.[127] The closely related ivory-billed woodpecker (Campephilus principalis) of the Southeast United States and Cuba approached similar sizes at up to 0.5 m (1.6 ft) in length, with a wingspan of 0.78 m (2.6 ft) and a mass of at least 530 g (1.17 lb). Despite possibilities that it has survived in some deep swamp forests in Arkansas or Florida, the ivory-billed is also generally considered to have gone extinct.[128] The great slaty woodpecker (Mulleripicus pulverulentus) of southeast Asia is the largest woodpecker certain to exist, with a weight of up to 500 g (1.1 lb) and a length of up to 0.58 m (1.9 ft).[129]
  • Less well-known than the woodpeckers and toucans, barbets can range up to 273 g (9.6 oz) and 33 cm (13 in) in the great barbet (Megalaima virens).[126]
  • The largest jacamar is the great jacamar (Jacamerops aureus). It measures 29.5 to 30 cm (11.61 to 11.81 in) in length and weighs between 63 and 70 g (2.22 and 2.47 oz).[130]

Grebes (Podicipediformes)

[edit]
  • The largest species of grebe is the South American great grebe (Podiceps major). It can reach a length of 0.8 m (2.6 ft), with a wingspan of 1 m (3.3 ft) and a weight of over 2 kg (4.4 lb).[131]
The wandering albatross is the largest seabird.

Tubenoses (Procellariiformes)

[edit]
  • The largest species of Procellariiformes is the wandering albatross (Diomedea exulans) of the sub-Antarctic oceans, which has the largest wingspan of any living bird. The maximum dimensions of this species are a length of 1.44 m (4.7 ft) and a wingspan of 3.65 m (12.0 ft).[47] Unverified specimens have been reported to measure 5.3 m (17 ft).[42] Immature wandering albatrosses have weighed as much as 15.9 kg (35 lb) at the time of their first flights, with the maximum reported weight of adults being 12.7 kg (28 lb).[42]
  • The Southern royal albatross (Diomedea epomophora) is slightly lesser in length, wingspan and weight.[117]
  • The largest procellarids is the southern giant petrel (Macronectes giganteus). It can reach a body length of 1 m (39 in), with a wingspan of 2.1 m (83 in) and a weight of 8 kg (17.6 lb).

Parrots (Psittaciformes)

[edit]
  • The largest parrot by length and wingspan is the endangered hyacinth macaw (Anodorhynchus hyacinthinus) of the neotropic lowlands, reaching a length of nearly 1.2 m (3.9 ft) with a wingspan of 1.4 m (4.6 ft) and weighing as little as 2 kg (4.4 lb).[132] The heaviest parrot is the nearly extinct kākāpō (Strigops habroptilus),[133] which is part of the New Zealand parrot family. The flightless kākāpō does not exceed 0.68 m (2.2 ft) in length, but weighs up to 4.1 kg (9.0 lb).[134]
  • The largest parakeet is the Alexandrine parakeet (Palaeornis eupatoria), reaching lengths of up to 60 centimetres and a mass of 250 grams.
  • The largest species in the cockatoo family is the Australasian palm cockatoo (Probosciger aterrimus), at up to 0.6 m (2.0 ft) long with a weight of 1.2 kg (2.6 lb).[135]

Sandgrouse (Pterocliformes)

[edit]
The emperor penguin is one of the heaviest living birds as well as the largest penguin.

Penguins (Sphenisciformes)

[edit]
  • The largest species of Sphenisciformes is the emperor penguin (Aptenodytes forsteri) of the Antarctic, with a maximum height of 1.35 m (4.4 ft) and a weight of 46 kg (101 lb).[42] The next largest living species is the king penguin, which grows to a maximum of 1 m (3.3 ft) in height and 18 kg (40 lb) in weight.[137] Now extinct, Anthropornis nordenskjoeldi, is believed to have reached a height of 1.8 m (5.9 ft) and a weight of up to 108 kg (238 lb).[138]
The Eurasian eagle-owl is one of the biggest owls.

Owls (Strigiformes)

[edit]
  • The most massive owl is certainly either the Eurasian eagle-owl (Bubo bubo) or the endangered and similarly sized Blakiston's fish owl (Bubo blakistoni) of coastal Russia and Japan. Record-sized specimens of both species have weighed approximately 4.5 kg (9.9 lb) and measured over 0.75 m (2.5 ft) long.[139] In either species, the wingspan can range up to 2 m (6.6 ft).[140][141] Longer still, but not as massive as the previous species (never more than 1.8 kg (4.0 lb) in weight), a large female great gray owl (Strix nebulosa) from the northern boreal forest can range up to 0.83 m (2.7 ft).[142]
  • The largest of the barn or masked owl family is the Tasmanian masked owl (Tyto novaehollandiae castanops), which weighs up to 1.4 kg (3.1 lb) and measures up to 0.6 m (2.0 ft).[141] The largest owl known to have existed was Ornimegalonyx oteroi of Cuba, a uniquely cursorial owl. The giant bird was estimated to stand over 1.1 m (3.6 ft) on the ground and to weigh at least 9.05 kg (20.0 lb).[143]
The grey tinamou ranks as the largest species of tinamou.

Ratites (Struthioniformes)

[edit]
  • The largest ratite is the ostrich (Struthio camelus), from the plains of Africa and Arabia. A large male ostrich can reach a height of 2.8 m (9.2 ft) and weigh over 156 kg (344 lb).[47] A mass of 200 kg (440 lb) has been cited for the ostrich but no wild ostriches of this weight have been verified.[2] Eggs laid by the ostrich are the largest in the world, weighing 1.4 kg (3.1 lb). The emu (Dromaius novaehollandiae) of Australia reaches 1 to 1.3 m (3.3–4.3 ft) at the shoulder with a full height of 150 to 190 cm (59–75 in). In length measured from the bill to the tail, emus range from 139 to 164 cm (55 to 65 in). The southern cassowary (Casuarius casuarius) from Australia and Papua New Guinea has a height of 127 to 190 cm (50 to 75 in).[1] The greater rhea (Rhea americana) from South America weighs up to 20–27 kg (44–60 lb) and often measures 127 to 140 cm (50 to 55 in) long from beak to tail with a height of approximately 1.5 m (4.9 ft).
  • The largest bird in the fossil record may be the extinct elephant birds (Vorombe/Aepyornis) of Madagascar, which were related to the ostrich. They exceeded 3 m (9.8 ft) in height and 500 kg (1,100 lb) in weight.[42] The last of the elephant birds became extinct approximately 1000 years ago.
  • The tallest bird ever was the South Island giant moa (Dinornis robustus), part of the moa family of New Zealand that went extinct about 500 years ago. The moa stood up to 3.7 m (12 ft) tall,[47] and weighed approximately half as much as a large elephant bird or mihirung due to its comparatively slender frame.[42]

Tinamous (Tinamiformes)

[edit]
  • The largest species of tinamou, a group of chunky, elusive ground-birds from neotropical forests, is the grey tinamou (Tinamus tao) of western South America. It can reach a weight over 2 kg (4.4 lb) and length of over 55 cm (22 in).[1]

Trogons (Trogoniformes)

[edit]
  • The resplendent quetzal (Pharomachrus mocinno) of the montane forest of Central America is the largest trogon, though a few other quetzals approach similar sizes. It can weigh more than 226 g (8.0 oz) and, in females and non-breeding or immature males, they can measure up to 0.4 m (1.3 ft) from the head to the tail. Upon developing tail streamers, adult males can reach lengths of up to 0.6 m (2.0 ft).[144]

See also

[edit]

Notes

[edit]
  1. ^ however this may be a vast overestimation based on robustness of the femur, A. maximus was more likely to have weighted around 275 kg (606 lb).[4]
  2. ^ Sexually dimorphic with larger males.[8]
  3. ^ Estimated to be the tallest bird ever, though this is based on fragmentary remains [10]
  4. ^ Disputed, was probably an Anseriformes but could still be an Cariamiformes
  5. ^ however this may be a vast overestimation based on robustness of the femur, A. hildebrandti was more likely to have weighted around 65 kg (143 lb).[4]
  6. ^ Sexually dimorphic with larger females.[17] D. novozealandae noted to be somewhat smaller than D. robustus.[18]
  7. ^ Heaviest known flying bird
  8. ^ Sexually dimorphic with larger males.[29]
  9. ^ Largest wingspan of any bird

References

[edit]
  1. ^ a b c d e f g h i j k l m n o p q r Davies, Stephen, Ratites and Tinamous. Oxford University Press (2002), ISBN 978-0-19-854996-3
  2. ^ a b Avian Medicine: Principles and Application Archived 2009-04-19 at the Wayback Machine. avianmedicine.net
  3. ^ "Largest egg from a bird (living, specimen)". Guinness World Records. Retrieved 2020-11-05.
  4. ^ a b c Hume, J. P.; Walters, M. (2012). Extinct Birds. London: A & C Black. pp. 19–21. ISBN 978-1-4081-5725-1.
  5. ^ a b c d Grealy, Alicia; Miller, Gifford H.; Phillips, Matthew J.; Clarke, Simon J.; Fogel, Marilyn; Patalwala, Diana; Rigby, Paul; Hubbard, Alysia; Demarchi, Beatrice; Collins, Matthew; Mackie, Meaghan; Sakalauskaite, Jorune; Stiller, Josefin; Clarke, Julia A.; Legendre, Lucas J. (2023-02-28). "Molecular exploration of fossil eggshell uncovers hidden lineage of giant extinct bird". Nature Communications. 14 (1): 914. Bibcode:2023NatCo..14..914G. doi:10.1038/s41467-023-36405-3. ISSN 2041-1723. PMC 9974994. PMID 36854679.
  6. ^ "Fossil of 'largest flying bird' identified". BBC News. 2014-07-07. Retrieved 2024-05-29.
  7. ^ ibid.
  8. ^ a b Handley, Warren D.; Chinsamy, Anusuya; Yates, Adam M.; Worthy, Trevor H. (2016-09-02). "Sexual dimorphism in the late Miocene mihirung Dromornis stirtoni (Aves: Dromornithidae) from the Alcoota Local Fauna of central Australia". Journal of Vertebrate Paleontology. 36 (5): e1180298. Bibcode:2016JVPal..36E0298H. doi:10.1080/02724634.2016.1180298. ISSN 0272-4634. S2CID 88784039.
  9. ^ Lazaro, Enrico de (28 June 2019). "Giant Birds Roamed Europe Two Million Years Ago | Paleontology | Sci-News.com". Sci.News: Breaking Science News. Retrieved 3 March 2023.
  10. ^ name="Giant Birds Roamed Europe Two Million Years Ago"
  11. ^ a b Degrange, F. J., Noriega, J. I., & Areta, J. I. (2012). Diversity and paleobiology of the Santacrucian birds. Early Miocene paleobiology in Patagonia: high-latitude paleocommunities of the Santa Cruz Formation, 138-155.
  12. ^ Buffetaut, Eric; Angst, Delphine (2021). "A Giant Ostrich from the Lower Pleistocene Nihewan Formation of North China, with a Review of the Fossil Ostriches of China". Diversity. 13 (2): 47. doi:10.3390/d13020047. hdl:1983/0d3c1bba-d496-47b6-a61c-e141a472612a. ISSN 1424-2818.
  13. ^ a b c d e f g Murray, P. F. & Vickers-Rich, P. (2004)
  14. ^ Buffetaut1, Eric; Angst, Delphine (2017). "How Large was the Giant Ostrich of China?". EVOLUÇÃO - Revista de Geistória e Pré-História. 2 (1): 6–8. Retrieved 9 January 2023.{{cite journal}}: CS1 maint: numeric names: authors list (link)
  15. ^ Brennan, Imogen (19 February 2016). "Bones from ancestor of 'giant duck' discovered in Australia". PM. www.abc.net.au.
  16. ^ Molina-Pérez, Rubén; Larramendi, Asier (2019). Dinosaurs Facts and Figures: The Theropods. Princeton University Press. p. 179. ISBN 978-0-691-18031-1.
  17. ^ a b Brassey, Charlotte A.; Holdaway, Richard N.; Packham, Abigail G.; Anné, Jennifer; Manning, Philip L.; Sellers, William I. (2013-12-18). Kupczik, Kornelius (ed.). "More than One Way of Being a Moa: Differences in Leg Bone Robustness Map Divergent Evolutionary Trajectories in Dinornithidae and Emeidae (Dinornithiformes)". PLOS ONE. 8 (12): e82668. Bibcode:2013PLoSO...882668B. doi:10.1371/journal.pone.0082668. ISSN 1932-6203. PMC 3867382. PMID 24367537.
  18. ^ Olson, Valérie A.; Turvey, Samuel T. (2013-06-07). "The evolution of sexual dimorphism in New Zealand giant moa ( Dinornis ) and other ratites". Proceedings of the Royal Society B: Biological Sciences. 280 (1760): 20130401. doi:10.1098/rspb.2013.0401. ISSN 0962-8452. PMC 3652460. PMID 23576789.
  19. ^ Tambussi, C. P., & Degrange, F. J. (2013). The dominance of zoophagous birds: just a cliché?. In South American and Antarctic Continental Cenozoic Birds (pp. 87-102). Springer, Dordrecht.
  20. ^ Gould, G. C.; Quitmyer, I. R. (2005). "Titanis walleri: Bones of Contention" (PDF). Bulletin of the Florida Museum of Natural History. 45 (4): 201–229. doi:10.58782/flmnh.xumx1681.
  21. ^ Eric Buffetaut und Delphine Angst: Stratigraphic distribution of large flightless birds in the Palaeogene of Europe and its palaeobiological and palaeogeographical implications. Earth-Science Reviews 32 (2), 2014, S. 394–408.
  22. ^ Rubén Molina-Pérez, Asier Larramendi, David Connolly, Gonzalo Ángel Ramírez Cruz, Andrey Atuchin (June 25, 2019). Dinosaur Facts and Figures: The Theropods and Other Dinosauriformes. Princeton University Press. p. 282. ISBN 9780691190594. Retrieved 30 August 2022.{{cite book}}: CS1 maint: multiple names: authors list (link)
  23. ^ Hospitaleche, Carolina A. (2014). "New giant penguin bones from Antarctica: Systematic and paleobiological significance". Comptes Rendus Palevol. 13 (7): 555–560. Bibcode:2014CRPal..13..555A. doi:10.1016/j.crpv.2014.03.008. hdl:11336/32571.
  24. ^ Ksepka, Daniel T. (2014). "Flights of fancy in avian evolution". American Scientist. 102 (1): 39. doi:10.1511/2014.106.36. ISSN 0003-0996. JSTOR 43707746.
  25. ^ a b c d e f g Meat Weights and Nutritional Yield Values for New Zealand Archaeofauna
  26. ^ Alexander, David E. (24 July 2007). "Ancient Argentavis soars again". Proceedings of the National Academy of Sciences of the United States of America. 104 (30): 12233–12234. Bibcode:2007PNAS..10412233A. doi:10.1073/pnas.0705515104. PMC 1941455. PMID 17640902.
  27. ^ a b c Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of Patagonia" [1], geredigeerd door Sergio F. Vizcaíno, Sergio F. Vizcno, Richard F. Kay en M. Susana Bargo
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