Squalius

Squalius
	Squalius carolitertii
	Squalius pyrenaicus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Actinopterygii
Order:	Cypriniformes
Family:	Cyprinidae
Subfamily:	Leuciscinae
Genus:	Squalius; Bonaparte, 1837
Type species
	Leuciscus squalus; Bonaparte, 1837
Synonyms
	Cephalopsis Fitzinger, 1873; Iberocypris Doadrio, 1980; Microlepis Bonaparte, 1846

Squalius is a genus of fish in the family Cyprinidae found in Europe and Asia.^[1] Hybridization is not rare in the Cyprinidae, including this genus. S. alburnoides is known to be of ancient hybrid origin, with the paternal lineage deriving from a prehistoric species related to Anaecypris; the latter mated with ancestral S. pyrenaicus. Present-day S. alburnoides mates with sympatric congeners of other species.^[2]

Species

There are currently 52 recognized species in this genus:

Squalius adanaensis Turan, Kottelat & Doğan, 2013 (Adana chub) ^[3]
Squalius agdamicus S. N. Kamensky, 1901
Squalius alburnoides (Steindachner, 1866)
Squalius albus (Bonaparte, 1838)
Squalius anatolicus (Bogutskaya, 1997) (Beyşehir chub) ^[4]
Squalius aphipsi (A. I. Aleksandrov, 1927) (Aphips chub)
Squalius aradensis (Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998)
Squalius aristotelis Özuluğ & Freyhof, 2011 (Tuzla chub) ^[4]
Squalius berak Heckel, 1843 (Mesopotamian chub) ^[5]
Squalius cappadocicus Özuluğ & Freyhof, 2011 (Cappadocian chub) ^[4]
Squalius carinus Özuluğ & Freyhof, 2011 (Chocolate chub) ^[4]
Squalius carolitertii (Doadrio, 1988)
Squalius castellanus Doadrio, Perea & F. M. Alonso, 2007
Squalius cephaloides (Battalgil, 1942) (Thick-lipped chub) ^[4]
Squalius cephalus (Linnaeus, 1758) (Common chub)
Squalius cii (J. Richardson, 1857) ^[4]
Squalius fellowesii (Günther, 1868) (Aegean chub) ^[4]
Squalius ghigii (Gianferrari, 1927) (Rhodes minnow)
Squalius illyricus Heckel & Kner, 1857
Squalius janae Bogutskaya & Zupančič, 2010
Squalius keadicus (Stephanidis, 1971)
Squalius kosswigi (M. S. Karaman (sr), 1972) ^[4]
Squalius kottelati Turan, Yılmaz & Kaya, 2009
Squalius laietanus Doadrio, Kottelat & de Sostoa, 2007
Squalius lepidus Heckel, 1843
Squalius lucumonis (Bianco, 1983)
Squalius malacitanus Doadrio & Carmona, 2006 (Málaga chub)
Squalius microlepis Heckel, 1843
Squalius moreoticus (Stephanidis, 1971) (Stymphalia chub)
Squalius namak Khaefi, Esmaeili, Sayyadzadeh, Geiger & Freyhof, 2016 ^[6]
Squalius orientalis (Nordmann, 1840)
Squalius orpheus Kottelat & Economidis, 2006 (Orpheus chub)
Squalius palaciosi (Doadrio, 1980) (Jándula chub)
Squalius pamvoticus (Stephanidis, 1939) (Pamvotis chub)
Squalius peloponensis (Valenciennes, 1844) (Peloponnese chub)
Squalius platyceps Zupančič, Marić, Naseka & Bogutskaya, 2010
Squalius prespensis (Fowler, 1977)
Squalius pursakensis (Hankó (hu), 1925) (Sakarya chub) ^[4]
Squalius pyrenaicus (Günther, 1868)
Squalius recurvirostris Özuluğ & Freyhof, 2011 (Akşehir chub) ^[4]
Squalius seyhanensis Turan, Kottelat & Doğan, 2013 (Seyhan chub) ^[3]
Squalius spurius Heckel, 1843 (Orontes chub)
Squalius squaliusculus Kessler, 1872 (Syr-Darya dace)
Squalius squalus (Bonaparte, 1837)
Squalius svallize Heckel & Kner, 1857
Squalius tenellus Heckel, 1843
Squalius torgalensis (Coelho, Bogutskaya, Rodrigues & Collares-Pereira, 1998)
Squalius turcicus De Filippi, 1865 (Transcaucasian chub)
Squalius ulanus Günther, 1899
Squalius valentinus Doadrio & Carmona, 2006 (Valencia chub)
Squalius vardarensis S. L. Karaman, 1928
Squalius zrmanjae S. L. Karaman, 1928

References

^ Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Squalius". FishBase. October 2016 version.
^ Collares-Pereira, M.J. & Coelho, M.M. (2010): Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex Squalius alburnoides. Journal of Fish Biology, 76 (3): 707–715.
^ ^a ^b Turan, D., Kottelat, M. & Doğan, E. (2013): Two new species of Squalius, S. adanaensis and S. seyhanensis (Teleostei: Cyprinidae), from the Seyhan River in Turkey. Zootaxa, 3637 (3): 308–324.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j Özuluğ, M. & Freyhof, J. (2011): Revision of the genus Squalius in Western and Central Anatolia, with description of four new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 22 (2): 107-148.
^ Esmaeili, H.R., Khaefi, R. & Zamanian Nejad, R. (2016): Historical review on the taxonomy of Squalius berak Heckel, 1843 (Teleostei: Cyprinidae). FishTaxa, 1 (3): 118-126.
^ Khaefi, R., Esmaeili, H.R., Sayyadzadeh, G., Geiger, M.F. & Freyhof, J. (2016): Squalius namak, a new chub from Lake Namak basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4169 (1): 145–159.

[FB-1] Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Squalius". FishBase. October 2016 version.

[Collares2010-2] Collares-Pereira, M.J. & Coelho, M.M. (2010): Reconfirming the hybrid origin and generic status of the Iberian cyprinid complex Squalius alburnoides. Journal of Fish Biology, 76 (3): 707–715.

[Turan2013-3] Turan, D., Kottelat, M. & Doğan, E. (2013): Two new species of Squalius, S. adanaensis and S. seyhanensis (Teleostei: Cyprinidae), from the Seyhan River in Turkey. Zootaxa, 3637 (3): 308–324.

[Ozulug2011-4] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j Özuluğ, M. & Freyhof, J. (2011): Revision of the genus Squalius in Western and Central Anatolia, with description of four new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 22 (2): 107-148.

[Esmaeili2016-5] Esmaeili, H.R., Khaefi, R. & Zamanian Nejad, R. (2016): Historical review on the taxonomy of Squalius berak Heckel, 1843 (Teleostei: Cyprinidae). FishTaxa, 1 (3): 118-126.

[Khaefi2016-6] Khaefi, R., Esmaeili, H.R., Sayyadzadeh, G., Geiger, M.F. & Freyhof, J. (2016): Squalius namak, a new chub from Lake Namak basin in Iran (Teleostei: Cyprinidae). Zootaxa, 4169 (1): 145–159.

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